The croaking gourami (''Trichopsis vittata'') is a species of small freshwater
labyrinth fish of the
gourami
Gouramis, or gouramies , are a group of freshwater anabantiform fishes that comprise the family Osphronemidae. The fish are native to Asia—from the Indian Subcontinent to Southeast Asia and northeasterly towards Korea. The name "gourami", of ...
family. They are native to still waters in
Southeast Asia
Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
and are distributed worldwide via the
aquarium trade
Fishkeeping is a popular hobby, practiced by aquarists, concerned with keeping fish in a home aquarium or garden pond. There is also a piscicultural fishkeeping industry, serving as a branch of agriculture.
Origins of fishkeeping
Fish hav ...
. Croaking gouramis are capable of producing a "croaking" noise using their pectoral fins.
General
Croaking gouramis can reach an average size of about 5 centimeters, though some individuals can grow as large as 6 or 7 centimeters. Coloration is highly variable, ranging from pale brown and green to dark purple with black or red spots on the fins. 2-4 brown or black stripes or rows of spots are present on their sides. Median fins have a thin iridescent blue coloration on their edges. The iris of the eye is bright blue or purple. Females tend to be paler than males, have a slightly rounded dorsal fin and a shorter anal fin. Most croaking gouramis live for about 2 years but with proper care can live as long as 5 in an aquarium setting. They are native to stillwater habitats including ponds, canals and paddy fields in
Java
Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mo ...
,
Borneo
Borneo (; id, Kalimantan) is the List of islands by area, third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java Isl ...
,
Sumatra,
Malaya,
Thailand
Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
,
Laos,
Cambodia
Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailan ...
, and
Vietnam
Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making it ...
.
A breeding population is known to exist in a series of drainage ditches in
Palm Beach County
Palm Beach County is a county located in the southeastern part of Florida and lies directly north of Broward County and Miami-Dade County. The county had a population of 1,492,191 as of the 2020 census, making it the third-most populous count ...
,
Florida
Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, a ...
, USA, almost certainly introduced there through the aquarium industry.
As their name suggests, croaking gouramis are capable of producing an audible grunting or chirping noise, accomplished through the use of specialized adaptations of their pectoral fins.
[Ladich, F., et al. (1992)] These noises are produced by both sexes during breeding displays and during the establishment of dominance hierarchies among males. A typical showdown between males consists of each fish circling the other, flaring fins, aggressively darting (though rarely making contact), and producing croaking noises. Well matched individuals may continue this behaviour for several hours at a time.
In aquaria
Croaking gouramis are fairly shy, peaceful fish that do well in most community
aquaria and do not cause problems with other tankmates. Males can be fairly territorial and aggressive with each other during breeding periods, but will not cause harm given a large enough aquarium, and usually males comfortably coexist. They require a tank no smaller than 40 litres (10 US
gallon
The gallon is a unit of volume in imperial units and United States customary units. Three different versions are in current use:
*the imperial gallon (imp gal), defined as , which is or was used in the United Kingdom, Ireland, Canada, Aus ...
s), preferably larger, and each pair of males will need about 20 inches of space to feel comfortable with one another. Croaking gouramis will prowl about all areas of the aquarium, preferring to lurk among reeds and under large leaves close to the water surface. At night, they might be found "sleeping" at the bottom, even resting on the substrate (which is not normally a cause for concern), or hovering motionless at the water surface.
Like all members of the suborder
Anabantoidei
The Anabantoidei are a suborder of anabantiform ray-finned freshwater fish distinguished by their possession of a lung-like labyrinth organ, which enables them to breathe air. The fish in the Anabantoidei suborder are known as anabantoids or la ...
, the croaking gourami can breathe atmospheric oxygen from above the water surface using a specialized
labyrinth organ if necessary. It is important, therefore, that the surface of the water be exposed to fresh air, usually accomplished by keeping them in an open-top tank or using a hood that allows air ventilation. If the tank has good air pumps, this is not always needed, since the air pumps will refresh the air above the water. Very cold air temperatures at the water surface may lead to infections of the labyrinth organ.
The aquarium should be heavily planted and have at least part of the surface shaded by broad leaves or floating plants. Croaking gouramis will become severely stressed in bare tanks without various hiding places. A darker substrate will make them feel comfortable and help show off their subtle colors. Like most gouramis, these fish are susceptible to diseases and infections, so regular water changes are a must. They are tolerant of fairly high
temperature
Temperature is a physical quantity that expresses quantitatively the perceptions of hotness and coldness. Temperature is measured with a thermometer.
Thermometers are calibrated in various temperature scales that historically have relied on ...
s. This can be used to eliminate
fish diseases
Like humans and other animals, fish suffer from diseases and parasites. Fish defences against disease are specific and non-specific. Non-specific defences include skin and scales, as well as the mucus layer secreted by the epidermis that traps ...
such as
ich
Ich may refer to:
* Ich, a German pronoun meaning ''I'', also a Middle English form of ''I''
* The ego, one of the psychic apparatus defined in Sigmund Freud's structural model of the psyche
* ''Ich'' (album), an album by German rapper Sido
* I ...
from the aquarium. Temperatures of 84 °F (29 °C) are easily tolerated though 26 °C seems to be close to optimal. A pH of 6.8 is about right and
peat
Peat (), also known as turf (), is an accumulation of partially decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, moors, or muskegs. The peatland ecosystem covers and is the most efficien ...
filtration is often recommended.
Croaking gouramis should not be kept with large, aggressive fish, but are compatible with other small, peaceful fish as well as with fellow gouramis. They will be targeted by male Bettas so they should not be kept with them. They are very sensitive to noise and the tank should be in a quiet area.
Diet
In the wild, croaking gouramis are mostly insectivorous, feeding on insects and insect larvae, however other food types, such as zooplankton, crustaceans, shrimp meat and occasionally plant matter, are eaten as well. In aquaria, a varied diet is important to their long-term health; standard flake foods along with regular supplements of freeze-dried bloodworms, tubifex worms,
brine shrimp
''Artemia'' is a genus of aquatic crustaceans also known as brine shrimp. It is the only genus in the family Artemiidae. The first historical record of the existence of ''Artemia'' dates back to the first half of the 10th century AD from Urmia L ...
, and some
algae
Algae ( , ; : alga ) are any of a large and diverse group of photosynthetic, eukaryotic organisms. The name is an informal term for a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from ...
-based flakes will provide these fish with proper nutrition. Occasional feedings of live brine shrimp offer the aquarist an opportunity to observe the natural hunting behaviour of croaking gouramis.
Breeding
Sex can be most reliably determined by
candling
Candling is a method used in embryology to study the growth and development of an embryo inside an egg. The method uses a bright light source behind the egg to show details through the shell, and is so called because the original sources of lig ...
but the more common approach is to observe the shape of the dorsal fin: the male's dorsal fin is pointed, while the female's is rounded. The male croaking gourami is a bubblenest builder, creating a small nest from air bubbles and mucous under a leaf. The water level should be reduced to 8 inches during spawning, circulation kept minimal, and the temperature should be approximately 28 °C (82 °F). Spawning occurs under the nest, with the female responding to the male's dance by rolling over, followed by the typical gourami embrace. About 5 to 10 eggs are released in a quick burst. The male will grab the eggs and spit them into the nest, often adding a few more bubbles for good measure. This act may be repeated a dozen times or more, until about 100 eggs are laid. Some large females may lay more than 200. After spawning the female should be moved to a different tank. The male will keep the bubblenest maintained and tend to the eggs and fry, but when the fry are 2–3 days old the male should also be removed. When first hatched, the fry should be fed infusoria, and later, baby brine shrimp and finely ground flakes. Freeze-dried tablets may also be fed to older fry. It is important that the breeding and grow-out tanks be covered and protected from cool drafts, as low air temperatures above the water surface may cause damage to the developing labyrinth organs of the young fish when they begin to take gulps of air.
Notes
References
*
*
* Ladich, F., W. Brtittenger, and H. Kratochvil (1992). ''Significance Of Agonistic Vocalization In The Croaking Gourami''.
* Shafland, P. L. (1996). "Exotic Fishes of Florida-1994". ''Reviews in Fisheries Science'' 4(2).
* Sterba, G. (1983). ''The Aquarium Fish Encyclopedia''. The MIT Press.
{{Taxonbar, from=Q4348592
Croaking gourami
Fishkeeping
Fish of Southeast Asia
Fish of Thailand
Freshwater fish of Indonesia
Freshwater fish of Malaysia
Fish described in 1831
Taxa named by Georges Cuvier