''Baccharis pilularis'', called coyote brush (or bush), chaparral broom, and bush baccharis, is a

shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

in the family

Asteraceae Asteraceae () is a large family (biology), family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the Order (biology), order Asterales. The number of species in Asteraceae is rivaled only by the Orchi ...

native to

California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...

Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...

Washington Washington most commonly refers to: * George Washington (1732–1799), the first president of the United States * Washington (state), a state in the Pacific Northwest of the United States * Washington, D.C., the capital of the United States ** A ...

, and

Baja California Baja California, officially the Free and Sovereign State of Baja California, is a state in Mexico. It is the northwesternmost of the 32 federal entities of Mexico. Before becoming a state in 1952, the area was known as the North Territory of B ...

. There are reports of isolated populations in

New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...

, most likely introduced.

Distribution and habitat

The plants are found in a variety of habitats, from coastal bluffs, oak woodlands, and grasslands, including on hillsides and in canyons, below . Coyote brush is known as a secondary pioneer plant in communities such as

coastal sage scrub Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland plant community of the California coastal sage and chaparral subecoregion, found in coastal California and northwestern coastal Baja California. It is ...

and

chaparral Chaparral ( ) is a shrubland plant plant community, community found primarily in California, southern Oregon, and northern Baja California. It is shaped by a Mediterranean climate (mild wet winters and hot dry summers) and infrequent, high-intens ...

. It does not regenerate under a closed shrub canopy because seedling growth is poor in the shade.

Coast live oak ''Quercus agrifolia'', the California live oak, or coast live oak, is an evergreen live oak native to the California Floristic Province. Live oaks are so-called because they keep living leaves on the tree all year, adding young leaves and sheddi ...

, California bay, ''

Rhus integrifolia ''Rhus integrifolia'', also known as lemonade sumac, lemonade berry, or lemonadeberry, is a shrub to small tree in the sumac genus '' Rhus''. It is native to the Transverse and Peninsular Ranges and the South Coast regions of Southern Califor ...

'', and other shade producing species replace coastal sage scrub and other coyote bush-dominated areas, particularly when there has not been a wildfire or heavy grazing. In California

grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...

s, it comes in late and invades and increases in the absence of fire or grazing. Coyote bush invasion of grasslands is important because it helps the establishment of other coastal sage species. However, establishment of coyote bush can be concerning because it also displaces highly biodiverse grassland habitat that are important to carbon storage and resilient to wildfires. After grassland restoration, coyote bush can be a major concern and plant invader that overtakes grassland habitat, especially if restoration activities are limited and nonperiodic.

Description

The ''Baccharis pilularis'' shrub is generally smaller than in height. Erect plants are generally mixed with (and sometimes grow to become) prostrate plants. Its leaves are smooth, without spines or hairs, and generally sticky to the touch. The stems are prostrate to erect with branches spreading or ascending. The leaves are long and are

entire Entire may refer to: * Entire function, a function that is holomorphic on the whole complex plane * Entire (animal), an indication that an animal is not neutered * Entire (botany) This glossary of botanical terms is a list of definitions o ...

to toothed and oblanceolate to obovate, with three principal veins. The

flower heads A pseudanthium (; : pseudanthia) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, composite flowers ...

are in a leafy

panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...

. The involucres are hemispheric to bell shaped. This species is

dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...

(

pistillate Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ...

and staminate flowers occur on separate plants). Both staminate and

heads are long. Phyllaries are in 4–6 series, ovate, and glabrous. The receptacles are convex to conic and honeycombed. The staminate flowers range from 20–30 and there are 19–43

flowers. This and other ''

Baccharis ''Baccharis'' is a genus of perennial plant, perennials and shrubs in the Asteraceae, aster family (Asteraceae). They are commonly known as baccharises but sometimes referred to as "brooms", because many members have small thin leaves resembling ...

'' species are nectar sources for most of the predatory wasps, native skippers (small butterflies), and native flies in their ranges. ;Subspecies *''Baccharis pilularis'' subsp. ''consanguinea'' (DC.) C.B.Wolf — primarily in coastal chaparral *''Baccharis pilularis'' subsp. ''pilularis'' — sandy coastal bluffs and beaches in California.

Cultivation

''Baccharis pilularis'' is cultivated as an

ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...

, and used frequently in drought tolerant,

native plant In biogeography, a native species is indigenous to a given region or ecosystem if its presence in that region is the result of only local natural evolution (though often popularised as "with no human intervention") during history. The term is equi ...

, and

wildlife garden A wildlife garden (or habitat garden or backyard restoration) is an Biophysical environment, environment created with the purpose to serve as a sustainable haven for surrounding wildlife. Wildlife gardens contain a variety of habitats that cater t ...

s, and in

natural landscaping Natural landscaping, also called native gardening, is the use of native plants including trees, shrubs, groundcover, and grasses which are local to the geographic area of the garden. Benefits Maintenance Natural landscaping is adapted to t ...

and

habitat restoration Ecological restoration, or ecosystem restoration, is the process of assisting the recovery of an ecosystem that has been degraded, damaged, destroyed or transformed. It is distinct from conservation in that it attempts to retroactively repair ...

projects. The

cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...

ground cover Groundcover or ground cover is any plant that grows low over an area of ground, which protects the topsoil from erosion and drought. In a terrestrial ecosystem, the ground cover forms the layer of vegetation below the shrub layer known as the ...

selections have various qualities of height and spread, leaf colors, and textures. The upright forms are useful for hedges and fence lines, and year-round foliage. Coyote brush is usually deer-resistant. The plants are also drought tolerant after maturity, requiring watering once a week until established, and then about once per month during the first summer. They can mature in one to two years. The plants prefer good drainage. Only male plants of ''Baccharis pilularis'' are cultivated for landscaping use. If these are substituted for ''Baccharis pilularis'' subsp. ''consanguinea'' in

ecological restoration Ecological restoration, or ecosystem restoration, is the process of assisting the recovery of an ecosystem that has been degraded, damaged, destroyed or transformed. It is distinct from conservation in that it attempts to retroactively repair ...

, there will not be as much seed set, nor recruitment of new individuals.

Cultivars

Cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...

s, often with the common name "dwarf coyote brush" or "dwarf baccharis" indicating

selections, include: * ''Baccharis pilularis'' 'Pigeon Point'—from Pigeon Point, California coast. * ''Baccharis pilularis'' 'Twin Peaks'—from coast along Sonoma to Monterey Counties. * ''Baccharis pilularis'' 'Santa Ana'Las Pilitas: ''Baccharis pilularis'' ' Santa Ana'
/ref>

References