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The Eurasian brown bear (''Ursus arctos arctos'') is one of the most common subspecies of the
brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is ...
, and is found in much of
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelag ...
. It is also called the European brown bear, common brown bear, common bear, and colloquially by many other names. The genetic diversity of present-day brown bears (''Ursus arctos'') has been extensively studied over the years and appears to be geographically structured into five main clades based upon analysis of the
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondrion, mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mit ...
.


Description

The Eurasian brown bear has brown fur, which ranges from yellowish-brown to dark brown, red-brown, and almost black in some cases; albinism has also been recorded. The fur is dense to varying degrees and the hair can grow up to in length. The head normally is quite round in shape and has relatively small rounded ears, a wide skull and a mouth equipped with 42 teeth, including predatory teeth. It has a powerful bone structure and large paws equipped with claws that can grow up to in length. The weight varies depending on habitat and the time of the year. A full-grown male weighs on average between ,''Mammals of the Soviet Union''
Vol.II Part 1a, SIRENIA AND CARNIVORA (Sea cows; Wolves and Bears), p. 611, V.G Heptner and N.P Naumov editors, Science Publishers, Inc. USA. 1998.
and reaches a maximum weight of and length of nearly . Females typically range between 150 and . They have a lifespan of 20 to 30 years in the wild.


History

Eurasian brown bears were used in
Ancient Rome In modern historiography, ancient Rome refers to Roman civilisation from the founding of the city of Rome in the 8th century BC to the collapse of the Western Roman Empire in the 5th century AD. It encompasses the Roman Kingdom (753–50 ...
for fighting in arenas. The strongest bears apparently came from
Caledonia Caledonia (; ) was the Latin name used by the Roman Empire to refer to the part of Great Britain () that lies north of the River Forth, which includes most of the land area of Scotland. Today, it is used as a romantic or poetic name for all ...
and
Dalmatia Dalmatia (; hr, Dalmacija ; it, Dalmazia; see names in other languages) is one of the four historical regions of Croatia, alongside Croatia proper, Slavonia, and Istria. Dalmatia is a narrow belt of the east shore of the Adriatic Sea, stre ...
. In antiquity, the Eurasian brown bear was largely carnivorous, with 80% of its diet consisting of animal matter. However, as its habitat increasingly diminished, the portion of meat in its diet decreased with it until by the late
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire ...
, meat consisted of only 40% of its dietary intake. Today, meat makes up little more than 10–15% of its diet. Whenever possible, the brown bear will consume sheep. Unlike in North America, where an average of two people a year are killed by bears, Scandinavia only has records of three fatal bear attacks within the last century. However, in late 2019, brown bears killed three men in Romania in just over a month.


Species origin

The oldest fossils are from the Choukoutien, China, and date back about 500,000 years. It is known from
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondrion, mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mit ...
studies that during the
Pleistocene The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...
ice age An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages and gre ...
it was too cold for the brown bear to survive in Europe except in three places:
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
, and the
Balkans The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
. Modern research has made it possible to track the origin of the subspecies. The species to which it belongs developed more than 500,000 years ago, and researchers have found that the Eurasian brown bear separated about 850,000 years ago, with one branch based in
Western Europe Western Europe is the western region of Europe. The region's countries and territories vary depending on context. The concept of "the West" appeared in Europe in juxtaposition to "the East" and originally applied to the ancient Mediterranean ...
and the other branch in Russia,
Eastern Europe Eastern Europe is a subregion of the European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russia, wh ...
and
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
. Through research of ''mitochondrial DNA'' (mtDNA), researchers have found that the European family has divided into two
clades A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term ...
—one in the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
and the Balkans, the other in Russia. There is a population in
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, Norway, and Swe ...
that includes bears of the western and eastern lineages. By analyzing the mtDNA of the southern population, researchers have found that they have probably come from populations in the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to ...
in
Southern France Southern France, also known as the South of France or colloquially in French as , is a defined geographical area consisting of the regions of France that border the Atlantic Ocean south of the Marais Poitevin,Louis Papy, ''Le midi atlantique'', A ...
and Spain and the
Cantabrian Mountains , etymology=Named after the Cantabri , photo=Cordillera Cantábrica vista desde el Castro Valnera.jpg , photo_caption=Cantabrian Mountains parallel to the Cantabrian Sea seen from Castro Valnera in an east-west direction. In the background, ...
(Spain). Bears from these populations spread to southern Scandinavia after the last ice age. The northern bear populations originate in the Finnish/Russian population. Probably their ancestors survived the ice age in the ice-free areas west of the
Ural Mountains The Ural Mountains ( ; rus, Ура́льские го́ры, r=Uralskiye gory, p=ʊˈralʲskʲɪjə ˈɡorɨ; ba, Урал тауҙары) or simply the Urals, are a mountain range that runs approximately from north to south through western ...
, and thereafter spread to Northern Europe.


Distribution

Brown bears could once be found across most of Eurasia, compared to the more limited range today. Although included as of
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. Th ...
on the 2006 IUCN Red List of Threatened Species (which refers to the global species, not to the Eurasian brown bear specifically), local populations are becoming increasingly scarce. As the IUCN itself adds: "Least Concern does not always mean that species are not at risk. There are declining species that are evaluated as Least Concern." The brown bear has long been extinct in Ireland, Britain, Denmark, the Netherlands and Belgium, but still exists in Northern Europe and in Russia. Brown bears were present in Britain until no later than AD 1000, by which time they had been exterminated through over-hunting. Globally, the largest population is found east of the Ural mountain range, in the large
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part o ...
n forests; brown bears are also present in smaller numbers in parts of central Asia (former Soviet states). The largest brown bear population in Europe is in
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
, where it has now recovered from an all-time low caused by intensive hunting. Populations in Baltoscandia are similarly, albeit slowly, increasing. They include almost 3,000 bears in Sweden, 2,000 in
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bo ...
, 1,100 in
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, and t ...
and around 100 in
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...
. Large populations can also be found in
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central, Eastern, and Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, Serbia to the southwest, Moldova to the east, a ...
(around 6,000),
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the ...
(around 1,200),
Bosnia and Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and ...
,
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...
(1,200),
Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, and ...
(500-700),
North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, is a country in Southeast Europe. It gained independence in 1991 as one of the successor states of Socialist Feder ...
,
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Mac ...
,
Poland Poland, officially the Republic of Poland, , is a country in Central Europe. Poland is divided into Voivodeships of Poland, sixteen voivodeships and is the fifth most populous member state of the European Union (EU), with over 38 mill ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
, and
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to t ...
. Small but still significant populations can also be found in
Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic The Adriatic Sea () is a body of water separating the ...
,
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wit ...
,
Serbia Serbia (, ; Serbian: , , ), officially the Republic of Serbia (Serbian: , , ), is a landlocked country in Southeastern and Central Europe, situated at the crossroads of the Pannonian Basin and the Balkans. It shares land borders with Hung ...
and
Montenegro ) , image_map = Europe-Montenegro.svg , map_caption = , image_map2 = , capital = Podgorica , coordinates = , largest_city = capital , official_languages = ...
. In 2005, there were an estimated 200 in
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian invas ...
; these populations are part of two distinct metapopulations: the Carpathian with over 5000 individuals, and the Dinaric-Pindos (Balkans) with around 3000 individuals. There is a small but growing population (at least 70 bears) in the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to ...
, on the border between
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
and
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan ar ...
, which was once on the edge of extinction, as well as two subpopulations in the
Cantabrian Mountains , etymology=Named after the Cantabri , photo=Cordillera Cantábrica vista desde el Castro Valnera.jpg , photo_caption=Cantabrian Mountains parallel to the Cantabrian Sea seen from Castro Valnera in an east-west direction. In the background, ...
in Spain (amounting to around 250 individuals). There are also
populations Population typically refers to the number of people in a single area, whether it be a city or town, region, country, continent, or the world. Governments typically quantify the size of the resident population within their jurisdiction using a ...
totalling around 100 bears in the
Abruzzo , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = , demographics1_footnotes = , demographics1_title1 = , demographics1 ...
,
South Tyrol it, Provincia Autonoma di Bolzano – Alto Adige lld, Provinzia Autonoma de Balsan/Bulsan – Südtirol , settlement_type = Autonomous province , image_skyline = , image_alt ...
and
Trentino Trentino ( lld, Trentin), officially the Autonomous Province of Trento, is an autonomous province of Italy, in the country's far north. The Trentino and South Tyrol constitute the region of Trentino-Alto Adige/Südtirol, an autonomous regio ...
regions of
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
. Bears from the aforementioned Italian regions occasionally cross over to bordering Switzerland, which has not hosted a native population since its last bear was shot and killed in Graubünden in 1904. Outside Europe and Russia/the CIS,
clades A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term ...
of brown bear persist in small, isolated and for the most part highly threatened populations in
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkm ...
,
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is bord ...
,
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
, parts of northwest
India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
and central China, and on the island of
Hokkaido is Japan, Japan's Japanese archipelago, second largest island and comprises the largest and northernmost Prefectures of Japan, prefecture, making up its own List of regions of Japan, region. The Tsugaru Strait separates Hokkaidō from Honshu; th ...
in Japan.


Cultural depictions

The historic distribution of bears and the impression the Eurasian brown bear has made on people are reflected in the names of a number of localities (some notable examples include
Berlin Berlin is Capital of Germany, the capital and largest city of Germany, both by area and List of cities in Germany by population, by population. Its more than 3.85 million inhabitants make it the European Union's List of cities in the European U ...
, Bern, Medvednica,
Otepää Otepää (formerly Nuustaku) is a town in Valga County, southern Estonia, it is the administrative centre of Otepää Parish. Otepää is a popular skiing resort, popularly known as the "winter capital" of Estonia (in contrast to the "summer c ...
and Ayu-Dag), as well as personal names—for example, Xiong,
Bernard Bernard ('' Bernhard'') is a French and West Germanic masculine given name. It is also a surname. The name is attested from at least the 9th century. West Germanic ''Bernhard'' is composed from the two elements ''bern'' "bear" and ''hard'' "brav ...
,
Arthur Arthur is a common male given name of Brythonic origin. Its popularity derives from it being the name of the legendary hero King Arthur. The etymology is disputed. It may derive from the Celtic ''Artos'' meaning “Bear”. Another theory, more w ...
, Ursula, Urs, Ursicinus,
Orsolya Orsolya is a Hungarian variant of Ursula meaning "little bear" or "bear cub", derived from a diminutive form of the Latin word ''ursa'' "she-bear". Saint Ursula was a legendary virgin princess of the 4th century who was martyred by the Huns while ...
, Björn, Nedved,
Medvedev Medvedev (russian: Медве́дев) and female Medvedeva (Медве́дева), from Russian ''medved’'' (медве́дь), meaning the animal "bear", are Slavic surnames. Notable bearers of the name include: Medvedev (male form): *Alexander ...
, and Otso. Bears of this subspecies appear very frequently in the fairy tales and fables of Europe, in particular tales collected by Jakob and Wilhelm Grimm. The European brown bear was once common in Germany and alpine lands like
Northern Italy Northern Italy ( it, Italia settentrionale, it, Nord Italia, label=none, it, Alta Italia, label=none or just it, Nord, label=none) is a geographical and cultural region in the northern part of Italy. It consists of eight administrative Regions ...
, Eastern France, and most of Switzerland, and thus appears in tales of various dialects of German. The bear is traditionally regarded as the symbol of Russian (military and political) might. It is also
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bo ...
's national animal; and in
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...
, a brown bear is depicted on the reverse of the Croatian 5 kuna coin, minted since 1993.Croatian National Bank
: ttp://www.hnb.hr/novcan/kovanice/e5kuna.htm?tsfsg=fa39efd7c6602f992d2ea120f7b64c3e 5 Kuna Coin. Retrieved on 31 March 2009.


See also

* Bear conservation *
Marsican brown bear The Marsican brown bear (''Ursus arctos arctos'', formerly ''Ursus arctos marsicanus''), also known as the "Apennine brown bear", and in Italian, is a critically endangered population or subspecies of the Eurasian brown bear, with a range rest ...
* Cantabrian brown bear * Syrian brown bear


References


External links


Det Skandinaviske Bjørneprosjektet

Large Carnivore Initiative for Europe
{{Authority control Carnivorans of Asia Carnivorans of Europe Mammals of the Middle East Least concern biota of Asia Least concern biota of Europe