''Caecosphaeroma'' is a
troglodytic isopod
Isopoda is an Order (biology), order of crustaceans. Members of this group are called isopods and include both Aquatic animal, aquatic species and Terrestrial animal, terrestrial species such as woodlice. All have rigid, segmented exoskeletons ...
genus in the family
Sphaeromatidae found in
cave
Caves or caverns are natural voids under the Earth's Planetary surface, surface. Caves often form by the weathering of rock and often extend deep underground. Exogene caves are smaller openings that extend a relatively short distance undergrou ...
s of NE and SW France. The genus was split off from
Monolistra by
Adrien Dollfus in 1896; in both genera, the female carries about 10 fertilized eggs in its external
marsupium (brood pouch); they are white in ''Monolistra'' but bluish-green in ''Caecosphaeroma''. ''C. burgundum'' is the most studied species.
Description

They measure from 2–20 mm long. As cave dwellers, they have lost their vision, but remain sensitive to light, which they shun. They are capable of
volvation (rolling themselves into a ball) to protect themselves, rest, or sleep. During copulation the male and female embrace takes the form of two concentric spheres.
Development
Larva remain in the marsupium about 12 months,
and the animals continue growing for several years, reaching a final length of 10–20 mm.
Evolution
The marine ancestors of ''Caecosphaeroma'' are believed to have migrated up the course of rivers and adapted to a subterranean environment that was relatively stable; meanwhile the marine environment of their ancestors changed, so ''Caecosphaeroma'' continued to differentiate as a separate lineage.
Volvation in ''C. burgundum''
''C. burgundum'' has evolved in ways to improve its ability to curl into a ball. The mandibular palps and antennae are able retract into two deep grooves of the face, and the posterior margin of
pleotelson
The telson () is the hindmost division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on account of ...
(the last body segment fused with the "tail") provides greater mechanical support for the head. Evolution of
volvation is seen in other subterranean isopods, but only ''C. burgundum'' is able to roll up into a hermetic sphere without any outward projections, and thus "approaches perfection in volvation".
[ (Article has English abstract.)]
References
{{Taxonbar, from=Q10439158
Sphaeromatidae
Isopod genera
Cave crustaceans
Fauna of France