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Darwin's rhea or the lesser rhea (''Rhea pennata'') is a large
flightless bird Flightless birds are birds that cannot Bird flight, fly, as they have, through evolution, lost the ability to. There are over 60 extant species, including the well-known ratites (ostriches, emus, cassowary, cassowaries, Rhea (bird), rheas, an ...
, the smaller of the two
extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...
species of rheas. It is found in the
Altiplano The Altiplano (Spanish language, Spanish for "high plain"), Collao (Quechuan languages, Quechua and Aymara language, Aymara: Qullaw, meaning "place of the Qulla people, Qulla") or Andean Plateau, in west-central South America, is the most extens ...
and
Patagonia Patagonia () is a geographical region that includes parts of Argentina and Chile at the southern end of South America. The region includes the southern section of the Andes mountain chain with lakes, fjords, temperate rainforests, and glaciers ...
in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
.


Description

The lesser rhea stands at tall. Length is and weight is . Like most
ratite Ratites () are a polyphyletic group consisting of all birds within the infraclass Palaeognathae that lack keels and cannot fly. They are mostly large, long-necked, and long-legged, the exception being the kiwi, which is also the only nocturnal ...
s, it has a small head and a small bill, the latter measuring , but has long legs and a long neck. It has relatively larger wings than other
ratite Ratites () are a polyphyletic group consisting of all birds within the infraclass Palaeognathae that lack keels and cannot fly. They are mostly large, long-necked, and long-legged, the exception being the kiwi, which is also the only nocturnal ...
s, enabling it to run particularly well. It can reach speeds of 60 km/h (37 mph), enabling it to outrun predators. The sharp claws on the toes are effective weapons. Their feathers are similar to those of ostriches, in that they have no aftershaft.Perrins, C. (1987) Their plumage is spotted brown and white, and the upper part of their tarsus is feathered. The tarsus is long and has 18 horizontal plates on the front.


Etymology

It is known as ''ñandú petiso'', or ''ñandú del norte'', in Argentina, where the majority live. Other names are ''suri'' and ''choique''. The name ''ñandú'' comes from the greater rhea's name in Guaraní, ''ñandu guazu'', meaning big spider, possibly in relation to their habit of alternately opening and lowering their wings when they run. In English, Darwin's rhea gets its
scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
from Rhea, a Greek goddess, and ''pennata'', meaning winged. The specific name was bestowed in 1837 by Darwin's contemporary and rival
Alcide d'Orbigny Alcide Charles Victor Marie Dessalines d'Orbigny (6 September 1802 – 30 June 1857) was a French naturalist who made major contributions in many areas, including zoology (including malacology), palaeontology, geology, archaeology and anthropol ...
, who first described the bird to Europeans from a specimen from the lower Río Negro south of
Buenos Aires Buenos Aires, controlled by the government of the Autonomous City of Buenos Aires, is the Capital city, capital and largest city of Argentina. It is located on the southwest of the Río de la Plata. Buenos Aires is classified as an Alpha− glob ...
,
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
.Davies, S.J.J.F. (2003) As late as 2008, it was classified in the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genus ''Pterocnemia''. This word is formed from two Greek words ''pteron'', meaning feathers, and ''knēmē'', meaning the leg between the knee and the ankle, hence feather-legged, alluding to their feathers that cover the top part of the leg. In 2008, the SACC subsumed ''Pterocnemia'' into the genus '' Rhea''.


Taxonomy

Three
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
have traditionally been recognized: * ''R. p. garleppi'' is found in the puna of southeastern
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
, southwestern
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
, and northwestern
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
.Clements, J (2007) * ''R. p. tarapacensis'' is found in the puna of northern
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
from the region of Arica and Parinacota to
Antofagasta Antofagasta () is a port city in northern Chile, about north of Santiago. It is the capital of Antofagasta Province and Antofagasta Region. According to the 2015 census, the city has a population of 402,669. Once claimed by Bolivia follo ...
.Jaramillo ''et al.'' (2003) * ''R. p. pennata'' is found in the Patagonian
steppes In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropical gr ...
of Argentina and Chile. The IUCN formerly considered the two northern
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
''R. p. tarapacensis'' and ''R. p. garleppi'' as a separate species, the puna rhea (''R. tarapacensis''). Both ''garleppi'' and ''tarapacensis'' were described by Charles Chubb in 1913.Brands, S. (2008) The IUCN now recognizes them as one species, but notes the need for more work on the taxonomy of this species. It is possible ''garleppi'' should be considered a
junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...
of ''tarapacensis''.


Behavior

The lesser rhea is mainly a herbivore, with the odd small animal (lizards, beetles, grasshoppers) eaten on occasion. It predominantly eats
saltbush Saltbush is a vernacular plant name that most often refers to ''Atriplex'', a genus of about 250 plants distributed worldwide from subtropical to subarctic regions. ''Atriplex'' species are native to Australia, North and South America, and Eurasia. ...
and fruits from cacti, as well as grasses, roots, seeds, and leaves. They tend to be quiet birds, except as chicks when they whistle mournfully, and as males looking for a female, when they emit a booming call. The males of this species become aggressive once they are incubating eggs, even towards females. The females thus lay the later eggs near the nest, rather than in it. Most of the eggs are moved into the nest by the male, but some remain outside, where they rot and attract flies. The male, and later the chicks, eat these flies. The incubation period is 30–44 days, and the clutch size is from 5–55 eggs. The eggs are and are greenish yellow. Chicks mature by three years of age. Outside the breeding season, Darwin's rhea is quite sociable: it lives in groups of from 5 to 30 birds, of both sexes and a variety of ages.


Distribution and habitat

Darwin's rhea lives in areas of open scrub in the grasslands of Patagonia and on the
Andean The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long and wide (widest between 18°S ...
plateau (the
Altiplano The Altiplano (Spanish language, Spanish for "high plain"), Collao (Quechuan languages, Quechua and Aymara language, Aymara: Qullaw, meaning "place of the Qulla people, Qulla") or Andean Plateau, in west-central South America, is the most extens ...
), through the countries of
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
,
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
,
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
, and
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
. All subspecies prefer grasslands, brushlands and marshland. However, the
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
prefers elevations less than , where the other subspecies typically range from , but locally down to in the south.Birdlife International (2016)


History of the discovery of the genus ''Rhea''

During the second voyage of HMS ''Beagle'', the young naturalist
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
made many trips on land, and around August 1833 heard from
gaucho A gaucho () or gaúcho () is a skilled horseman, reputed to be brave and unruly. The figure of the gaucho is a folk symbol of Argentina, Paraguay, Uruguay, Rio Grande do Sul in Brazil, the southern part of Bolivia, and the south of Chilean Patago ...
s in the Río Negro area of Northern Patagonia about the existence of a smaller rhea, "a very rare bird which they called the Avestruz Petise". He continued searching fruitlessly for this bird, and the ''Beagle'' sailed south, putting in at Port Desire in southern Patagonia on 23 December. On the following day, Darwin shot a
guanaco The guanaco ( ; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids; the other species is the vicuña, which lives at higher elevations. Etymology The gua ...
(similar to a
llama The llama (; or ) (''Lama glama'') is a domesticated South American camelid, widely used as a List of meat animals, meat and pack animal by Inca empire, Andean cultures since the pre-Columbian era. Llamas are social animals and live with ...
) which provided them with a Christmas meal, and in the first days of January, the artist Conrad Martens shot a rhea which they enjoyed eating before Darwin realised that this was the elusive smaller rhea rather than a juvenile, and preserved the head, neck, legs, one wing, and many of the larger feathers. As with his other collections, these were sent to John Stevens Henslow in Cambridge. On 26 January the ''Beagle'' entered the
Straits of Magellan The Strait of Magellan (), also called the Straits of Magellan, is a navigable sea route in southern Chile separating mainland South America to the north and the Tierra del Fuego archipelago to the south. Considered the most important natural ...
, and at St Gregory's Bay Darwin met Patagonians he described as "excellent practical naturalists". A half Indian, who had been born in the Northern Provinces, told him that the smaller rheas were the only species this far south, while the larger rheas kept to the north. On an expedition up the Santa Cruz River, they saw several of the smaller rheas, which were too wary to be approached closely or caught. In 1837, Darwin's rhea was described as ''Rhea darwinii'' (later synonymized with ''R. pennata'') by the ornithologist
John Gould John Gould (; 14 September 1804 – 3 February 1881) was an English ornithologist who published monographs on birds, illustrated by plates produced by his wife, Elizabeth Gould (illustrator), Elizabeth Gould, and several other artists, includ ...
in a presentation to the
Zoological Society of London The Zoological Society of London (ZSL) is a charity and organization devoted to the worldwide animal conservation, conservation of animals and their habitat conservation, habitats. It was founded in 1826. Since 1828, it has maintained London Zo ...
, in which he was followed by Darwin reading a paper on the eggs and distribution of the two species of rheas.Darwin, C (1837) When Gould classified Darwin's rhea and the greater rhea as separate species, he confirmed a serious problem for Darwin. These birds mainly live in different parts of Patagonia, but there is also an overlapping zone where the two species coexist. As every living being had been created in a fixed form, as accepted by the science of his time, they could only change their appearance by a perfect adaptation to their way of life, but would still be the same species. But now he had to deal with two different species. This started to form his idea that species were not fixed at all, but that another mechanism might be at work.Herbert, S (1980)


Conservation

Darwin's rhea is categorized as
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
by the
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
. The former southern nominate subspecies remains relatively widespread and locally fairly common. Its range is estimated at . The situation for the two former northern subspecies is more worrying, with their combined population estimated as being possibly as low as in the hundreds. However, they are classified as ''Rhea tarapacensis'' by the IUCN, which regards it as being
near threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
, with the primary threats being hunting, egg-collecting, and fragmentation of its habitat due to conversion to farmland or pastures for cattle-grazing. Patagonia National Park in Chile's
Aysén Region The Aysén del General Carlos Ibáñez del Campo Region (, , '), often shortened to Aysén Region or Aisén,Examples of name usage1, official regional government site refers to the region as "Región de Aysén"., Chile's official meteorological ...
hosts the Centro de Reproducción para la Conservación del Ñandú ('Reproduction Centre for Darwin's rhea Conservation'). The centre is run by Tompkins Conservation with the support of the National Forest Corporation.


Footnotes


Notes


References

* * * * * * * * * * * * * * * * * * {{Authority control Rheidae Ratites Domesticated birds Flightless birds Birds of the Altiplano Birds of Patagonia Birds of Argentina Birds of Peru Birds of Chile Birds of Bolivia Darwin's rhea Taxa named by Alcide d'Orbigny