The chalaza (; ; : chalazas or chalazae ) is a structure inside

bird egg Bird eggs are laid by the females and range in quantity from one (as in condors) to up to seventeen (the grey partridge). Avian clutch size, Clutch size may vary latitudinally within a species. Some birds lay eggs even when the eggs have not been ...

s and plant

ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the sporangium, megasporangium), ...

s. It attaches or suspends the

yolk Among animals which produce eggs, the yolk (; also known as the vitellus) is the nutrient-bearing portion of the egg whose primary function is to supply food for the development of the embryo. Some types of egg contain no yolk, for example bec ...

nucellus In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fe ...

within the larger structure.

In animals

In the eggs of most birds (though not in eggs of reptiles), the chalazae are two spiral bands of tissue that suspend the yolk in the center of the white (the albumen). The function of the chalazae is to hold the yolk in place. In baking, the chalazae are sometimes removed in order to ensure a uniform texture.

In plants

In plant ovules, the chalaza is located opposite the micropyle opening of the integuments. It is the tissue where the

integument In biology, an integument is the tissue surrounding an organism's body or an organ within, such as skin, a husk, Exoskeleton, shell, germ or Peel (fruit), rind. Etymology The term is derived from ''integumentum'', which is Latin for "a coverin ...

s and

are joined. Nutrients from the plant travel through vascular tissue in the funiculus and outer integument through the chalaza into the nucellus. During the development of the embryo sac inside a

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...

ovule, the three cells at the chalazal end become the antipodal cells. Embryosac-en

Chalazogamy

In most flowering plants, the

pollen tube A pollen tube is a tubular structure produced by the male gametophyte of seed plants when it germinates. Pollen tube elongation is an integral stage in the plant life cycle. The pollen tube acts as a conduit to transport the male gamete cells fr ...

enters the ovule through the micropyle opening in the integuments for fertilization (porogamy). In chalazogamous fertilization, the

s penetrate the ovule through the chalaza rather than the micropyle opening. Chalazogamy was first discovered in

monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system comparable with gynomonoecy, andromonoecy and trimonoecy, and contras ...

plant species of the family

Casuarinaceae The Casuarinaceae are a family of dicotyledonous flowering plants placed in the order Fagales, consisting of four genera and 91 species of trees and shrubs native to eastern Africa, Australia, Southeast Asia, Malesia, Papuasia, and the Pacific ...

by Melchior Treub, but has since then also been observed in others, for example in

pistachio The pistachio (, ; ''Pistacia vera'') is a small to medium-sized tree of the Anacardiaceae, cashew family, originating in Iran. The tree produces nut (fruit)#Culinary definition and uses, seeds that are widely consumed as food. In 2022, world ...

and

walnut A walnut is the edible seed of any tree of the genus '' Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. They are accessory fruit because the outer covering of the fruit is technically an i ...

Notes

Oology Plant morphology Plant anatomy Pollination {{Plant-morphology-stub