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The Coscinodiscophyceae are a proposed class(s) of
diatom A diatom (Neo-Latin ''diatoma'') is any member of a large group comprising several Genus, genera of algae, specifically microalgae, found in the oceans, waterways and soils of the world. Living diatoms make up a significant portion of Earth's B ...
s. They are similar to the Centrales, a traditional,
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
subdivision of the
heterokont The stramenopiles, also called heterokonts, are protists distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, an ...
algae Algae ( , ; : alga ) is an informal term for any organisms of a large and diverse group of photosynthesis, photosynthetic organisms that are not plants, and includes species from multiple distinct clades. Such organisms range from unicellular ...
known as
diatom A diatom (Neo-Latin ''diatoma'') is any member of a large group comprising several Genus, genera of algae, specifically microalgae, found in the oceans, waterways and soils of the world. Living diatoms make up a significant portion of Earth's B ...
s. The order is named for the shape of the
cell walls A cell wall is a structural layer that surrounds some cell types, found immediately outside the cell membrane. It can be tough, flexible, and sometimes rigid. Primarily, it provides the cell with structural support, shape, protection, and functi ...
(or valves or frustules) of centric diatoms, which are circular or ellipsoid in valve view. The valves often bear
radially symmetrical Symmetry in biology refers to the symmetry observed in organisms, including plants, animals, fungi, and bacteria. External symmetry can be easily seen by just looking at an organism. For example, the face of a human being has a plane of symme ...
ornamental patterns that can appear as dots when viewed with an
optical microscope The optical microscope, also referred to as a light microscope, is a type of microscope that commonly uses visible light and a system of lenses to generate magnified images of small objects. Optical microscopes are the oldest design of micros ...
. Some also bear spines on their valves, which may either increase cell surface area and reduce sinking, or act as a deterrent to
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
grazers. Unlike pennate diatoms, centric diatoms never have a raphe. In terms of
cell cycle The cell cycle, or cell-division cycle, is the sequential series of events that take place in a cell (biology), cell that causes it to divide into two daughter cells. These events include the growth of the cell, duplication of its DNA (DNA re ...
, vegetative cells are
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, ...
and undergo
mitosis Mitosis () is a part of the cell cycle in eukaryote, eukaryotic cells in which replicated chromosomes are separated into two new Cell nucleus, nuclei. Cell division by mitosis is an equational division which gives rise to genetically identic ...
during normal
cell division Cell division is the process by which a parent cell (biology), cell divides into two daughter cells. Cell division usually occurs as part of a larger cell cycle in which the cell grows and replicates its chromosome(s) before dividing. In eukar ...
. In sexual species,
oogamous Oogamy is a form of anisogamy where the gametes differ in both size and form. In oogamy the large female gamete (also known as ovum) is immotile, while the small male gamete (also known as spermatozoon) is mobile. Oogamy is a common form of anis ...
meiosis Meiosis () is a special type of cell division of germ cells in sexually-reproducing organisms that produces the gametes, the sperm or egg cells. It involves two rounds of division that ultimately result in four cells, each with only one c ...
produces
haploid Ploidy () is the number of complete sets of chromosomes in a cell (biology), cell, and hence the number of possible alleles for Autosome, autosomal and Pseudoautosomal region, pseudoautosomal genes. Here ''sets of chromosomes'' refers to the num ...
gamete A gamete ( ) is a Ploidy#Haploid and monoploid, haploid cell that fuses with another haploid cell during fertilization in organisms that Sexual reproduction, reproduce sexually. Gametes are an organism's reproductive cells, also referred to as s ...
s, either
ova , abbreviated as OVA and sometimes as OAV (original animation video), are Japanese animated films and special episodes of a series made specially for release in home video formats without prior showings on television or in theaters, though the ...
or
sperm cell Sperm (: sperm or sperms) is the male reproductive cell, or gamete, in anisogamous forms of sexual reproduction (forms in which there is a larger, female reproductive cell and a smaller, male one). Animals produce motile sperm with a tail k ...
s. These fuse to produce a
zygote A zygote (; , ) is a eukaryote, eukaryotic cell (biology), cell formed by a fertilization event between two gametes. The zygote's genome is a combination of the DNA in each gamete, and contains all of the genetic information of a new individ ...
which expands in size to develop into an
auxospore Auxospores are specialised cells in diatoms that are produced at key stages in their cell cycle or life history. Auxospores typically play a role in growth processes, sexual reproduction or dormancy.Hoek, C. van den, Mann, D. G. and Jahns, H. M. ...
from which full-sized vegetative cells are produced. In some
taxonomy image:Hierarchical clustering diagram.png, 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme o ...
schemes, the centric diatoms are known instead as order Coscinodiscophyceae, and in some schemes as order Biddulphiales. However, diatom taxonomy is changing due to the development of new molecular and genetical analysis tools.


See also

*
Pennales The order Pennales is a traditional subdivision of the heterokont algae known as diatoms. The order is named for the shape of the cell wall#Diatom cell walls, cell walls (or valves or frustules) of pennate diatoms, which are elongated in valve vi ...


References


Further reading


External links

{{Taxonbar, from=Q2698449 Ochrophyte classes