Echium Tenue
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''Echium'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Boraginaceae Boraginaceae, the Borago, borage or forget-me-not family, includes about 2,000 species of shrubs, trees, and herbs in 146 to 154 genus, genera with a worldwide distribution. The APG IV system from 2016 classifies the Boraginaceae as single famil ...
that contains about 70
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
and several
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
. Species of ''Echium'' are
native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...
to North Africa, mainland Europe to Central Asia, and the
Macaronesia Macaronesia (; ) is a collection of four volcanic archipelagos in the Atlantic Ocean#North Atlantic, North Atlantic Ocean, off the coast of North Africa and Europe. Each archipelago is made up of a number of list of islands in the Atlantic Oc ...
n islands where the genus reaches its maximum diversity. Twenty-nine species of ''Echium'' are endemic to the Canary, Madeira, and Cape Verde archipelagos. The continental species are
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of ...
, whereas many of the endemic species of the Macaronesian islands are woody perennial shrubs.


Etymology

The Latin genus name ''Echium'' comes from the
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
''echion'', referring to ''
Echium plantagineum ''Echium plantagineum'', commonly known as purple viper's-bugloss, Paterson's curse or Salvation Jane, is a species of the genus '' Echium'' native to western and southern Europe (from southern England south to Iberia and east to the Crimea), no ...
'' and itself deriving from ''echis'' (
viper Vipers are snakes in the family Viperidae, found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, New Zealand, Ireland, and various other isolated islands. They are venomous and have long (relative to non-vipe ...
); the Greek term dates to
Dioscorides Pedanius Dioscorides (, ; 40–90 AD), "the father of pharmacognosy", was a Greek physician, pharmacologist, botanist, and author of (in the original , , both meaning "On Materia medica, Medical Material") , a 5-volume Greek encyclopedic phar ...
, who noted a resemblance between the shape of the nutlets and a viper's head. The genus ''Echium'' was published by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1753.


Cultivation and uses

Many species are used as ornamental and garden plants and may be found in suitable climates throughout the world. In
Crete Crete ( ; , Modern Greek, Modern: , Ancient Greek, Ancient: ) is the largest and most populous of the Greek islands, the List of islands by area, 88th largest island in the world and the List of islands in the Mediterranean#By area, fifth la ...
, '' Echium italicum'' is called ''pateroi'' (πάτεροι) or ''voidoglosses'' (βοϊδόγλωσσες) and its tender shoots are eaten boiled or steamed. ''Echium'' species are used as food plants by the
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e of some
Lepidoptera Lepidoptera ( ) or lepidopterans is an order (biology), order of winged insects which includes butterflies and moths. About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organ ...
species including '' Coleophora onosmella'' and orange swift. In some countries ''Echium'' extract has been used as cure for various diseases and is believed to have beneficial properties.


''Echium'' seed oil

The seed oil from ''E. plantagineum'' contains high levels of alpha-linolenic acid (ALA), gamma-linolenic acid, and stearidonic acid, making it valuable in cosmetic and skin-care applications, with further potential as a functional food, as an alternative to fish oils. However, despite its high ALA content, ''Echium'' seed oil does not increase
docosahexaenoic acid Docosahexaenoic acid (DHA) is an omega−3 fatty acid that is an important component of the human brain, cerebral cortex, skin, and retina. It is given the fatty acid notation 22:6(''n''−3). It can be synthesized from alpha-linolenic acid or ...
and
eicosapentaenoic acid Eicosapentaenoic acid (EPA; also icosapentaenoic acid) is an omega−3 fatty acid. In physiological literature, it is given the name 20:5(''n''−3). It also has the trivial name timnodonic acid. In chemical structure, EPA is a carboxylic acid wi ...
levels.


Invasiveness

Some species have been widely naturalized in Mediterranean climates, including South Africa, Australia, New Zealand, and parts of South America and the United States. For example, ''
Echium plantagineum ''Echium plantagineum'', commonly known as purple viper's-bugloss, Paterson's curse or Salvation Jane, is a species of the genus '' Echium'' native to western and southern Europe (from southern England south to Iberia and east to the Crimea), no ...
'' has become a major
invasive species in Australia Invasive species are a serious threat to endemic Australian species, and an ongoing cost to Australian agriculture. Numerous species arrived with European maritime exploration and colonisation of Australia and steadily since then. There is ...
.


Species

68 species are accepted. #'' Echium acanthocarpum'' #'' Echium aculeatum'' #'' Echium albicans'' #'' Echium amoenum'' #'' Echium anchusoides'' #'' Echium angustifolium'' #'' Echium arenarium'' #'' Echium asperrimum'' #'' Echium auberianum'' #'' Echium × bailaderense'' #'' Echium bethencourtii'' #'' Echium boissieri'' #'' Echium × bond-spraguei'' #'' Echium bonnetii'' #'' Echium brevirame'' #'' Echium callithyrsum'' #'' Echium candicans'' #'' Echium canum'' #'' Echium clandestinum'' #'' Echium creticum'' #'' Echium decaisnei'' #'' Echium flavum'' #'' Echium gaditanum'' #'' Echium giganteum'' #'' Echium glomeratum'' #''
Echium handiense ''Echium'' is a genus of flowering plants in the family (biology), family Boraginaceae that contains about 70 species and several subspecies. Species of ''Echium'' are native plant, native to North Africa, mainland Europe to Central Asia, ...
'' #'' Echium hierrense'' #'' Echium horridum'' #'' Echium humile'' #'' Echium hypertropicum'' #'' Echium italicum'' #'' Echium judaeum'' #'' Echium khuzistanicum'' #'' Echium × lemsii'' #'' Echium leucophaeum'' #'' Echium × lidii'' #'' Echium longifolium'' #'' Echium lusitanicum'' #'' Echium modestum'' #'' Echium montenielluense'' #'' Echium nervosum'' #'' Echium onosmifolium'' #'' Echium pabotii'' #'' Echium parviflorum'' #'' Echium perezii'' #'' Echium petiolatum'' #'' Echium pininana'' #'' Echium pitardii'' #''
Echium plantagineum ''Echium plantagineum'', commonly known as purple viper's-bugloss, Paterson's curse or Salvation Jane, is a species of the genus '' Echium'' native to western and southern Europe (from southern England south to Iberia and east to the Crimea), no ...
'' #'' Echium portosanctense'' #'' Echium rauwolfii'' #'' Echium rosulatum'' #'' Echium rubrum'' #'' Echium sabulicola'' #'' Echium salmanticum'' #'' Echium simplex'' #'' Echium spurium'' #'' Echium stenosiphon'' #'' Echium strictum'' #'' Echium suffruticosum'' #'' Echium sventenii'' #'' Echium x taibiquense'' #'' Echium tenue'' #'' Echium thyrsiflorum'' #'' Echium triste'' #'' Echium trygorrhizum'' #''
Echium tuberculatum ''Echium tuberculatum'' is a species of flowering plant native to Portugal, Morocco, Algeria and Libya in the Iberian Peninsula and North Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the A ...
'' #'' Echium velutinum'' #'' Echium virescens'' #'' Echium vulcanorum'' #''
Echium vulgare ''Echium vulgare'', known as viper's bugloss and blueweed,Dickinson, T.; Metsger, D.; Bull, J.; & Dickinson, R. (2004) ROM Field Guide to Wildflowers of Ontario. Toronto:Royal Ontario Museum, p. 203. is a species of flowering plant in the borag ...
'' #'' Echium webbii'' #'' Echium wildpretii''


Formerly placed here

* '' Megacaryon orientale'' (as ''Echium orientale'' ) * '' Pontechium maculatum'' (as ''Echium maculatum'' )


References

{{Authority control Boraginaceae genera Taxa named by Carl Linnaeus