Ejaculation is the discharge of

semen Semen, also known as seminal fluid, is a bodily fluid that contains spermatozoon, spermatozoa which is secreted by the male gonads (sexual glands) and other sexual organs of male or hermaphrodite, hermaphroditic animals. In humans and placen ...

(the ''ejaculate''; normally containing

sperm Sperm (: sperm or sperms) is the male reproductive Cell (biology), cell, or gamete, in anisogamous forms of sexual reproduction (forms in which there is a larger, female reproductive cell and a smaller, male one). Animals produce motile sperm ...

) from the

penis A penis (; : penises or penes) is a sex organ through which male and hermaphrodite animals expel semen during copulation (zoology), copulation, and through which male placental mammals and marsupials also Urination, urinate. The term ''pen ...

through the urethra. It is the final stage and natural objective of male sexual stimulation, and an essential component of natural conception. After forming an

erection An erection (clinically: penile erection or penile tumescence) is a Physiology, physiological phenomenon in which the penis becomes firm, engorged, and enlarged. Penile erection is the result of a complex interaction of psychological, neural, ...

, many men emit pre-ejaculatory fluid during stimulation prior to ejaculating. Ejaculation involves involuntary contractions of the pelvic floor and is normally linked with orgasm. It is a normal part of male human sexual development. Ejaculation can occur spontaneously during

sleep Sleep is a state of reduced mental and physical activity in which consciousness is altered and certain Sensory nervous system, sensory activity is inhibited. During sleep, there is a marked decrease in muscle activity and interactions with th ...

(a nocturnal emission or "wet dream") or in rare cases because of prostatic disease. '' Anejaculation'' is the condition of being unable to ejaculate. '' Dysejaculation'' is an ejaculation that is painful or uncomfortable. Retrograde ejaculation is the backward flow of semen from the urethra into the bladder.

Premature ejaculation Premature ejaculation (PE) is a male sexual dysfunction that occurs when a male Ejaculation, expels semen (and most likely experiences orgasm) soon after beginning sexual activity, and with minimal penile stimulation. It has also been called ''e ...

happens shortly after initiating sexual activity, and hinders prolonged

sexual intercourse Sexual intercourse (also coitus or copulation) is a sexual activity typically involving the insertion of the Erection, erect male Human penis, penis inside the female vagina and followed by Pelvic thrust, thrusting motions for sexual pleasure ...

. A

vasectomy Vasectomy is an elective surgical procedure that results in male sterilization, often as a means of permanent contraception. During the procedure, the male vasa deferentia are cut and tied or sealed so as to prevent sperm from entering into ...

alters the composition of the ejaculate as a form of birth control.

Phases

Stimulation

The normal precursor to ejaculation is sexual arousal of the male, leading to the

of the

, though not all arousals or erections lead to ejaculation, and ejaculation does not require erection. Penile sexual stimulation during

masturbation Masturbation is a form of autoeroticism in which a person Sexual stimulation, sexually stimulates their own Sex organ, genitals for sexual arousal or other sexual pleasure, usually to the point of orgasm. Stimulation may involve the use of han ...

or vaginal, anal, oral, manual, or non-penetrative sexual activity may provide the necessary stimulus for a man to achieve orgasm and ejaculation. With regard to intravaginal ejaculation latency, men typically reach orgasm five to seven minutes after the start of penile-vaginal intercourse, taking into account their desire and that of their partners, but 10 minutes is also a common intravaginal ejaculation latency. Prolonged stimulation either through foreplay (kissing, petting and direct stimulation of erogenous zones before penetration during intercourse) or stroking (during masturbation) leads to adequate arousal and production of pre-ejaculatory fluid. Infectious agents (including HIV) can be present in pre-ejaculate.

Emission phase

Once the penis has achieved sufficient stimulation for the man to reach orgasm, ejaculation begins. The initial stage of ejaculation, called emission, is controlled by a reflex in the sympathetic spinal cord. Sperm undergo their final developmental changes within the epididymis, where they are held until being ejaculated.

Expulsion phase

Ejaculation reaches its peak in the expulsion phase, which involves the discharge of semen from the urethral opening. This ejection is driven by coordinated contractions of the pelvic muscles, including the bulbospongiosus and pubococcygeus muscles. For the semen to be expelled out of the penis, the bladder neck stays shut while the external

urethral sphincter The urethral sphincters are two muscles used to control the exit of urine in the urinary bladder through the urethra. The two muscles are either the external sphincter muscle of male urethra, male or external sphincter muscle of female urethra, fe ...

is relaxed. These rhythmic contractions are part of the male orgasm under the control of a

spinal reflex The stretch reflex (myotatic reflex), or more accurately ''muscle stretch reflex'', is a muscle contraction in response to stretching a muscle. The function of the reflex is generally thought to be maintaining the muscle at a constant length but t ...

at the level of the spinal nerves S2–4 via the pudendal nerve. Although the external sphincter and pelvic muscles can be voluntarily controlled, any voluntary control during semen expulsion is not evident. The expulsion phase is considered an extension of the emission phase, triggered by reaching a certain level of spinal nerve activation. The typical male orgasm lasts several seconds.

is when ejaculation occurs before it is desired. Otherwise, if a man is unable to ejaculate after prolonged sexual stimulation in spite of his desire, it is called delayed ejaculation or anorgasmia. An orgasm that is not accompanied by ejaculation is known as a dry orgasm. At start of orgasm, pulses of semen begin to flow from the urethra, reach a peak of discharge and then diminish in flow. The typical orgasm consists of 10 to 15 contractions, although the man may not be consciously aware of so many. After the first contraction, ejaculation continues to completion involuntarily. During this stage ejaculation cannot be stopped. The rate of contractions gradually slows throughout the orgasm. Initial contractions occur on average every 0.6 seconds with an increasing increment of 0.1 seconds per contraction. Contractions of most men proceed at regular rhythmic intervals through their duration. Many men also experience irregular contractions at the end of the orgasm. Ejaculation usually begins during the first or second contraction of orgasm. For most men, the first ejection occurs during the second contraction, which is typically the largest, expelling 40% or more of total semen discharge. After this peak, the quantity of semen emitted by the penis diminishes as the contractions lessen in intensity. The muscle contractions of the orgasm can continue after ejaculation with no additional semen discharge. A small sample study of seven men showed an average of seven spurts of semen followed by an average of 10 more contractions with no semen expelled. This study also found a high correlation between number of spurts of semen and total ejaculate volume, i.e., larger semen volumes resulted from additional pulses of semen rather than larger individual spurts. Alfred Kinsey measured the distance of ejaculation, in "some hundreds" of men. In three-quarters of men tested, ejaculate "is propelled with so little force that the liquid is not carried more than a minute distance beyond the tip of the penis." In contrast to those test subjects, Kinsey noted "In other males the semen may be propelled from a matter of some inches to a foot or two, or even as far as five or six and (rarely) eight feet". Masters and Johnson report ejaculation distance to be no greater than . During the series of contractions that accompany ejaculation, semen is propelled from the urethra at , close to .

Refractory period

Most men experience a refractory period immediately following an orgasm, during which they are unable to achieve another erection, and a longer period before they are capable of achieving another ejaculation. During this time a male feels a deep and often pleasurable sense of relaxation, usually in the groin and thighs. The length of the refractory period varies considerably, even for a given individual. Age affects the recovery time, with younger men recovering faster than older men, though not always. Whereas some men have refractory periods of 15 minutes or more, others are able to experience sexual arousal immediately after ejaculation. A short recovery period may allow partners to continue sexual play relatively uninterrupted by ejaculation. Some men may experience their penis becoming hypersensitive to stimulation after ejaculation, which can make sexual stimulation unpleasant even while they may be sexually aroused. Some men are able to achieve multiple orgasms, with or without the typical sequence of ejaculation and refractory period. Some of those men report not noticing refractory periods, or are able to maintain erection by "sustaining sexual activity with a full erection until they passed their refractory time for orgasm when they proceeded to have a second or third orgasm".

Volume

The force and amount of semen that is ejected during ejaculation varies widely among men, containing between 0.1 and 10 milliliters (for comparison, a teaspoon holds 5 ml and a tablespoon, 15 ml). Adult semen volume is affected by the time that has passed since his previous ejaculation; larger semen volumes develop with longer abstinence. The duration of the stimulation leading to ejaculation can affect the volume. Abnormally low semen volume is known as hypospermia and abnormally high semen volume is called hyperspermia. One possible underlying cause of low volume or complete lack of semen is ejaculatory duct obstruction. It is normal for the amount of semen to diminish with age.

Quality

The number of sperm in an ejaculation varies widely, depending on many factors including the time since the previous ejaculation, age, stress levels, and

testosterone Testosterone is the primary male sex hormone and androgen in Male, males. In humans, testosterone plays a key role in the development of Male reproductive system, male reproductive tissues such as testicles and prostate, as well as promoting se ...

. Longer time of sexual stimulation immediately preceding ejaculation can result in higher concentration of sperm. An unusually low sperm count, distinguished from low semen volume, is known as oligospermia, and the absence of any sperm from the semen is termed azoospermia.

Development

During puberty

The first ejaculation in males often occurs about 12 months after the onset of

puberty Puberty is the process of physical changes through which a child's body matures into an adult body capable of sexual reproduction. It is initiated by hormonal signals from the brain to the gonads: the ovaries in a female, the testicles i ...

, generally through

or nocturnal emission (wet dreams). This first

volume is small. The typical ejaculation over the following three months produces less than 1 ml of semen. The semen produced during early puberty is also typically clear. After ejaculation this early semen remains jellylike and, unlike semen from mature males, fails to liquefy. A summary of semen development is shown in Table 1. Most first ejaculations (90%) lack sperm. Of the few early ejaculations that do contain sperm, the majority of sperm (97%) lack motion. The remaining sperm (3%) have abnormal motion. As the male proceeds through puberty, the semen develops mature characteristics with increasing quantities of normal sperm. Semen produced 12 to 14 months after the first ejaculation liquefies after a short period of time. Within 24 months of the first ejaculation, the semen volume and the quantity and characteristics of the sperm match that of adult male semen. Ejaculate is jellylike and fails to liquefy.
Most samples liquefy. Some remain jellylike.
Ejaculate liquefies within an hour.

Control from the central nervous system

There is a central pattern generator in the

spinal cord The spinal cord is a long, thin, tubular structure made up of nervous tissue that extends from the medulla oblongata in the lower brainstem to the lumbar region of the vertebral column (backbone) of vertebrate animals. The center of the spinal c ...

, made up of groups of spinal interneurons, that is involved in the rhythmic response of ejaculation. This is known as the ''spinal generator for ejaculation''. To map the neuronal activation of the brain during the ejaculatory response, researchers have studied the expression of c-Fos, a proto-oncogene expressed in neurons in response to stimulation by hormones and neurotransmitters. Expression of c-Fos in the following areas has been observed: * medial preoptic area (MPOA) * lateral septum, bed nucleus of the stria terminalis * paraventricular nucleus of hypothalamus (PVN) * ventromedial nucleus of the hypothalamus, medial

amygdala The amygdala (; : amygdalae or amygdalas; also '; Latin from Greek language, Greek, , ', 'almond', 'tonsil') is a paired nucleus (neuroanatomy), nuclear complex present in the Cerebral hemisphere, cerebral hemispheres of vertebrates. It is c ...

* ventral premammillary nuclei * ventral tegmental area * central tegmental field * mesencephalic central gray * peripeduncular nuclei * parvocellular subparafascicular nucleus (SPF) within the posterior

thalamus The thalamus (: thalami; from Greek language, Greek Wikt:θάλαμος, θάλαμος, "chamber") is a large mass of gray matter on the lateral wall of the third ventricle forming the wikt:dorsal, dorsal part of the diencephalon (a division of ...

Hands-free ejaculation

Although uncommon, some men can achieve ejaculations during masturbation without any manual stimulation. Such men usually do it by tensing and flexing their abdominal and buttocks muscles along with vigorous fantasizing. Others may do it by relaxing the area around the

, which may result in harder erections especially when extremely aroused. Hands-free ejaculation can also be achieved by prostate stimulation alone, either internally (with the use of

sex toys A sex toy is an object or device that is primarily used to facilitate Sexual stimulation, sexual pleasure, such as a dildo, artificial vagina or Vibrator (sex toy), vibrator. Many popular sex toys are designed to resemble human genitals, and ma ...

, fingers or performing

anal sex Anal sex or anal intercourse principally means the insertion and pelvic thrusting, thrusting of the Erection, erect human penis, penis into a person's Human anus, anus, or anus and rectum, for sexual pleasure.Sepages 270–271for anal sex inform ...

or pegging) or externally (such as

perineum The perineum (: perineums or perinea) in placentalia, placental mammals is the space between the anus and the genitals. The human perineum is between the anus and scrotum in the male or between the anus and vulva in the female. The perineum is ...

massages), although prostate orgasms without ejaculation (dry orgasms) are also possible.

Perineum pressing and retrograde ejaculation

Perineum The perineum (: perineums or perinea) in placentalia, placental mammals is the space between the anus and the genitals. The human perineum is between the anus and scrotum in the male or between the anus and vulva in the female. The perineum is ...

pressing results in an ejaculation which is purposefully held back by pressing on either the perineum or the urethra to force the seminal fluid to remain inside. In such a scenario, the seminal fluid stays inside the body and goes to the bladder. Some people do this to avoid making a mess by keeping all the semen inside. As a medical condition, it is called retrograde ejaculation.

Health issues

For most men, no detrimental health effects have been determined from ejaculation itself or from frequent ejaculations, though sexual activity in general can have health or psychological consequences. A small fraction of men have a disease called postorgasmic illness syndrome (POIS), which causes severe muscle pain throughout the body and other symptoms immediately following ejaculation. The symptoms last for up to a week. Some doctors speculate that the frequency of POIS "in the population may be greater than has been reported in the academic literature", and that many POIS sufferers are undiagnosed. It is not clear whether frequent ejaculation has any effect on the risk of

prostate cancer Prostate cancer is the neoplasm, uncontrolled growth of cells in the prostate, a gland in the male reproductive system below the bladder. Abnormal growth of the prostate tissue is usually detected through Screening (medicine), screening tests, ...

. Two large studies examining the issue were "Ejaculation Frequency and Subsequent Risk of Prostate Cancer" and "Sexual Factors and Prostate Cancer". These suggest that frequent ejaculation after puberty offers some reduction of the risk of prostate cancer. The US study involving 29,342 US men aged 46 to 81 years suggested that "high ejaculation frequency was related to decreased risk of total prostate cancer". An Australian study involving 1,079 men with prostate cancer and 1,259 healthy men found that "there is evidence that the more frequently men ejaculate between the ages of 20 and 50, the less likely they are to develop prostate cancer":

Other animals

In mammals and birds, multiple ejaculation is commonplace. During copulation, the two sides of a short-beaked echidna's penis are used sequentially. Alternating between the two sides allows for persistent stimulation to induce ejaculation without impeding the refractory period. In stallions, ejaculation is accompanied by a motion of the tail known as "tail flagging". When a male

wolf The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been recognized, includin ...

ejaculates, his final pelvic thrust may be slightly prolonged. A male rhesus monkey usually ejaculates less than 15 seconds after

sexual penetration Sexual penetration is the insertion of a body part or other object into a body orifice, such as the mouth, vagina or anus, as part of human sexual activity or sexual behavior in non-human animals. The term is most commonly used in statute la ...

. The first report and footage of spontaneous ejaculation in an aquatic mammal was recorded in a wild Indo-Pacific bottlenose dolphin near Mikura Island, Japan, in 2012. In horses, sheep, and cattle, ejaculation occurs within a few seconds, but in

boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a Suidae, suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The speci ...

s, it can last for five to thirty minutes. Ejaculation in boars is stimulated when the spiral-shaped penis interlocks with the female's cervix. A mature boar can produce of semen during one ejaculation. In llamas and alpacas, ejaculation occurs continuously during copulation. The semen of male dogs is ejaculated in three separate phases. The last phase of a male canine's ejaculation occurs during the copulatory tie, and contains mostly prostatic fluid.