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__NOTOC__ Euryapsida is a
polyphyletic A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...
(unnatural, as the various members are not closely related) group of sauropsids that are distinguished by a single temporal fenestra, an opening behind the
orbit In celestial mechanics, an orbit (also known as orbital revolution) is the curved trajectory of an object such as the trajectory of a planet around a star, or of a natural satellite around a planet, or of an artificial satellite around an ...
, under which the post-orbital and squamosal bones articulate. They are different from Synapsida, which also have a single opening behind the orbit, by the placement of the fenestra. In synapsids, this opening is below the articulation of the post-orbital and squamosal bones. It is now commonly believed that euryapsids (particularly sauropterygians) are in fact diapsids (which have two fenestrae behind the orbit) that lost the lower temporal fenestra. Euryapsids are usually considered entirely extinct, although
turtle Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...
s might be part of the sauropterygian clade while other authors disagree. Euryapsida may also be a synonym of Sauropterygia ''sensu lato''. The ichthyosaurian skull is sometimes described as having a ''metapsid'' (or ''parapsid'') condition instead of a truly euryapsid one. In ichthyosaurs, the squamosal bone is never part of the fenestra's margin. Parapsida was originally a taxon consisting of ichthyosaurs, squamates, protorosaurs, araeoscelidans and pleurosaurs. Historically, a variety of reptiles with upper fenestrae, either alone or with a lower emargination, have been considered euryapsid or parapsid, and to have had their patterns of fenestration originate separately from those of diapsids. This includes araeoscelidans,
mesosaur Mesosaurs ("middle lizards") were a group of small aquatic reptiles that lived during the early Permian period ( Cisuralian), roughly 299 to 270 million years ago. Mesosaurs were the first known aquatic reptiles, having apparently returned to a ...
s,
squamate Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest Order (biology), order of reptiles; most members of which are commonly known as Lizard, lizards, with the group also including Snake, snakes. With over 11,991 species, it i ...
s, pleurosaurids, weigeltisaurids, protorosaurs, and trilophosaurs. With the exception of mesosaurs, which only have the lower temporal opening, all of these are universally agreed to be diapsids which either secondarily closed the lower opening (araeoscelids, trilophosaurs) or lost the lower bar (squamates, pleurosaurs, protorosaurs). Euryapsida was proposed by Edwin H. Colbert as a substitute for the earlier term Synaptosauria, originally created by Edward D. Cope for a taxon including sauropterygians, turtles and rhynchocephalians. Baur removed the rhynchocephalians from Synaptosauria and Williston later resurrected the taxon, including only Sauropterygia ( Nothosauria and
Plesiosaur The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...
ia) and
Placodontia Placodonts ("Tablet (pharmacy), tablet tooth, teeth") are an Extinction, extinct order (biology), order of marine reptiles that lived during the Triassic period, becoming extinct at the end of the period. They were part of Sauropterygia, the gro ...
in it. The terms Enaliosauria and Halisauria have also been used for a taxon including ichthyosaurs and sauropterygians. Some 21st century studies have found that ichthyosaurs, thalattosaurs and sauropterygians were close relatives, either as stem-archosaurs or as stem-saurians.


See also

* Anapsid * Diapsid * Synapsida


References

Polyphyletic groups Prehistoric marine reptiles Prehistoric reptile taxonomy {{paleo-reptile-stub