Clavatula Filograna

''Clavatula'' is a genus of sea snails, marine gastropod mollusks in the family Clavatulidae.

Description

In this genus, the shell is fusiform, with a well-produced spire. The whorls are coronated with tubercles or short spines at the suture;. The aperture is oval. The columellar lip is smooth, arcuate, and callous behind. The outer lip is thin and arcuated, with the anal sinus situated below the sutural coronal. The operculum is semioval with its nucleus about the middle, on the inner side.

The foot is large, short and obtuse behind. The eyes are placed externally near the extremity of the tentacles. The radula contains a small unicuspid central tooth with single sharp laterals (formula :1-1-1).

The species mostly inhabits the west coast of Africa.

Species

Species within the genus ''Clavatula'' include:

* † '' Clavatula agassizi'' (Bellardi, 1847)
* † '' Clavatula agatensis'' Bellardi, 1877
* '' Clavatula ahuiri'' Cossignani & Ardovini, 2014
* '' Clavatula albasulcata'' R. Aiken, 2021
* † '' Clavatula albucianensis'' Bellardi, 1877
* † '' Clavatula anaglypta'' (Millet, 1865)
* † '' Clavatula apenninica'' Bellardi, 1877
* † '' Clavatula apolloniae'' (R. Hoernes & Auinger, 1891)
* †'' Clavatula aralica'' (Luković, 1924) (an extinct species from the Eocene in western Kazakhstan (Tshegan Formation) )
* † '' Clavatula ariejansseni'' Harzhauser, Landau & R. Janssen, 2022
* '' Clavatula asamusiensis'' Nomura & Zinbo, 1940
* †'' Clavatula asperulata'' (Lamarck, 1822)
* † '' Clavatula atatuerki'' Harzhauser, Landau & R. Janssen, 2022
* † '' Clavatula auingeri'' Finlay, 1927
* † '' Clavatula barnabasi'' Kovács, Leél-Őssy & Vicián, 2023
* † '' Clavatula basilica'' Bellardi, 1877
* † '' Clavatula beauforti'' Oostingh, 1938
* † '' Clavatula berauensis'' Beets, 1941
* † '' Clavatula bicarinata'' Bellardi, 1877
* '' Clavatula bimarginata'' (Lamarck, 1822)
* † '' Clavatula boreoromana'' Kautsky, 1925
* † '' Clavatula burdigalensis'' Cossmann, 1895
* '' Clavatula caerulea'' (Weinkauff & Kobelt, 1875)
* † '' Clavatula camillae'' (R. Hoernes & Auinger, 1891)
* †'' Clavatula carinifera'' Grateloup, 1832
* † '' Clavatula circumclusa'' Bellardi, 1877
* '' Clavatula colini'' Von Maltzan, 1883
* † '' Clavatula complanata'' Bellardi, 1877
* †'' Clavatula concatenata'' (Grateloup, 1832)
* '' Clavatula congoensis'' Nolf & Verstraeten, 2008
* † '' Clavatula consimilis'' Bellardi, 1877
* † '' Clavatula coppii'' Bellardi, 1877
* '' Clavatula coronata'' Lamarck, 1801
* '' Clavatula cossignanii'' Ardovini, 2004
* † '' Clavatula danuvii'' Vicián, Z. Kovács & Stein, 2019
* '' Clavatula debilis'' (R.B. Hinds, 1843) (taxon inquirendum)
* '' Clavatula decorata'' Sowerby III, 1916
* † '' Clavatula defossa'' Bellardi, 1877
* † '' Clavatula defrancii'' Bellardi, 1877
* '' Clavatula diadema'' (Kiener, 1840)
* † '' Clavatula djocdjocartae'' (K. Martin, 1885)
* † '' Clavatula dodecapunctata'' Vera-Peláez & Lozano-Francisco, 2001
* † '' Clavatula dujardini'' Peyrot, 1938
* † '' Clavatula eichwaldi'' Bellardi, 1877
* † '' Clavatula excavata'' Bellardi, 1877
* '' Clavatula filograna'' Odhner, 1923
* '' Clavatula flammulata'' Knudsen, 1952
* † '' Clavatula flexicosta'' Bellardi, 1877
* † '' Clavatula francisci'' (Toula, 1901) (accepted > unreplaced junior homonym)
* '' Clavatula gabonensis'' Melvill, 1923
* † '' Clavatula geniculata'' Bellardi, 1877
* † '' Clavatula glaberrima'' J.P.S. Grateloup, 1832
** †'' Clavatula glaberrima praecedens'' (Peyrot, 1931)
* † '' Clavatula gothica'' Bellardi, 1877
* '' Clavatula gracilior'' Sowerby II, 1870
* † '' Clavatula gracilis'' Erünal-Erentöz, 1958
* † '' Clavatula gradata'' (Defrance, 1826)
* '' Clavatula hattenbergeri'' Nolf & Verstraeten, 2008
* '' Clavatula helena'' Bartsch, 1915
* †'' Clavatula helwerdae'' Ceulemans, Van Dingenen & Landau, 2018
* † '' Clavatula heros'' (Mayer, 1858)
* † '' Clavatula hirmetzli'' Z. Kovács & Vicián, 2021
* †'' Clavatula iberoechinata'' Vera-Peláez & Lozano-Francisco, 2001
* '' Clavatula imperialis'' Lamarck, 1816
* † '' Clavatula implexa'' Bellardi, 1877
* † '' Clavatula inedita'' Bellardi, 1877
* † '' Clavatula inornata'' Bellardi, 1877
* † '' Clavatula interrupta'' (Brocchi, 1814) (accepted > unreplaced junior homonym)
* † '' Clavatula irisae'' Harzhauser, Landau & R. Janssen, 2022
* † '' Clavatula isseli'' Bellardi, 1877
* † '' Clavatula istvani'' Kovács, Leél-Őssy & Vicián, 2023
* † '' Clavatula jarzynkae'' Harzhauser, Landau & R. Janssen, 2022
* † '' Clavatula juliae'' (R. Hoernes & Auinger, 1891)
* † '' Clavatula justinae'' (R. Hoernes & Auinger, 1891)
* '' Clavatula knudseni'' Nolf & Verstraeten, 2007
* ''Clavatula kraepelini'' (Strebel, 1914)
* † '' Clavatula labiolirata'' Landau, Harzhauser, İslamoğlu & C. M. Silva, 2013
* † '' Clavatula laciniata'' Bellardi, 1877
* † '' Clavatula landaui'' Vera-Peláez & Lozano-Francisco, 2001
* '' Clavatula lelieuri'' (Récluz, 1851)
* † '' Clavatula ligeriana'' (Peyrot, 1938)
* '' Clavatula malala'' Bozzetti, 2018
* † '' Clavatula manzonii'' Bellardi, 1877
* † '' Clavatula mariae'' (R. Hoernes & Auinger, 1891)
* '' Clavatula martensi'' Von Maltzan, 1883
* † '' Clavatula martinelli'' Vera-Peláez & Lozano-Francisco, 2001
* '' Clavatula matthiasi'' Nolf, 2008
* ''Clavatula milleti'' (Petit de la Saussaye, 1851)
* '' Clavatula muricata'' (Lamarck, 1822)
* '' Clavatula mystica'' (Reeve, 1843)
* '' Clavatula nathaliae'' Nolf, 2006
* † '' Clavatula neogradata'' Glibert, 1954
* † '' Clavatula nodosa'' Bellardi, 1877
* † '' Clavatula obtruta'' (Millet, 1865)
* † '' Clavatula olgae'' (R. Hoernes & Auinger, 1891)
* † '' Clavatula oostinghi'' Beets, 1942
* † '' Clavatula orientoromana'' Báldi, 1960
* † '' Clavatula palatina'' Strausz, 1954
* † '' Clavatula pettkoi'' Biskupič, 2023
* '' Clavatula petzyae'' Boyer & Ryall, 2006
* † '' Clavatula pusilla'' Bellardi, 1877
* † '' Clavatula pyramidum'' Cuvillier, 1933
* '' Clavatula regia'' (Röding, 1798)
* †'' Clavatula reginae'' Hoernes & Auinger, 1891
* '' Clavatula retusa'' R.B. Hinds, 1843 (taxon inquirendum)
* † '' Clavatula romana'' (Defrance, 1826)
* '' Clavatula rubrifasciata'' (Reeve, 1845)
* † '' Clavatula rugata'' Bellardi, 1877
* † '' Clavatula ruida'' Bellardi, 1877
* † '' Clavatula rusticula'' (Mayer-Eymar, 1886)
* '' Clavatula santhai'' Z. Kovács & Vicián, 2021 †
* † '' Clavatula sceauxensis'' Landau, Van Dingenen & Ceulemans, 2020
* † '' Clavatula seminuda'' Bellardi, 1877
* † '' Clavatula seyithasanensis'' Landau, Harzhauser, İslamoğlu & C. M. Silva, 2013
* '' Clavatula smithi'' Knudsen, 1952
* '' Clavatula solangeae'' Bozzetti, 2006
* † '' Clavatula sophiae'' (R. Hoernes & Auinger, 1891)
* † '' Clavatula sorini'' Harzhauser, Landau & R. Janssen, 2022
* † '' Clavatula sotterii'' (Michelotti, 1847)
* '' Clavatula strebeli'' Knudsen, 1952
* '' Clavatula taxea'' (Röding, 1798)
* † '' Clavatula theodori'' (Toula, 1901)
* '' Clavatula tripartita'' (Weinkauff & Kobelt, 1876)
* † '' Clavatula turriculata'' (Grateloup, 1846)
* † '' Clavatula turriculoides'' Bellardi, 1877
* † '' Clavatula unicostata'' Bellardi, 1877
* † '' Clavatula veronicae'' (R. Hoernes & Auinger, 1891)
* '' Clavatula virgineus'' (Dillwyn, 1817)
* † '' Clavatula vitalisi'' Strausz, 1955
* '' Clavatula xanteni'' Nolf & Verstraeten, 2006

;Nomen dubium:
* ''Clavatula erecta'' W. H. Turton, 1932

;Species brought into synonymy :
* ''Clavatula albicans'' Hinds, 1843: synonym of ''Splendrillia albicans'' (Hinds, 1843)
* ''Clavatula amabilis'' Hinds, 1843: synonym of '' Tritonoturris amabilis'' (Hinds, 1843)
* ''Clavatula argillacea'' Hinds, 1843: synonym of '' Etrema argillacea'' (Hinds, 1843)
* ''Clavatula auriculifera'' Lamarck, J.B.P.A. de M. 1816: synonym of '' Clavus canalicularis'' (Röding, 1798)
* ''Clavatula bella'' Hinds, 1843: synonym of '' Bellacythara bella'' (Hinds, 1843)
* ''Clavatula boreointerrupta'' Kautsky, 1925: synonym of † '' Granulatocincta boreointerrupta'' (Kautsky, 1925)
*'' Clavatula candida'' Hinds, 1843: synonym of '' Glyphostoma candida'' (Hinds, 1843)
* ''Clavatula candida'' (Philippi, 1848): synonym of ''Pusionella valida'' (Dunker, 1852)
* ''Clavatula christianae'' Nolf, 2011: synonym of '' Paraclavatula christianae'' (Nolf, 2011) (original combination)
*'' Clavatula coelata'' Hinds, 1843: synonym of '' Crassispira coelata'' (Hinds, 1843)
* ''Clavatula crenularis'' Lamarck, 1816: synonym of '' Ptychobela nodulosa'' (Gmelin, 1791)
* ''Clavatula delphinae'' Nolf, 2008: synonym of '' Paraclavatula delphinae'' (Nolf, 2008) (original combination)
* ''Clavatula echinata'' Lamarck, 1816: synonym of '' Clavus flammulatus'' Montfort, 1810
* ''Clavatula ericea'' Hinds, 1843: synonym of '' Lioglyphostoma ericea'' (Hinds, 1843)
* ''Clavatula fimbriata'' Hinds, 1843: synonym of '' Daphnellopsis fimbriata'' (Hinds, 1843)
* ''Clavatula flammea'' Hinds, 1843: synonym of '' Daphnella flammea'' (Hinds, 1843)
* ''Clavatula flavidula'' (Lamarck, 1822): synonym of '' Clathrodrillia flavidula'' (Lamarck, 1822)
* ''Clavatula fulva'' Hinds, 1843: synonym of '' Clavus fulva'' (Hinds, 1843)
* ''Clavatula gravis'' (Hinds, 1843): synonym of '' Makiyamaia gravis'' (Hinds, 1843)
* ''Clavatula griffithii'' Gray, 1834: synonym of '' Ptychobela griffithii'' (Gray, 1834)
* ''Clavatula haliplex'' Bartsch, 1915: synonym of '' Toxiclionella haliplex'' (Bartsch, 1915)
* ''Clavatula halistrepta'' Bartsch, 1915: synonym of'' Clionella halistrepta'' (Bartsch, 1915)
* ''Clavatula hottentota'' (E. A. Smith, 1882): synonym of '' Clavus hottentotus'' (E. A. Smith, 1882)
* ''Clavatula impressa'' Hinds, 1843: synonym of '' Kylix impressa'' (Hinds, 1843)
* ''Clavatula kraussii'' (E. A. Smith, 1877): synonym of '' Clionella kraussii'' (E. A. Smith, 1877)
* ''Clavatula laeta'' Hinds, 1843: synonym of '' Splendrillia laeta'' (Hinds, 1843)
* ''Clavatula lineata'' Lamarck, 1816: synonym of '' Perrona lineata'' (Lamarck, 1816)
* ''Clavatula lobatopsis'' Barnard, 1963: synonym of '' Clionella lobatopsis'' (Barnard, 1963)
* ''Clavatula luctuosa'' (Hinds, 1843): synonym of '' Crassispira kluthi'' E. K. Jordan, 1936
* ''Clavatula marmarina'' (Watson, 1881): synonym of '' Fenimorea marmarina'' (Watson, 1881)
* ''Clavatula merita'' Hinds, 1843: synonym of '' Tenaturris merita'' (Hinds, 1843)
* ''Clavatula metula'' Hinds, 1843: synonym of '' Anarithma metula'' (Hinds, 1843)
* ''Clavatula micans'' Hinds, 1843: synonym of '' Globidrillia micans'' (Hinds, 1843)
* ''Clavatula neglecta'' Hinds, 1843: synonym of '' Glyphostoma neglecta'' (Hinds, 1843)
* ''Clavatula nitens'' Hinds, 1843: synonym of '' Agladrillia nitens'' (Hinds, 1843)
* ''Clavatula obesa'' Reeve: synonym of '' Perrona obesa'' (Reeve, 1842)
* ''Clavatula occata'' Hinds, 1843: synonym of '' Diptychophlia occata'' (Hinds, 1843)
* ''Clavatula pardalis'' Hinds, 1843: synonym of '' Anachis pardalis'' (Hinds, 1843)
* ''Clavatula parilis'' E. A. Smith, 1902: synonym of ''Clavatula tripartita'' (Weinkauff, 1876)
* ''Clavatula perronii'' (Reeve, 1843): synonym of '' Perrona perron'' (Gmelin, 1791)
* ''Clavatula pfefferi'' (Strebel, 1912): synonym of '' Fusiturris pfefferi'' (Strebel, 1912)
* ''Clavatula plumbea'' Hinds, 1843: synonym of '' Kurtziella plumbea'' (Hinds, 1843)
* ''Clavatula pseudomystica'' Nolf, 2008: synonym of '' Paraclavatula pseudomystica'' (Nolf, 2008) (original combination)
* ''Clavatula pudica'' Hinds, 1843: synonym of '' Agladrillia pudica'' (Hinds, 1843)
* ''Clavatula pungens'' Gould, 1860: synonym of '' Veprecula pungens'' (Gould, 1860)
* ''Clavatula pyramis'' Hinds, 1843: synonym of '' Pseudorhaphitoma pyramis'' (Hinds, 1843)
* ''Clavatula quinteni'' Nolf & Verstraeten, 2006: synonym of '' Perrona quinteni'' (Nolf & Verstraeten, 2006)
* ''Clavatula quisqualis'' Hinds, 1843: synonym of '' Leptadrillia quisqualis'' (Hinds, 1843)
* ''Clavatula rava'' Hinds, 1843: synonym of '' Clathurella rava'' (Hinds, 1843)
* ''Clavatula rigida'' Hinds, 1843: synonym of '' Clathurella rigida'' (Hinds, 1843)
* ''Clavatula robusta'' Hinds, 1843: synonym of '' Cheungbeia robusta'' (Hinds, 1843)
* ''Clavatula rubida'' Hinds, 1843: synonym of '' Lienardia rubida'' (Hinds, 1843)
* ''Clavatula rubiginosa'' Hinds, 1843: synonym of'' Otitoma rubiginosa'' (Hinds, 1843)
* ''Clavatula rubrofasciata'' Reeve: synonym of '' Clavatula rubrifasciata'' (Reeve, 1845)
* ''Clavatula sacerdos'' (Reeve, 1845): synonym of '' Clavatula mystica'' (Reeve, 1843)
* ''Clavatula scalaris'' Hinds, 1843: synonym of '' Tritonoturris scalaris'' (Hinds, 1843)
* ''Clavatula sculpta'' Hinds, 1843: synonym of '' Rimosodaphnella sculpta'' (Hinds, 1843)
* ''Clavatula semicostata'' (Kiener, 1840): synonym of '' Clionella semicostata'' (Kiener, 1840)
* ''Clavatula sinensis'' Hinds, 1843: synonym of '' Crassispira sinensis'' (Hinds, 1843)
* ''Clavatula sinuata'' (Born, 1778): synonym of '' Clionella sinuata'' (Born, 1778)
* ''Clavatula spicata'' Hinds, 1843: synonym of '' Inquisitor spicata'' (Hinds, 1843)
* ''Clavatula spirata'' Lamarck: synonym of '' Perrona spirata'' (Lamarck, 1816)
* ''Clavatula spurca'' Hinds, 1843: synonym of '' Etrema spurca'' (Hinds, 1843)
* ''Clavatula striata'' Gray, 1826: synonym of '' Epideira striata'' (Gray, 1826)
* ''Clavatula subspirata'' (von Martens, 1902): synonym of '' Perrona subspirata'' (von Martens, 1902)
* ''Clavatula subventricosa'' (E. A. Smith, 1877): synonym of '' Clionella subventricosa'' (E. A. Smith, 1877)
* ''Clavatula taxus'' : synonym of ''Clavatula taxea'' (Röding, 1798) (incorrect subsequent spelling)
* ''Clavatula tessellata'' Hinds, 1843: synonym of '' Kermia tessellata'' (Hinds, 1843)
* ''Clavatula tumida'' Sowerby II, 1870: synonym of '' Toxiclionella tumida'' (Sowerby II, 1870)
* ''Clavatula turriplana'' (G.B. Sowerby III, 1903): synonym of '' Turricula turriplana'' (Sowerby III, 1903)

References

External links

{{Taxonbar, from=Q3722726