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Burchell's zebra (''Equus quagga burchellii'') is a southern
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of the
plains zebra The plains zebra (''Equus quagga'', formerly ''Equus burchellii'') is the most common and geographically widespread species of zebra. Its range is fragmented, but spans much of southern and eastern Africa south of the Sahara. Six or seven subspec ...
. It is named after the British explorer and naturalist
William John Burchell __NOTOC__ William John Burchell (23 July 1781 – 23 March 1863) was an English explorer, naturalist, traveller, artist, and author. His thousands of plant specimens, as well as field journals from his South African expedition, are held by Kew ...
. Common names include bontequagga, Damaraland zebra, and Zululand zebra (
Gray Grey (more frequent in British English) or gray (more frequent in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning that it has no chroma. It is the color of a cloud-covered s ...
, 1824). Burchell's zebra is the only subspecies of zebra which may be legally farmed for human consumption.


Physical characteristics

Like most plains zebras, females and males are about the same size, standing at the shoulder. They weigh between . Year-round reproduction observed in this subspecies in
Etosha National Park Etosha National Park is a national park in northwestern Namibia and one of the largest national parks in Africa. It was proclaimed a game reserve in March 1907 in Ordinance 88 by the Governor of German South West Africa, Friedrich von Lindequist. ...
, Namibia, concludes synchronization of a time budget between males and females, possibly explaining the lack of
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
. Burchell's zebras are described as being striped on the head, the neck, and the flanks, and sparsely down the upper segments of the limbs then fading to white. One or two shadow stripes rest between the bold, broad stripes on the haunch. This main distinguishing characteristic sets the Burchell's zebra apart from the other subspecies. Gray (1824) observed a distinct
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...
line, the tail only bristly at the end, and the body distinctly white. The dorsal line is narrow and becomes gradually broader toward the rear, distinctly margined with white on each side.


Behaviour

Like most plains zebras, Burchell's live in small family groups. These can be either harem or bachelor groups, with harem groups consisting of one stallion and one to six mares and their most recent foals, and bachelor groups containing two to eight unattached stallions. The males in bachelor herds are often the younger or older stallions of the population, as they are most likely not experienced enough or strong enough to defend breeding rights to a group of females from challengers. These small groups often congregate in larger herds around water and food sources, but still maintain their identity as family units while in the population gatherings.


Range and adaptation

Formerly, the Burchell's zebra range was centred north of the Vaal/Orange river system, extending northwest via southern
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
to Etosha and the Kaokoveld, and southeast to
Eswatini Eswatini, formally the Kingdom of Eswatini, also known by its former official names Swaziland and the Kingdom of Swaziland, is a landlocked country in Southern Africa. It is bordered by South Africa on all sides except the northeast, where i ...
and
KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
. Now extirpated in the middle portion, it survives at the northwestern and southeastern ends of the distribution. Burchell's zebra migrates the longest distance of any terrestrial animal in Africa, making a round trip of . They migrate from the
Chobe River The Cuando (or Kwando) is a river in south-central Africa flowing through Angola and Namibia's Caprivi Strip and into the Linyanti Swamp on the northern border of Botswana. Below the swamp, the river is called the Linyanti River and, farther east ...
in
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
to Nxai Pan National Park in Botswana. Their migration follows a straight north–south route almost entirely within the Kavango–Zambezi Transfrontier Conservation Area (KAZA).


Controversial introductions outside its historical range

From 2001 until 2016 the Kissama Foundation reintroduced wildlife in the Quiçama National Park of Angola. The project was dubbed Operation Noah's Ark. Amongst animals such as
blue wildebeest The blue wildebeest (''Connochaetes taurinus''), also called the common wildebeest, white-bearded gnu or brindled gnu, is a large antelope and one of the two species of wildebeest. It is placed in the genus ''Connochaetes'' and Family (biology), ...
,
waterbuck The waterbuck (''Kobus ellipsiprymnus'') is a large antelope found widely in sub-Saharan Africa. It is placed in the genus ''Kobus (antelope), Kobus'' of the family Bovidae. It was first Scientific description, described by Irish naturalist Will ...
, Cape giraffe, bush elephants, gemsbok, eland,
nyala The lowland nyala or simply nyala (''Tragelaphus angasii'') is a spiral-horned artiodactyl antelope native to Southern Africa. The species is part of the family Bovidae and the genus '' Tragelaphus'' (formerly placed in the genus ''Nyala''). It ...
and
ostrich Ostriches are large flightless birds. Two living species are recognised, the common ostrich, native to large parts of sub-Saharan Africa, and the Somali ostrich, native to the Horn of Africa. They are the heaviest and largest living birds, w ...
were also Burchell's zebras. And from 2017 until 2019 Wildlife Vets Namibia exported wildlife to the Democratic Republic of the Congo's capital city Kinshasa to introduce animals into the Parc de la Vallée de la Nsele. Wildlife Vets Namibia in partnership with Institut Congolais pour la Conservation de la Nature also established a Burchell's zebra population on Île de Mateba, which is originally a rainforest island. Both introductions in west Angola and west DRC are controversial since the park service bodies from both countries did not opt to obtain the native Grant's zebra from for example countries such as Zambia, Tanzania or Kenya.


Extinct subpopulation

Like other plains zebras, Burchell's zebras must have populated the African plains in impressive numbers. Associations of thousands have been reported. The wild herds were thought to have disappeared by 1910, and the last known captive individual died in the
Berlin Zoo The Berlin Zoological Garden (, ) is the oldest surviving and best-known zoo in Germany. Opened in 1844, it covers and is located in Berlin's Tiergarten. With about 1,380 different species and over 20,200 animals, the zoo presents one of the ...
in 1918. As European settlement spread northward from the Cape to colonial southern
Rhodesia Rhodesia ( , ; ), officially the Republic of Rhodesia from 1970, was an unrecognised state, unrecognised state in Southern Africa that existed from 1965 to 1979. Rhodesia served as the ''de facto'' Succession of states, successor state to the ...
, this subspecies was thought to have been hunted to extinction. However, Groves and Bell concluded in their 2004 publication that "the extinct true Burchell's zebra" is a phantom. Careful study of the original zebra populations in Zululand and
Eswatini Eswatini, formally the Kingdom of Eswatini, also known by its former official names Swaziland and the Kingdom of Swaziland, is a landlocked country in Southern Africa. It is bordered by South Africa on all sides except the northeast, where i ...
, and of skins harvested on game farms in Zululand and Natal, has revealed that a certain small proportion shows similarity to what now is regarded as typical ''burchellii''. The type localities of the two subspecies ''Equus quagga burchellii'' (Burchell's zebra) and ''Equus quagga antiquorum'' (Damaraland zebra) are so close to each other that they suggest that the two are in fact one, and therefore the older of the two names should take precedence over the younger. They therefore say that the correct name for the southernmost subspecies must be ''burchellii'', not ''antiquorum''. The subspecies ''Equus quagga burchellii'' still exists in
KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
and in Etosha. ''Equus quagga burchellii'' can be found in a number of zoos in the United States, including the Cincinnati Zoo,
Columbus Zoo The Columbus Zoo and Aquarium is a non-profit zoo located near Powell, Ohio, Powell in Liberty Township, Delaware County, Ohio, United States, north of the city of Columbus, Ohio, Columbus. The land lies along the eastern banks of the O'Shaughnes ...
, Naples Zoo, Nashville Zoo, Woodland Park Zoo.


References


Sources

* Duncan, P. (ed.). 1992. ''Zebras, Asses, and Horses: An Action Plan for the Conservation of Wild Equids''. IUCN/SSC Equid Specialist Group. IUCN, Gland, Switzerland. * Maas, P. 2005
"Burchell's Zebra – ''Equus quagga burchellii''"The Extinction Website
Downloaded on 21 January 2006. * Moehlman, P.D. 2002. Equids
Zebras, Asses and Horses. Status Survey and Conservation Action Plan
IUCN/SSC Equid Specialist Group. IUCN, Gland, Switzerland.


External links

* {{Authority control Zebras Mammals described in 1824 Taxa named by John Edward Gray Mammals of Angola Mammals of Botswana Mammals of Lesotho Mammals of Namibia Mammals of South Africa Mammals of Eswatini