In
biological classification, class () is a
taxonomic rank
In biology, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of or ...
, as well as a taxonomic unit, a
taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are
domain,
kingdom,
phylum,
order,
family,
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
, and
species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
, with class ranking between phylum and order.
History
The class as a distinct rank of biological classification having its own distinctive name – and not just called a ''top-level genus'' ''(genus summum)'' – was first introduced by
French botanist Joseph Pitton de Tournefort in the classification of plants that appeared in his ''
Eléments de botanique'' of 1694.
Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine a distinct ''grade'' of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct ''type'' of construction, which is to say a particular layout of organ systems. This said, the composition of each class is ultimately determined by the subjective judgment of
taxonomists.
In the first edition of his ''
Systema Naturae'' (1735),
Carl Linnaeus
Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
divided all three of his
kingdoms of nature (
minerals,
plant
Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
s, and
animal
Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
s) into classes. Only in the animal kingdom are Linnaeus's classes similar to the classes used today; his classes and orders of plants were never intended to represent natural groups, but rather to provide a
convenient "artificial key" according to his ''
Systema Sexuale'', largely based on the arrangement of flowers. In botany, classes are now rarely discussed. Since the first publication of the
APG system in 1998, which proposed a taxonomy of the
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s up to the level of orders, many sources have preferred to treat ranks higher than orders as informal
clades. Where formal ranks have been assigned, the ranks have been reduced to a very much lower level, e.g. class Equisitopsida for the land plants, with the major divisions within the class assigned to subclasses and superorders.
The class was considered the highest level of the taxonomic hierarchy until
George Cuvier's ''embranchements'', first called
Phyla by
Ernst Haeckel,
[Collins, A.G., Valentine, J.W. (2001)]
"Defining phyla: evolutionary pathways to metazoan body plans"
. ''Evol. Dev''. 3: 432–442. were introduced in the early nineteenth century.
See also
*
Cladistics
*
List of animal classes
*
Phylogenetics
*
Systematics
Systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: phylogenetic trees, phylogenies). Phy ...
*
Taxonomy
Explanatory notes
References
{{Authority control
Bacterial nomenclature
Zoological nomenclature
Class
Plant taxonomy