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''Corydalis cava'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Papaveraceae The Papaveraceae are an economically important family of about 42 genera and approximately 775 known species of flowering plants in the order Ranunculales, informally known as the poppy family. The family is cosmopolitan, occurring in temperat ...
,
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to moist, shady, woodland habitats throughout most of mainland Europe, although commonest in central and southeast Europe. Its range extends from Spain in the west to Ukraine, Belarus and the Caucasus in the east and as far north as Sweden. It is absent from (though may sometimes be found in a naturalised state in) Iceland, the UK, the Netherlands, Norway, Finland, Russia and Greece.


Description

''Corydalis cava'' grows to tall. It is a spring ephemeral—foliage that grows in the spring dies down to its
tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...
ous rootstock in summer. It has long-spurred flowers which appear in spring. The flowers may be mauve, purple, red, or white. The seeds contain an
elaiosome Elaiosomes ( grc, ἔλαιον ''élaion'' "oil" + ''sóma'' "body") are fleshy structures that are attached to the seeds of many plant species. The elaiosome is rich in lipids and proteins, and may be variously shaped. Many plants have elaio ...
that attracts ants, which transport the seeds into their ant colony. This seed transportation is called
myrmecochory Myrmecochory ( (sometimes myrmechory); from grc, μύρμηξ, mýrmēks ("ant") and ''khoreíā'' ("circular dance") is seed dispersal by ants, an ecologically significant ant–plant interaction with worldwide distribution. Most myrmeco ...
.


Toxicity

Many of the species in ''Corydalis'' contain alkaloids such as
canadine (''S'')-Canadine, also known as (''S'')-tetrahydroberberine and xanthopuccine, is a benzylisoquinoline alkaloid (BIA), of the protoberberine structural subgroup, and is present in many plants from the family Papaveraceae, such as '' Corydalis yan ...
and corydaline, which blocks
calcium Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to ...
. The majority of these belong to the isoquinoline alkaloid group. All parts of the plant are alkaloidal but the highest concentrations are present in the hollow root tuber. ''Corydalis cava'' and some other tuberous species contain the
alkaloid Alkaloids are a class of basic, naturally occurring organic compounds that contain at least one nitrogen atom. This group also includes some related compounds with neutral and even weakly acidic properties. Some synthetic compounds of similar ...
bulbocapnine Bulbocapnine is an alkaloid found in ''Corydalis'' (notably the European species Corydalis cava, C. cava) and ''Dicentra'', genera of the plant family Fumariaceae which have caused (notably the American species ''Corydalis caseana'') the fatal poi ...
, which is occasionally used in medicine but for which scientific evidence is lacking in regard to correct dosage and side effects.


Gallery

File:Dymnivka dutá Podkomorské lesy.JPG, Young shoot pushing up through leaf litter. Přírodní park, Podkomorské forest,
South Moravia The South Moravian Region ( cs, Jihomoravský kraj; , ; sk, Juhomoravský kraj) is an administrative unit () of the Czech Republic, located in the south-western part of its historical region of Moravia (an exception is Jobova Lhota which trad ...
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Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
. File:20150322Corydalis cava3.jpg, Developing inflorescence. Lower Lußhart near
Neulußheim Neulußheim is a town in the district of Rhein-Neckar in Baden-Württemberg in Germany, with about 7,100 inhabitants. Larger cities in the surrounding area include Speyer, Mannheim and Heidelberg. It was founded in 1711 at a crossroads by Julius ...
,
Baden-Württemberg Baden-Württemberg (; ), commonly shortened to BW or BaWü, is a German state () in Southwest Germany, east of the Rhine, which forms the southern part of Germany's western border with France. With more than 11.07 million inhabitants across a ...
, Germany. File:Corydalis cava Bulb.jpg, Mature specimen from a wood in
Vicenza Vicenza ( , ; ) is a city in northeastern Italy. It is in the Veneto region at the northern base of the ''Monte Berico'', where it straddles the Bacchiglione River. Vicenza is approximately west of Venice and east of Milan. Vicenza is a th ...
(
Veneto Veneto (, ; vec, Vèneto ) or Venetia is one of the 20 regions of Italy. Its population is about five million, ranking fourth in Italy. The region's capital is Venice while the biggest city is Verona. Veneto was part of the Roman Empire unt ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
) uprooted at flowering time to show large, hollow tuber. File:Corydalis cava illustration (03).jpg, Dutch coloured plate showing anatomical detail (under original Linnaean name ''Fumaria bulbosa'' i.e. "bulbous fumitory"). File:Corydalis cava PID770-1.jpg, Hollow tuber bisected to show central void. Forest floor,
Rak Škocjan Rak Škocjan ( sl, Rakov Škocjan; ) is a valley and a landscape park, part of Inner Carniola Regional Park in southwestern Slovenia. Administratively, it belongs to the settlement of Rakov Škocjan. Rak Škocjan has been protected since 1949 ...
Landscape park,
Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, an ...
. File:Corydalis cava root tuber (01).jpg, Ants gathered in the void in an exposed tuber, seemingly as attracted by odour of tuber as by that of seed elaiosome. File:Corydalis cava plant (16).jpg, Specimen of white-flowered form blooming among ivy leaves. File:Odvas keltike (Corydalis cava).JPG, Dense carpet of flowering plants (both pink and white-flowered forms) on forest floor. File:Corydalis cava plant (19).JPG, Flowering plants thriving in damp conditions on a stream bank. File:Corydalis cava MichaD 2.jpg, Comparison of inflorescences of pink-flowered and white-flowered forms.
Rosenfeld Rosenfeld is a German name meaning "rose field" or "field of roses" It may refer to: Places * Rosenfeld, Germany, a town in Baden-Württemberg * Rosenfeld, Manitoba, a Canadian village in the Rural Municipality of Rhineland * Rosenfeld (Melk), a ...
,
Baden-Württemberg Baden-Württemberg (; ), commonly shortened to BW or BaWü, is a German state () in Southwest Germany, east of the Rhine, which forms the southern part of Germany's western border with France. With more than 11.07 million inhabitants across a ...
, southern Germany. File:CorydalisSolida+Cava.jpg, Comparison of inflorescences of ''Corydalis cava'' (right, with simple bracts) and the closely related ''C. solida'' (left, with digitate bracts). File:Corydalis cava inflorescence (32).jpg, Inflorescence of pink-flowered form, back-lit by bright sunlight, silhouetting structures within flower spurs. File:Corydalis cava 01.jpg, Inflorescence of white-flowered form, back-lit by bright sunlight. File:Corydalis cava inflorescence (18).jpg, Front-lit inflorescence of white-flowered form. File:Corydalis cava inflorescence (36).jpg , Front view of individual flower of white-flowered form. File:Corydalis cava flowers.JPG, Front view of individual flowers of wild plant, Alte aare nr.
Aarberg Aarberg is a historic town and a municipality in the Seeland administrative district in the canton of Bern in Switzerland. Aarberg lies 20 kilometers from Bern above the river Aare. With an area of , Aarberg borders Bargen, Kappelen, Lyss, Rade ...
,
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
. File:Corydalis cava MichaD 1.jpg, Back-lit inflorescence, showing clearly striated venation of bracts. File:Corydalis cava inflorescence (20).jpg, Close-up of flowers of pale pink form in profile. File:Corydalis cava fg01.jpg, Side view of flower showing hole chewed in nectary by nectar-stealing insect. Selzerbrunnen,
Frankfurt Frankfurt, officially Frankfurt am Main (; Hessian: , "Frank ford on the Main"), is the most populous city in the German state of Hesse. Its 791,000 inhabitants as of 2022 make it the fifth-most populous city in Germany. Located on its na ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
. File:Corydalis cava MichaD 3.jpg, Central structure of interior of single flower.
Rosenfeld Rosenfeld is a German name meaning "rose field" or "field of roses" It may refer to: Places * Rosenfeld, Germany, a town in Baden-Württemberg * Rosenfeld, Manitoba, a Canadian village in the Rural Municipality of Rhineland * Rosenfeld (Melk), a ...
,
Baden-Württemberg Baden-Württemberg (; ), commonly shortened to BW or BaWü, is a German state () in Southwest Germany, east of the Rhine, which forms the southern part of Germany's western border with France. With more than 11.07 million inhabitants across a ...
southern Germany. File:Corydalis cava blatt.jpeg, Leaves of specimen growing at Pölchow nr.
Rostock Rostock (), officially the Hanseatic and University City of Rostock (german: link=no, Hanse- und Universitätsstadt Rostock), is the largest city in the German state of Mecklenburg-Vorpommern and lies in the Mecklenburgian part of the state, c ...
,
Mecklenburg-Vorpommern Mecklenburg-Vorpommern (MV; ; nds, Mäkelborg-Vörpommern), also known by its anglicized name Mecklenburg–Western Pomerania, is a state in the north-east of Germany. Of the country's sixteen states, Mecklenburg-Vorpommern ranks 14th in po ...
, Germany. File:Corydalis cava fruit (06).jpg, Infructescence. File:Corydalis cava fruit (04).jpg, Close-up of single pendent fruit with its accompanying bract. File:Compilation of seeds with elaiosomes 3.jpg, Single seed, greatly magnified, showing its
elaiosome Elaiosomes ( grc, ἔλαιον ''élaion'' "oil" + ''sóma'' "body") are fleshy structures that are attached to the seeds of many plant species. The elaiosome is rich in lipids and proteins, and may be variously shaped. Many plants have elaio ...
(= oily, ant-attracting appendage). File:Corydalis cava Pollen 400x.jpg,
Photomicrograph A micrograph or photomicrograph is a photograph or digital image taken through a microscope or similar device to show a magnified image of an object. This is opposed to a macrograph or photomacrograph, an image which is also taken on a mic ...
of pollen grains.


References

{{Taxonbar, from=Q161556 cava Flora of Europe Ephemeral plants