In
mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
ian
oral anatomy, the canine teeth, also called cuspids, dogteeth, eye teeth, vampire teeth, or fangs, are the relatively long, pointed
teeth
A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
. In the context of the upper jaw, they are also known as ''
fangs''. They can appear more flattened, however, causing them to resemble
incisor
Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s and leading them to be called ''incisiform''. They developed and are used primarily for firmly holding food in order to tear it apart, and occasionally as weapons. They are often the largest teeth in a mammal's mouth. Individuals of most species that develop them normally have four, two in the upper jaw and two in the lower, separated within each jaw by incisors; humans and dogs are examples. In most species, canines are the anterior-most teeth in the
maxilla
In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
ry bone.
The four canines in humans are the two upper
maxillary canines and the two lower
mandibular canine
The mandibular canine is the tooth located distally (away from the midline of the face) from both mandibular lateral incisors of the mouth but mesially (toward the midline of the face) from both mandibular first premolars. Both the maxillary and m ...
s.
They are specially prominent in
dog
The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers. ...
s (
Canidae
Canidae (; from Latin, ''canis'', "dog") is a family (biology), biological family of caniform carnivorans, constituting a clade. A member of this family is also called a canid (). The family includes three subfamily, subfamilies: the Caninae, a ...
), hence the name.
Details
There are generally four canine teeth: two in the upper (maxillary) and two in the lower (mandibular) arch. A canine is placed laterally to (outside of) each
lateral
Lateral is a geometric term of location which may also refer to:
Biology and healthcare
* Lateral (anatomy), a term of location meaning "towards the side"
* Lateral cricoarytenoid muscle, an intrinsic muscle of the larynx
* Lateral release ( ...
incisor
Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
and
mesial to (inwards of) the
premolar
The premolars, also called premolar Tooth (human), teeth, or bicuspids, are transitional teeth located between the Canine tooth, canine and Molar (tooth), molar teeth. In humans, there are two premolars per dental terminology#Quadrant, quadrant in ...
s. They are larger and stronger than the incisors, and their roots sink deeply into the bones, causing well-marked prominences upon the surface. They are the only teeth in dentition with a single cusp, their crowns are roughly triangular from a mesial viewpoint and trapezoidal from a
buccal viewpoint.
The crown is large and conical, very convex on its
labial surface, a little hollowed and uneven on its
lingual surface, and tapering to a blunted point or cusp, which projects beyond the level of the other teeth. The root is single, but longer and thicker than that of the incisors, conical in form, compressed laterally, and marked by a slight groove on each side. The lingual surface also presents two depressions on either side of the surface separated by a ridge in between; these depressions are known as mesial and distal lingual fossae.
Eruption
In humans, the upper canine teeth (popularly called eye teeth, from their position under the eyes) are larger and longer than the lower, and usually present a distinct basal ridge. Eruption typically occurs between the ages of eleven and twelve years for upper canines and between nine and ten years for lower canines. Upper
deciduous
In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
canines also known as the baby tooth, typically erupt between the ages of sixteen and twenty-two months and shed between ten and twelve years. Lower deciduous canines typically erupt between the ages of seventeen and twenty-two months and shed between nine and twelve years.
Developmental defects
Transposition (positional interchange of two adjacent teeth) is a development defect that most commonly found in the permanent canine, with the maxillary being more commonly seen than mandibular. The upper canine frequently transposed with the first premolar whilst the lower canine transposed with the lateral incisor. Occasionally canines are congenitally missing.
Maxillary canine morphology
From a facial aspect, maxillary canines are approximately one millimetre narrower than the central incisor. Their mesial aspects resemble the adjacent lateral incisors, while their distal aspects anticipate the first premolars. They are slightly darker and more yellow in color than the other anterior teeth. From a lingual aspect, they have well-developed mesial and distal marginal ridges and a well-developed cingulum. A prominent lingual ridge divides the lingual aspect in half and creates the mesial and distal lingual fossae between the lingual ridge and the marginal ridges. From a proximal aspect, they resemble the incisors, but are more robust, especially in the cingulum region. Incisally, they are visibly asymmetrical, as the mesial incisal edge is slightly shorter than the distal incisal edge, which places the cusp slightly mesial to the long axis of the tooth. They are also thicker
labiolingually than mesiodistally. Because of the disproportionate incisal edges, the contacts are also asymmetrical. Mesially, the contact sits at the junction of the incisal and middle third of the crown, while distally, the contact as more cervical, in the middle of the middle third of the crown. The root of the maxillary canines are the longest root of any tooth and conical in shape.
[Oral Anatomy, Histology and Embryology Barry K. B. Berkovitz, G. R. Holland, and Bernard J. Moxham Chapter 2.]
Mandibular canine morphology
The lower canine teeth are placed nearer the middle line than the upper, so that their summits correspond to the intervals between the upper canines and the lateral incisors. From a facial aspect, the mandibular canine is notably narrower mesiodistally than the maxillary one, the root is compressed mesiodistally with well-marked grooves on both sides and may be just as long as the maxillary (and at times bifurcated).
A distinctive feature is the nearly straight outline this tooth has compared to the maxillary canine which is slightly more bowed. As in the maxillary canine, the mesial incisal edge (or cusp ridge) is shorter than the distal side, however, the cusp is displaced slightly lingual relative to the cusp of the maxillary canine.
Lingually, the surface of the tooth is much smoother compared to the very pronounced surface of the maxillary canine, and the cingulum is noted as less developed. The cusp may be lost with attrition over time and may resemble an upper second permanent incisor.
Sexual dimorphism

In many species the canine teeth in the upper or lower jaw, or in both jaws, are much larger in males than in females, where they are sometimes hidden or completely absent. Animals where this occurs include
antelope
The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...
s,
musk-deer,
camel
A camel (from and () from Ancient Semitic: ''gāmāl'') is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. Camels have long been domesticated and, as livestock, they provid ...
s,
horse
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
s, wild
boar
The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a Suidae, suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The speci ...
, some apes, seals,
narwhal
The narwhal (''Monodon monoceros'') is a species of toothed whale native to the Arctic. It is the only member of the genus ''Monodon'' and one of two living representatives of the family Monodontidae. The narwhal is a stocky cetacean with a ...
, and
walrus
The walrus (''Odobenus rosmarus'') is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. It is the only extant species in the family Odobeni ...
. Male dogs have larger canines with different contour than do females.
Humans have the proportionately smallest male canine teeth among all anthropoids and exhibit relatively little sexual dimorphism in canine tooth size. It has been proposed that the receding canine teeth in human males was likely to be a result of
sexual selection
Sexual selection is a mechanism of evolution in which members of one sex mate choice, choose mates of the other sex to mating, mate with (intersexual selection), and compete with members of the same sex for access to members of the opposite sex ...
for less aggressive partners by female humans. The dimorphism is also similarly less pronounced in
bonobo
The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee (less often the dwarf chimpanzee or gracile chimpanzee), is an endangered great ape and one of the two species making up the genus ''Pan (genus), Pan'' (the other bei ...
s and
chimpanzee
The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
s.
Non-synapsids
In non-
synapsid
Synapsida is a diverse group of tetrapod vertebrates that includes all mammals and their extinct relatives. It is one of the two major clades of the group Amniota, the other being the more diverse group Sauropsida (which includes all extant rept ...
s, such as reptiles and crocodiles, teeth similar to canines may be termed "caniniform" ("canine-shaped") teeth. Teeth or appendages with similar appearances found in many snakes and invertebrates, such as spiders, are referred to as
fangs, but are usually modified to inject venom.
[''The Teeth of Non-Mammalian Vertebrates: Form, Function, Development and Growth'' by Barry Berkovitz, Peter Shellis – Academic Press 2023 Page 335--355]
Additional images
File:3D Medical Animation Still Showing Types of Teeth.jpg, Medical animation showing Canine teeth and their arrangement in the mouth of an adult human being.
File:Illu mouth.jpg, Mouth (oral cavity)
File:Gray157.png, Left maxilla. Outer surface.
File:Gray187.png, Base of skull. Inferior surface.
File:Gray1001.png, Unerupted permanent teeth underlie the deciduous teeth
Deciduous teeth or primary teeth, also informally known as baby teeth, milk teeth, or temporary teeth,Fehrenbach, MJ and Popowics, T. (2026). ''Illustrated Dental Embryology, Histology, and Anatomy'', 6th edition, Elsevier, page 287–296. are ...
.
See also
*
Carnassial
Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified f ...
*
Incisor
Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
*
Molar
*
Premolar
The premolars, also called premolar Tooth (human), teeth, or bicuspids, are transitional teeth located between the Canine tooth, canine and Molar (tooth), molar teeth. In humans, there are two premolars per dental terminology#Quadrant, quadrant in ...
References
External links
*
{{Authority control
Teeth
Mammal anatomy
Types of teeth