''Engelhardia spicata'' is a species of plant in the

Juglandaceae The Juglandaceae are a plant family known as the walnut family. They are trees, or sometimes shrubs, in the order Fagales. Members of this family are native to the Americas, Eurasia, and Southeast Asia. The nine or ten genera in the family have ...

family. It is recognized to include the variety ''E. spicata'' var. ''integra'' (Kurz) W.E. Manning ex Steen.

Distribution and habitat

Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's fifth-most populous country, with a population of almost 243 million people, and has the world's second-lar ...

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

Nepal Nepal (; ne, :ne:नेपाल, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in S ...

Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainou ...

, China,

Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...

Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...

, Laos,

Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...

Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...

Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...

New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...

. From near sea level to 2100 m, in forests on mountain slopes and in valleys. In

Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mos ...

it is found in primary evergreen forests and seems to prefer the mountains up to 2500 m, especially frequent in the ''

Casuarina ''Casuarina'' is a genus of 17 tree species in the family Casuarinaceae, native to Australia, the Indian subcontinent, southeast Asia, islands of the western Pacific Ocean, and eastern Africa. It was once treated as the sole genus in the fa ...

'' forests on the volcanoes in Central and

East Java East Java ( id, Jawa Timur) is a Provinces of Indonesia, province of Indonesia located in the easternmost hemisphere of Java island. It has a land border only with the province of Central Java to the west; the Java Sea and the Indian Ocean bord ...

. It is known to locally form pure stands on the western side of Mount Jang in

. Similar local dominance has been observed on Mount Rindjani in

Lombok Lombok is an island in West Nusa Tenggara province, Indonesia. It forms part of the chain of the Lesser Sunda Islands, with the Lombok Strait separating it from Bali to the west and the Alas Strait between it and Sumbawa to the east. It is ...

, it has also been observed pioneering in mountain

savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...

s composed of ''

Pittosporum ''Pittosporum'' ( or ) is a genus of about 200 species of flowering plants in the family Pittosporaceae. The genus is probably Gondwanan in origin; its present range extends from Australasia, Oceania, eastern Asia and some parts of Africa. ''Cit ...

'', '' Homalanthus'' gigantheus, ''

Vernonia arborea ''Vernonia arborea'' is a species of mid-level rainforest tree in the aster family (Asteraceae, or Compositae) that is found in Western Ghats of India and Sri Lanka and is also widespread in the East Indies. It is the tallest of all composite t ...

'', ''

Dodonaea ''Dodonaea'' is a genus of about 70 species of flowering plants, often known as hop-bushes, in the soapberry family, Sapindaceae. It has a cosmopolitan distribution in tropical, subtropical and warm temperate regions of Africa, the Americas, so ...

'' and '' Wendlandia'', It is often

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...

for a short time and then flowering, not in definite periods.

Fossil record

Pollen

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s of ''Engelhardia spicata'' have been recovered from strata of Messinian stage of the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

epoch in Western

Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...

in the

Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia (country), Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range ...

region.The History of the Flora and Vegetation of Georgia by Irina Shatilova, Nino Mchedlishvili, Luara Rukhadze, Eliso Kvavadze,

Georgian National Museum The Georgian National Museum ( ka, საქართველოს ეროვნული მუზეუმი, tr) unifies several leading museums in Georgia. The museum was established within the framework of structural, institutional, and ...

Institute of

Paleobiology Paleobiology (or palaeobiology) is an interdisciplinary field that combines the methods and findings found in both the earth sciences and the life sciences. Paleobiology is not to be confused with geobiology, which focuses more on the interactio ...

Tbilisi Tbilisi ( ; ka, თბილისი ), in some languages still known by its pre-1936 name Tiflis ( ), is the capital and the largest city of Georgia, lying on the banks of the Kura River with a population of approximately 1.5 million p ...

2011,

Gallery

Engelhardia spicata AJT Johnsingh P1100584.JPG, ''Engelhardia spicata'' in Ramnagar Forest,

Uttarakhand Uttarakhand ( , or ; , ), also known as Uttaranchal ( ; the official name until 2007), is a state in the northern part of India. It is often referred to as the "Devbhumi" (literally 'Land of the Gods') due to its religious significance and ...

, India Engelhardia spicata seeds, by Omar Hoftun.jpg, Seeds File:Naturalis Biodiversity Center - L.2096246 - Sixtus, Ernst Philipp - Engelhardtia spicata Blume - Artwork.jpeg, Artwork by Ernst Philipp Sixtus of ''Engelhardia spicata'' flowers

References

spicata Flora of China Flora of tropical Asia {{Fagales-stub