
The cat righting reflex is a
cat's innate ability to orientate itself as it falls in order to land on its feet. The
righting reflex begins to appear at 3–4 weeks of age, and is perfected at 6–9 weeks. Cats are able to do this because they have an unusually flexible
backbone and no functional
clavicle
The clavicle, collarbone, or keybone is a slender, S-shaped long bone approximately long that serves as a strut between the scapula, shoulder blade and the sternum (breastbone). There are two clavicles, one on each side of the body. The clavic ...
(collarbone). The tail seems to help but cats without a tail also have this ability, since a cat mostly turns by moving its legs and twisting its spine in a certain sequence.
While cats provide the most famous example of this reflex, they are not the only animal known to have a mid-air righting capability. Similar phenomenons have been observed in other small vertebrates such as
rabbits,
rats,
lizards
Lizard is the common name used for all squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The ...
, and certain invertebrate tailed
arthropods
Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
(e.g.
stick insects).
Technique

After determining down from up visually or with their
vestibular apparatus (in the
inner ear
The inner ear (internal ear, auris interna) is the innermost part of the vertebrate ear. In vertebrates, the inner ear is mainly responsible for sound detection and balance. In mammals, it consists of the bony labyrinth, a hollow cavity in the ...
), cats twist themselves to face downward. They are able to accomplish this within the physical law of
conservation of angular momentum
Angular momentum (sometimes called moment of momentum or rotational momentum) is the rotational analog of Momentum, linear momentum. It is an important physical quantity because it is a Conservation law, conserved quantity – the total ang ...
with these key steps:
#Bend in the middle so that the front half of their body rotates about a different axis from the rear half.
#Tuck their front legs in to reduce the
moment of inertia of the front half of their body and extend their rear legs to increase the moment of inertia of the rear half of their body so that they can rotate their front by as much as 90° while the rear half rotates in the opposite direction as little as 10°.
#Extend their front legs and tuck their rear legs so that they can rotate their rear half further while their front half rotates in the opposite direction less.
Depending on the cat's flexibility and initial angular momentum, if any, the cat may need to perform steps two and three repeatedly to complete a full 180° rotation.
Terminal velocity
In addition to the righting reflex, cats have other features that reduce damage from a fall. Their small size, light bone structure, and thick fur decrease their
terminal velocity. While falling, a cat spreads out its body to increase drag.
An average-sized cat with its limbs extended achieves a terminal velocity of about , around half that of an average-sized man, who reaches a terminal velocity of about . A 2003 study of feline
high-rise syndrome found that cats "orient
heir
Inheritance is the practice of receiving private property, titles, debts, entitlements, privileges, rights, and obligations upon the death of an individual. The rules of inheritance differ among societies and have changed over time. Offi ...
limbs horizontally after achieving maximum velocity so that the impact is more evenly distributed throughout the body".
Injury
With their righting reflex, cats often land uninjured. However, this is not always the case, since cats can still break bones or die from extreme falls. In a 1987 study, published in the ''Journal of the
American Veterinary Medical Association
The American Veterinary Medical Association (AVMA) is an American not-for-profit association founded in 1863 that represents more than 105,000 veterinarians.
The AVMA provides information resources, continuing education opportunities, publicat ...
'', of 132 cats that were brought into the
New York Animal Medical Center after having fallen from buildings, it was found that injuries per cat increased positively with altitude until a height of seven stories, at which point injuries decreased. One cat fell 40 stories without injury, having apparently bounced off a canopy and into a planter.
The study's authors speculated that, after falling five stories, the cats reached terminal velocity, at which point they relaxed and spread their bodies out to increase
drag. However, critics of the study have questioned the conclusion that mortality rates decrease as height increases due to
survivorship bias
Survivorship bias or survival bias is the logical error of concentrating on entities that passed a selection process while overlooking those that did not. This can lead to incorrect conclusions because of incomplete data.
Survivorship bias is ...
; falls that resulted in instant death were not included as a deceased cat would not be brought to a vet.
[ A 2003 study of 119 cats concluded that "Falls from the seventh or higher stories, are associated with more severe injuries and with a higher incidence of thoracic trauma."]
See also
* Falling cat problem
The falling cat problem is a problem that consists of explaining the underlying physics behind the observation of the cat righting reflex.
Although amusing and trivial to pose, the solution of the problem is not as straightforward as its state ...
– the mathematical problem of explaining the physics of the cat righting reflex
* High-rise syndrome – veterinary terminology for injuries sustained by cats typically caused by falls from significant heights
* Buttered cat paradox – a humorous combination of two observations, the cat righting reflex and the buttered toast phenomenon
References
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Further reading
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External links
The surprisingly complicated physics of why cats always land on their feet
National Geographic video on the cat righting reflex
Slow Motion Flipping Cat Physics
{{Cat nav
Cat behavior
Reflexes
Articles containing video clips