''Baurutitan'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

that lived during the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

in what is now

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

, ''Baurutitan britoi'', was described in 2005 by Kellner and colleagues, although the fossil remains had already been discovered in 1957. ''Baurutitan'' is classified as a

lithostrotia Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' an ...

titanosaur Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still th ...

, and is distinguished from related genera based on its distinctive caudal

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

e. This South American dinosaur was found in the

Serra da Galga Formation The Serra da Galga Formation is a geological formation in Minas Gerais state of southeastern Brazil. Its strata date back to the Maastrichtian, and are part of the Bauru Group.It was originally considered a member of the Marília Formation. Dinos ...

near

Uberaba Uberaba () is a city in the state of Minas Gerais, southeast Brazil. It is located in the Brazilian Highlands at above sea level on the Uberaba River, and away from the state capital, Belo Horizonte. The city status was granted in 1856, and i ...

, in the Brazilian state of

Minas Gerais Minas Gerais () is a state in Southeastern Brazil. It ranks as the second most populous, the third by gross domestic product (GDP), and the fourth largest by area in the country. The state's capital and largest city, Belo Horizonte (literally ...

Discovery

The holotype of ''Baurutitan'' were found in 1957 by

Llewellyn Ivor Price Llewellyn Ivor Price (October 9, 1905 – June 9, 1980) was one of the first Brazilian paleontologists. His work contributed not only to the development of Brazilian but also to global paleontology. He collected '' Staurikosaurus'', the first ...

, the famous Brazilian paleontologist, in the region of Peirópolis,

. However, it was not until 2005 that ''Baurutitan'' was officially published and named. The works of Price in Peirópolis began in 1947 after Jesuíno Felicíssimo Junior, from the Instituto Geográfico e Geológico of São Paulo, told him about the presence of fossils in the region. Price then conducted excavations in an old quarry, known as Quarry Caieira, on the in São Luis farm. Recovered fossils included those of

turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked tu ...

crocodylomorphs Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, cr ...

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

s and

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

, freshwater invertebrates (

gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...

s and

bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...

s),

trace fossils A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, ...

, fragments of

eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...

, and plant debris. Dynamite was occasionally used to remove materials embedded in the matrix. There were expeditions to Caieira between 1949 and 1961, after which fieldwork was not continued due to a lack of new relevant findings. A series of 19 vertebrae (the last sacral and 18 caudals) found at this locality were recognized as belong to those of a titanosaur; they were initially named 'Series "C"', and subsequently became the holotype of the new genus ''Baurutitan'' (specimen number MCT 1490-R). All material found was stored in the collection of Museu de Ciência da Terra (Earth Science Museum – MCT) of the Departamento Nacional de Produção Mineral (National Department of Mineral Production) in

Rio de Janeiro Rio de Janeiro ( , , ; literally 'River of January'), or simply Rio, is the capital of the state of the same name, Brazil's third-most populous state, and the second-most populous city in Brazil, after São Paulo. Listed by the GaWC as a b ...

. In addition to Caieira Quarry, Price also worked in the "Point 6" or "Rodovia" site, also from the

, about 1.5 km east of Peirópolis. Field work during the 1980s and 1990s discovered many titanosaur remains, including vertebrae and limb bones. A preliminary report found the cervical and dorsal vertebrae from the site similar to ''Trigonosaurus,'' while the caudal vertebrae were similar to ''Baurutitan.'' In 2022, the remains were described, and were found to share characteristics with both ''Baurutitan'' and ''Trigonosaurus''. This led the describing authors to synonymize the two taxa, while separating the unique paratype of ''Trigonosaurus'' as the new genus '' Caieiria.''

Etymology

The name ''Baurutitan'' comes from the junction of the word "bauru", which alludes to the geographical region of the finding, within the

Bauru Group The Bauru Group is a geological group of the Bauru Sub-basin, Paraná Basin in Minas Gerais, São Paulo, General Salgado, Itapecuru-Mirim, Mato Grosso, Brazil whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils ...

, and the word "titan" of Greek myths. The specific name "britoi" is given in honor of Ignácio Aureliano Machado Brito, a Brazilian

paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

who advised the study and description of this

, whose fossils have remained stored for many years.

Description

''Baurutitan'' was a

which was estimated to have measured long and had a height of around . The holotype specimen was found in the Marilia Formation, dating to the

Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...

epoch, some 72-66 million years ago. ''Baurutitan'' was the fourth sauropod described in

, after ''

Antarctosaurus ''Antarctosaurus'' (; meaning "southern lizard") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. The type species, ''Antarctosaurus wichmannianus'', and a second species, ''Antarctosau ...

'', ''

Gondwanatitan ''Gondwanatitan'' (meaning "giant from Gondwana") was a titanosaurian sauropod dinosaur. ''Gondwanatitan'' was found in Brazil, at the time part of the southern supercontinent Gondwana, in the late Cretaceous Period (70 mya). Like some other ...

'' and '' Amazonsaurus''. A 2017 cladistic analysis by José Carballido and colleagues places ''Baurutitan'' as a member of Lithostrotia.José L. Carballido; Diego Pol; Alejandro Otero; Ignacio A. Cerda; Leonardo Salgado ; Alberto C. Garrido ; Jahandar Ramezani ; Néstor R. Cúneo ; Javier M. Krause (2017). "A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs". Proceedings of the Royal Society B: Biological Sciences. 284 (1860): 20171219. doi:10.1098/rspb.2017.1219.

References

External links

''Baurutitan'' on the Dinosaur Mailing List
{{Taxonbar, from=Q134648 Titanosaurs Maastrichtian life Late Cretaceous dinosaurs of South America Cretaceous Brazil Fossils of Brazil Marília Formation Fossil taxa described in 2005 Taxa named by Alexander Kellner