Dsungaripterus
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''Dsungaripterus'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
dsungaripterid Dsungaripteridae is a group of pterosaurs within the suborder Pterodactyloidea. They were robust pterosaurs with good terrestrial abilities and flight honed for inland settings, and were commonly interpreted as durophagous and possibly piscivorou ...
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
which lived during the
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
in what is now
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
and possibly
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia. It constitutes the southern half of the Korea, Korean Peninsula and borders North Korea along the Korean Demilitarized Zone, with the Yellow Sea to the west and t ...
. Its fossils come from the Wuerho Pterosaur Fauna group (WPF) of the
Tugulu Group The Tugulu Group () is a geological Group in Xinjiang, China whose strata date back to the Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the ...
, comprising sections of the Lianmuqin and Shengjinkou Formations of the
Junggar Basin The Junggar Basin (), also known as the Dzungarian Basin or Zungarian Basin, is one of the largest sedimentary basins in Northwest China. It is located in Dzungaria in northern Xinjiang, and enclosed by the Tarbagatai Mountains of Kazakhstan in t ...
. Both formations have been given various age estimates, with data suggesting at least some of the WPF date to the middle
Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 137.05 ± 0.2 Ma and 132.6 ± 0.2 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretac ...
, but the upper and lower bounds of these pterosaur bearing deposits remain unclear.


Description

''Dsungaripterus weii'' had a wingspan of . Like most dsungaripteroids it had a rather robust skeleton with thick walls and stouty bodily proportions, suggesting a mostly terrestrial lifestyle. The flight style of these animals is unclear, but it was probably punctuated by abrupt landings and extensive flapping. The skull of ''Dsungaripterus'', measuring over long, bore a low bone crest that ran down from the base of the skull to halfway to the beak. ''Dsungaripteruss head and neck were together almost long. Its most notable feature are its long, narrow, upcurved jaws with a pointed tip. It had no teeth in the front part of its jaws, which were probably used to remove prey from cracks in rocks and/or the sandy, muddy inland environments it inhabited. It had knobbly flat teeth more to the back of the jaw that were well suited for crushing the armor of shellfish or other hard objects. Thus, it is commonly interpreted that dsungaripterids were
durophagous Durophagy is the eating behavior of animals that consume hard-shelled or exoskeleton-bearing organisms, such as corals, shelled mollusks, or crabs. It is mostly used to describe fish, but is also used when describing reptiles, including fossil t ...
and possibly
piscivorous A piscivore () is a carnivorous animal that primarily eats fish. Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted rept ...
pterosaurs. Additionally, ''Dsungaripterus'' also had a palate similar to that of
azhdarchoid Azhdarchoidea (International Phonetic Alphabet, , meaning "azhdarchid-like forms") is a group of pterosaurs within the suborder Pterodactyloidea. Pterosaurs belonging to this group lived throughout the Early Cretaceous, Early and Late Cretaceous ...
s.


History of discovery

''Dsungaripterus'' was described in 1964 named by
Yang Zhongjian Yang Zhongjian, also Yang Chung-chien (; 1 June 1897 – 15 January 1979), courtesy name Keqiang (), also known as C.C. (Chung Chien) Young, was a Chinese paleontologist and zoologist. He was one of China's foremost vertebrate paleontologists. ...
. The genus name combines a reference to the Junggar Basin with a Latinized Greek ''pteron'', "wing". The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Dsungaripterus weii'', the
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honoring paleontologist C.M. Wei of the Palaeontological Division, Institute of Science, Bureau of Petroleum of
Xinjiang Xinjiang,; , SASM/GNC romanization, SASM/GNC: Chinese postal romanization, previously romanized as Sinkiang, officially the Xinjiang Uygur Autonomous Region (XUAR), is an Autonomous regions of China, autonomous region of the China, People' ...
. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
is IVPP V-2776, a partial skull and skeleton from the
Lianmuqin Formation The Lianmuqin Formation, also transcribed as Lianmugin Formation,Shengjinkou Formation The Shengjinkou Formation is an Early Cretaceous-aged Konservat-Lagerstätte composed of "interbedded red green and yellow variegated mudstones and siltstones" that is part of the larger Tugulu Group of China. Dinosaur and pterosaur remains have ...
, which includes almost complete skulls. In 1980
Peter Galton Peter Malcolm Galton (born 14 March 1942 in London) is a British vertebrate paleontologist who has to date written or co-written about 190 papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosau ...
renamed ''Pterodactylus brancai'' (Reck 1931) from the
Tendaguru Formation The Tendaguru Formation, or Tendaguru Beds are a highly List of fossiliferous stratigraphic units in Tanzania, fossiliferous Lithostratigraphy, formation and Lagerstätte located in the Lindi Region of southeastern Tanzania. The formation represe ...
into ''Dsungaripterus brancai'', but the identification is now commonly rejected. In 1982 Natasha Bakhurina named a new species, ''Dsungaripterus parvus'', based on a smaller skeleton from
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
. Later, this was renamed into "Phobetor", a preoccupied name, and in 2009 concluded to be identical to ''
Noripterus ''Noripterus'' (meaning "lake wing" from Mongolian language, Mongolian ''nuur'', "lake" and Greek ''pteron'', "wing") is a genus of dsungaripteridae, dsungaripterid pterodactyloidea, pterodactyloid pterosaur from Lower Cretaceous-age Wuerho Ptero ...
''. A
dsungaripterid Dsungaripteridae is a group of pterosaurs within the suborder Pterodactyloidea. They were robust pterosaurs with good terrestrial abilities and flight honed for inland settings, and were commonly interpreted as durophagous and possibly piscivorou ...
wing finger phalanx was reported in 2002 from the
Hasandong Formation The Hasandong Formation () is an Early Cretaceous (Aptian to Albian) geologic Formation (geology), formation in South Korea. It has been dated to the late Aptian and earliest Albian, between 118.0 ± 2.6 Ma and 112.4 ± 1.3 Ma. Dinosaur ...
of
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia. It constitutes the southern half of the Korea, Korean Peninsula and borders North Korea along the Korean Demilitarized Zone, with the Yellow Sea to the west and t ...
, and was identified in 2015 and 2018 as ''Dsungaripterus''? cf. ''D. weii''.


Classification

''Dsungaripterus'' was classified by Yang as a member of the
Dsungaripteridae Dsungaripteridae is a group of pterosaurs within the suborder Pterodactyloidea. They were robust pterosaurs with good terrestrial abilities and flight honed for inland settings, and were commonly interpreted as durophagous and possibly piscivoro ...
. Below is a
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
showing the results of a
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analysis presented by Andres and colleagues in 2014. They recovered ''Dsungaripterus'' within the clade
Dsungaripteromorpha Azhdarchoidea (International Phonetic Alphabet, , meaning "azhdarchid-like forms") is a group of pterosaurs within the suborder Pterodactyloidea. Pterosaurs belonging to this group lived throughout the Early Cretaceous, Early and Late Cretaceous ...
(a subgroup within the Azhdarchoidea), more specifically within the Dsungaripteridae, sister taxon to ''
Domeykodactylus ''Domeykodactylus'' is an extinct genus of dsungaripterid pterosaur from the Early Cretaceous (Berriasian-Albian)-age Santa Ana Formation at Quebrada de la Carreta in Antofagasta, Chile. Discovery and naming The genus was named in 2000 by Dav ...
''. Their cladogram is shown below. In 2019, a different topology, this time by Kellner and colleagues, was published. In this study, ''Dsungaripterus'' was recovered outside the Azhdarchoidea, within the larger group
Tapejaroidea Pterodactyloidea ( ; derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contai ...
, sister taxon to ''Noripterus''. The cladogram of the analysis is shown below.


See also

*
List of pterosaur genera This list of pterosaurs is a comprehensive listing of all Genus, genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considere ...
*
Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and Biological taxonomy, taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic Era (ge ...


References

{{Portal bar, Paleontology, China Dsungaripteridae Early Cretaceous pterosaurs of Asia Taxa named by Yang Zhongjian Fossil taxa described in 1964