Carpinus

Hornbeams are hardwood trees in the plant genus ''Carpinus'' in the family Betulaceae. Its species occur across much of the temperate regions of the Northern Hemisphere.

Common names

The common English name ''hornbeam'' derives from the hardness of the woods (likened to horn) and the Old English ''beam'', "tree" (cognate with Dutch ''Boom'' and German ''Baum'').

The American hornbeam is also occasionally known as blue-beech, ironwood, or musclewood, the first from the resemblance of the bark to that of the American beech ''Fagus grandifolia'', the other two from the hardness of the wood and the muscled appearance of the trunk and limbs.

The botanical name for the genus, ''Carpinus'', is the original Latin name for the European species, although some etymologists derive it from the Celtic for a yoke.

Description

Hornbeams are small, slow-growing, understory trees with a natural, rounded form growing tall and wide; the exemplar species—the European hornbeam—reaches a maximum height of .

Leaves are deciduous, dark-green, alternate and simple with a coarsely-serrated margin, varying from in length. In autumn, leaves turn various shades of yellow, orange and red. Hornbeam saplings, stressed trees, and the lower branches of mature trees may exhibit marcescence—where leaves wither with autumn but abscission (leafdrop) is delayed until spring.

The smooth, gray trunk and larger branches of a mature tree exhibit a distinctive muscle-like fluting.

As with other members of the birch family, hornbeam flowers are wind-pollinated pendulous catkins, produced in spring. Male and female flowers are on separate catkins, but on the same tree (monoecious). Female flowers give way to distinctive clusters of winged seeds that somewhat resemble the hops-like seeds of ironwood.

The fruit is a small nut about long, held in a leafy bract; the bract may be either trilobed or simple oval, and is slightly asymmetrical. The asymmetry of the seedwing makes it spin as it falls, improving wind dispersal. The shape of the wing is important in the identification of different hornbeam species. Typically, 10–30 seeds are on each seed catkin.

Taxonomy

Formerly some taxonomists segregated them with the genera '' Corylus'' ( hazels) and ''Ostrya'' ( hop-hornbeams) in a separate family, Corylaceae. Modern botanists place ''Carpinus'' in the subfamily Coryloideae of the family Betulaceae. Species of ''Carpinus'' are often grouped into two subgenera ''Carpinus'' subgenus ''Carpinus'' and ''Carpinus'' subgenus '' Distegicarpus''.

Phylogentic analyses have shown that ''Ostrya'' likely evolved from a ''Carpinus'' ancestor somewhere in ''C.'' subg. '' Distegicarpus'' making ''Carpinus'' paraphyletic. The fossil record of the genus extends back to the Early Eocene, Ypresian of northwestern North America, with the species '' Carpinus perryae'' described from fossil fruits found in the Klondike Mountain Formation of Republic, Washington.

Species

43 species are currently accepted.

* '' Carpinus austrobalcanica'' N.Kuzmanović, D.Lakušić, I.Stevanoski, P.Schönswetter, B.Frajman – Southern Albania, Northwestern Greece
*''Carpinus betulus'' – European hornbeam - Europe to Western Asia; naturalized in North America.
*'' Carpinus caroliniana'' – American hornbeam - Eastern North America
*'' Carpinus chuniana'' – Guangdong, Guizhou, Hubei
*'' Carpinus cordata'' – Sawa hornbeam - Primorye, China, Korea, Japan
*'' Carpinus dayongiana'' – Hunan
*'' Carpinus faginea'' – Nepal, Himalayas of northern India
*'' Carpinus fangiana'' – Sichuan, Guangxi
*'' Carpinus fargesiana'' – central and east-central China
*'' Carpinus firmifolia'' – Guizhou: Guiyang Shi
*'' Carpinus gigabracteatus'' – Yunnan
*'' Carpinus hebestroma'' – Taiwan
*'' Carpinus henryana'' – southern China
*'' Carpinus insularis'' – Hong Kong
*'' Carpinus japonica'' – Japanese hornbeam - Japan
*'' Carpinus kawakamii'' – Taiwan, southeastern China
*'' Carpinus kweichowensis'' – Guizhou, Yunnan
*'' Carpinus langaoensis'' – Shaanxi, China
*'' Carpinus laxiflora'' – Aka-shide hornbeam - Japan, Korea
*'' Carpinus lipoensis'' – Guizhou
*'' Carpinus londoniana'' – southern China, northern Indochina
*'' Carpinus luochengensis'' – Guangxi
*'' Carpinus mengshanensis'' – Shandong
*'' Carpinus microphylla'' – Guangxi
*'' Carpinus mollicoma'' – Tibet, Sichuan, Yunnan
*'' Carpinus monbeigiana'' – Tibet, Yunnan
*'' Carpinus omeiensis'' – Sichuan, Guizhou
*'' Carpinus orientalis'' – Oriental hornbeam - Hungary, Balkans, Italy, Crimea, Turkey, Iran, Caucasus
*'' Carpinus paohsingensis'' – China
*†'' Carpinus perryae'' - Ypresian, Klondike Mountain Formation
*'' Carpinus polyneura'' – southern China
*'' Carpinus pubescens'' – China, Vietnam
*'' Carpinus purpurinervis'' – Guizhou, Guangxi
*'' Carpinus putoensis'' – Putuo hornbeam - Zhejiang
*'' Carpinus rankanensis'' – Taiwan
*'' Carpinus rupestris'' – Yunnan, Guangxi, Guizhou
*'' Carpinus × schuschaensis'' (''C. betulus'' × ''C. orientalis'') – Caucasus and northern Iran
*'' Carpinus shensiensis'' – Gansu, Shaanxi
*'' Carpinus shimenensis'' – Hunan
*†'' Carpinus tengshongensis'' – Pliocene
*'' Carpinus tibetana'' – Tibet
*'' Carpinus tientaiensis'' – Zhejiang: Tianmu Shan
*'' Carpinus tropicalis'' – Mexico, Central America
*'' Carpinus tsaiana'' – Yunnan, Guizhou
*'' Carpinus tschonoskii'' – Asian Hornbeam, Chonosuki's Hornbeam - southern China, Korea, Japan
*'' Carpinus turczaninovii'' – Korean hornbeam - China, Korea, Japan
*'' Carpinus viminea'' – China, Korea, Himalayas, northern Indochina

Distribution and habitat

The 43 species occur across much of the temperate regions of the northern hemisphere, with the greatest number of species in east Asia, particularly China. Only three species occur in Europe, only one in eastern North America, and one in Mesoamerica. ''Carpinus betulus'' can be found in Europe, Turkey and Ukraine.

Ecology

Hornbeams are used as food plants by the larvae of some Lepidoptera species, including autumnal moth, common emerald, feathered thorn, walnut sphinx, Svensson's copper underwing, and winter moth (recorded on European hornbeam) as well as the ''Coleophora'' case-bearers ''C. currucipennella'' and ''C. ostryae''.

Uses

Hornbeams yield a very hard timber, giving rise to the name "ironwood".Archived a
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Dried heartwood billets are nearly white and are suitable for decorative use. For general carpentry, hornbeam is rarely used, partly due to the difficulty of working it.

The wood is used to construct carving boards, tool handles, handplane soles, coach wheels, piano actions, shoe lasts, and other products where a very tough, hard wood is required.

The wood can also be used as gear pegs in simple machines, including traditional windmills. It is sometimes coppiced to provide hardwood poles. It is also used in parquet flooring and for making chess pieces.

References

External links

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Taxa described in 1753
Taxa named by Carl Linnaeus