Columella (genus)
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''Columella'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of very small, air-breathing land snails,
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate gastropod
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s in the family
Truncatellinidae Truncatellinidae is a family of minute, air-breathing land snails, terrestrial pulmonate gastropod molluscs or micromollusks in the superfamily Chondrinoidea. MolluscaBase eds. (2020). MolluscaBase. Truncatellinidae Steenberg, 1925. Accessed ...
, the whorl snails and allies.


Taxonomy

''Columella'' is commonly classified within the
Vertiginidae Vertiginidae, common name the whorl snails, is a family of minute, air-breathing land snails, terrestrial pulmonate gastropod molluscs or micromollusks in the superfamily Pupilloidea. Distribution The distribution of the Vertiginidae is in the ...
. However, as shown by Nekola & Coles (2016), the genus is not that closely related to the Vertiginidae, but may rather be affiliated with the
Chondrinidae Chondrinidae is a family of mostly minute air-breathing land snails, terrestrial pulmonate gastropod mollusks in the order Pulmonata. Anatomy In this family, the number of haploid chromosomes lies between 26 and 30 (according to the values i ...
or form a pupilloid family of its own. A systematic revision is pending.


Species

Species within the genus ''Columella'' include: * '' Columella acicularis'' Almuhambetova, 1979 * '' Columella aspera'' Waldén, 1966 * '' Columella columella'' ( G. v. Martens, 1830) * '' Columella edentula'' ( Draparnaud, 1805) - toothless column * '' Columella hartmutnordsiecki'' Schlickum & Geissert, 1980 * '' Columella hasta'' (Hanna, 1911) * '' Columella intermedia'' Schileyko & Almuhambetova, 1984 * '' Columella microspora'' (R.T. Lowe, 1852) * '' Columella nymphaepratensis'' Hlaváč & Pokryszko, 2009 * '' Columella polvonense'' (Pilsbry, 1894) * '' Columella simplex'' (Gould, 1840) * '' Columella talgarica'' Schileyko & Rymzhanov, 2010 * '' Columella wyciski'' Schlickum & Strauch, 1972 ;Species brought into synonymy: * ''Columella inornata'' (Michaud, 1831): synonym of ''Columella edentula'' (Draparnaud, 1805) (junior synonym) * ''Columella ninagongonis'' Pilsbry, 1935: synonym of '' Truncatellina ninagongonis'' (Pilsbry, 1935) (original combination) * ''Columella tridentata'' A. B. Leonard, 1946: synonym of '' Gastrocopta ruidosensis'' (Cockerell, 1899)


References

{{DEFAULTSORT:Columella (Genus) Truncatellinidae