
Carcharhiniformes ( from
Classical Greek
Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archa ...
' (karcharos) 'sharp/jagged' and ' (rhinos) 'nose', plus
Latin
Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
''forme'' 'shape'), commonly known as ground sharks, are the largest order of
shark
Sharks are a group of elasmobranch cartilaginous fish characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the ...
s, with over 270 species. They include a number of common types, such as
catsharks,
swellsharks, and
requiem sharks.
Members of this order are characterized by the presence of a
nictitating membrane
The nictitating membrane (from Latin '' nictare'', to blink) is a transparent or translucent third eyelid present in some animals that can be drawn across the eye from the medial canthus to protect and moisten it while maintaining vision. Most ...
over the eye, two
dorsal fin
A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
s, an
anal fin
Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
, and five
gill
A gill () is a respiration organ, respiratory organ that many aquatic ecosystem, aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow r ...
slits.
The families in the order Carcharhiniformes are expected to be revised; recent
DNA
Deoxyribonucleic acid (; DNA) is a polymer composed of two polynucleotide chains that coil around each other to form a double helix. The polymer carries genetic instructions for the development, functioning, growth and reproduction of al ...
studies show that some of the conventional groups are not
monophyletic
In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria:
# the grouping contains its own most recent co ...
.
The oldest members of the order appeared during the Middle-Late
Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
, which have teeth and body forms that are morphologically similar to living
catsharks.
Carchariniformes first underwent major diversification during the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
, initially as mostly small-sized forms, before radiating into medium and large body sizes during the
Cenozoic
The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three g ...
.
Families
Eschmeyer's Catalog of Fishes
Catalog of Fishes is a comprehensive on-line database and reference work on the scientific names of fish species and genera. It is global in its scope and is hosted by the California Academy of Sciences. It has been compiled and is continuously up ...
classifies the Carcharhiniformes into two suborders and nine families:
*Suborder
Scyliorhinoidei
**Family
Scyliorhinidae
Scyliorhinidae is a Family (biology), family of sharks, one of a few families whose members share the common name catsharks, belonging to the Order (biology), order Carcharhiniformes, the ground sharks. Although they are generally known as cats ...
Gill
A gill () is a respiration organ, respiratory organ that many aquatic ecosystem, aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow r ...
, 1862 (catsharks)
*Suborder
Carcharhinoidei
**Family
Atelomycteridae
Atelomycteridae, the coloured catsharks, is a family of sharks belonging to the order Carcharhiniformes. These sharks are found in the Indian, Pacific and Western Atlantic Oceans.
Genera
Atelomycteridae contains the following genera and specie ...
E. G. White, 1936 (coloured catsharks)
**Family
Pentanchidae Smith, 1912 (deepwater catsharks)
**Family
Dichichthyidae W. T. White, Stewart, Reuben
O'Neill & Naylor, 2024 (bristle sharks)
**Family
Pseudotriakidae Gill, 1893 (false catsharks)
**Family
Proscylliidae
The finback catsharks are a small family (biology), family, the Proscylliidae, of Carcharhiniformes, ground sharks. They can be found in warm seas worldwide and are often the most numerous and common shark in tropical regions. They are generally ...
Fowler, 1941 (finback catsharks)
**Family
Leptochariidae Gray
Grey (more frequent in British English) or gray (more frequent in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning that it has no chroma. It is the color of a cloud-covered s ...
, 1851 (barbeled houndsharks)
**Family
Triakidae Gray, 1851 (houndsharks)
***Subfamily
Triakinae Gray, 1851 (smouth-hounds)
***Subfamily
Galeorhininae Gill, 1862 (school sharks)
**Family
Hemigaleidae J.C.F. Hasse, 1878 (weasel sharks)
**Family
Carcharhinidae D. S. Jordan & Evermann, 1896 (requiem sharks)
**Family
Galeocerdonidae Poey, 1875 (tiger sharks)
**Family
Sphyrnidae Bonaparte, 1840 (hammerhead sharks)
Fossil genera
The following fossil genera of Carcharhiniformes are also known:
* Genus †''
Altusmirus''
Fuchs, Engelbrecht, Lukeneder & Kriwet, 2017
* Genus †''
Corysodon''
Saint-Seine, 1949
* Genus †''
Platyrhizodon''
Guinot, Underwood, Cappetta & Ward, 2013
* Genus †''
Vectiscyllium''
Duffin & Batchelor, 2024
* Family
Carcharhinidae
** Genus †''
Abdounia''
Cappetta, 1980
** Genus †''
Archaeogaleus''
Guinot, Cappetta & Adnet, 2014
** Genus †''
Danogaleus''
Noubhani & Cappetta, 1997
** Genus †''
Eogaleus''
Cappetta, 1975
** Genus †''
Eorhincodon''
Li, 1997
** Genus †''
Fornicatus''
Fuchs, Engelbrecht, Lukeneder & Kriwet, 2018
** Genus †''
Kruckowlamna''
Laurito Mora, 1999
** Genus †''
Misrichthys''
Case & Cappetta, 1990
** Genus †''
Pseudabdounia''
Ebersole, Cicimurri & Stringer, 2019
* Family †
Florenceodontidae Cappetta, Morrison & Adnet, 2019
** Genus †''
Florenceodon''
Cappetta, Morrison & Adnet, 2019
* Family
Hemigaleidae
** Genus †''
Moerigaleus''
Underwood & Ward, 2011
* Family
Galeocerdonidae
** Genus †''
Physogaleus''
Cappetta, 1980
* Family †
Pseudoscyliorhinidae Stumpf, Scheer & Kriwet, 2019
** Genus †''
Diprosopovenator''
Stumpf, Scheer & Kriwet, 2019
** Genus †''
Pseudoscyliorhinus''
Müller & Diedrich, 1991
* Family
Pentanchidae
** Genus †''
Pseudoapristurus''
Pollerspöck & Straube, 2017
* Family
Proscyllidae
** Genus †''
Praeproscyllium''
Underwood & Ward, 2004
* Family
Scyliorhinidae
Scyliorhinidae is a Family (biology), family of sharks, one of a few families whose members share the common name catsharks, belonging to the Order (biology), order Carcharhiniformes, the ground sharks. Although they are generally known as cats ...
** Genus †''
Bavariscyllium''
Thies, 2005
** Genus †''
Cadiera''
Guinot, Cappetta & Adnet, 2014
** Genus †''
Casieria''
Noubhani & Cappetta, 1997
** Genus †''
Crassescyliorhinus''
Underwood & Ward, 2008
** Genus †''
Cretascyliorhinus''
Underwood & Mitchell, 1999
** Genus †''
Foumtizia''
Noubhani & Cappetta, 1997
** Genus †''
Megascyliorhinus''
Cappetta & Ward, 1977
** Genus †''
Microscyliorhinus''
Case, 1994
** Genus †''
Orthodon
The Sacramento blackfish (''Orthodon microlepidotus'') is a species of freshwater ray-finned fish belongingto the family Leuciscidae, which includes the daces, chubs, Eurasian minnows and related fishes. This species is endemic to central Califor ...
''
Coquand, 1860
** Genus †''
Pachyscyllium''
Reinecke, Moths, Grant & Breitkreuz, 2005
** Genus †''
Palaeoscyllium''
Wagner, 1857
** Genus †''
Platyrhizoscyllium''
Adnet, 2006
** Genus †''
Porodermoides''
Noubhani & Cappetta, 1997
** Genus †''
Premontreia''
Cappetta, 1992
** Genus †''
Prohaploblepharus''
Underwood & Ward, 2008
** Genus †''
Protoscyliorhinus''
Herman, 1977
** Genus †''
Pteroscyllium''
Cappetta, 1980
** Genus †''
Scyliorhinotheca''
Kiel, Peckmann & Simon, 2013
** Genus †''
Sigmoscyllium''
Guinot, Underwood, Cappetta & Ward, 2013
** Genus †''
Stenoscyllium''
Noubhani & Cappetta, 1997
** Genus †''
Thiesus''
Guinot, Cappetta & Adnet, 2014
** Genus †''
Tingaleus''
Cappetta, 1993
* Family
Triakidae
** Genus †''
Archaeotriakis''
Case, 1978
** Genus †''
Gomphogaleus''
Adnet & Cappetta, 2008
** Genus †''
Kallodentis''
Engelbrecht, Mörs, Reguero & Kriwet, 2017
** Genus †''
Khouribgaleus''
Noubhani & Cappetta, 1997
** Genus †''
Meridiogaleus''
Engelbrecht, Mörs, Reguero & Kriwet, 2017
** Genus †''
Pachygaleus''
Cappetta, 1992
** Genus †''
Palaeogaleus''
Gurr, 1962
** Genus †''
Palaeotriakis''
Guinot, Underwood, Cappetta & Ward, 2013
** Genus †''
Rhaibodus''
Böhm, 1926
** Genus †''
Squatigaleus''
Cappetta, 1989
** Genus †''
Xystrogaleus''
Adnet, 2006
References
Further reading
* Froese, Rainer, and Daniel Pauly, eds. (2013
Fish Identification: Ground sharksin
FishBase
FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfish on the web. . March 2013 version.
*
External links
Order Carcharhiniformes
{{Authority control
Articles which contain graphical timelines
Cartilaginous fish orders
Taxa named by Leonard Compagno
Extant Albian first appearances