
The annual growth cycle of grapevines is the process that takes place in the vineyard each year, beginning with
bud
In botany, a bud is an undeveloped or Plant embryogenesis, embryonic Shoot (botany), shoot and normally occurs in the axil of a leaf or at the tip of a Plant stem, stem. Once formed, a bud may remain for some time in a dormancy, dormant conditi ...
break in the spring and culminating in
leaf fall in autumn followed by winter
dormancy
Dormancy is a period in an organism's Biological life cycle, life cycle when growth, development, and (in animals) physical activity are temporarily stopped. This minimizes metabolism, metabolic activity and therefore helps an organism to conserv ...
. From a
winemaking
Winemaking, wine-making, or vinification is the production of wine, starting with the selection of the fruit, its Ethanol fermentation, fermentation into alcohol, and the bottling of the finished liquid. The history of wine-making stretches over ...
perspective, each step in the process plays a vital role in the development of
grape
A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters.
The cultivation of grapes began approximately 8,0 ...
s with ideal characteristics for making
wine
Wine is an alcoholic drink made from Fermentation in winemaking, fermented fruit. Yeast in winemaking, Yeast consumes the sugar in the fruit and converts it to ethanol and carbon dioxide, releasing heat in the process. Wine is most often made f ...
.
Viticulturalists and vineyard managers monitor the effect of climate,
vine disease and pests in facilitating or impeding the
vine's progression from bud break,
flowering,
fruit
In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering.
Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
set,
veraison,
harvesting, leaf fall and dormancy – reacting if need be with the use of viticultural practices like
canopy management,
irrigation
Irrigation (also referred to as watering of plants) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has bee ...
,
vine training and the use of
agrochemicals
An agrochemical or agrichemical, a contraction of ''agricultural chemical'', is a chemical product used in industrial agriculture. Agrichemical typically refers to biocides (pesticides including insecticides, herbicides, fungicides and nematicid ...
. The stages of the annual growth cycle usually become observable within the first year of a vine's life. The amount of time spent at each stage of the growth cycle depends on a number of factors – most notably the type of climate (warm or cool) and the characteristics of the
grape variety
This list of grape varieties includes cultivated grapes, whether used for wine, or eating as a table grape, fresh or dried (raisin, Zante currant, currant, sultana (grape), sultana). For a complete list of all grape species, including those unimp ...
.
[J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 741–742 Oxford University Press 2006 ]
Bud break

The grape starts its annual growth cycle in the spring with bud break. In the Northern Hemisphere, this stage begins around March while in the
Southern Hemisphere it begins around September when daily temperatures begin to surpass . If the vine had been
pruned during the winter, the start of this cycle is signaled by a "bleeding" of the vine. This bleeding occurs when the soil begins to warm and
osmotic forces pushes water, containing a low concentration of
organic acid
An organic acid is an organic compound with acidic properties. The most common organic acids are the carboxylic acids, whose acidity is associated with their carboxyl group –COOH. Sulfonic acids, containing the group –SO2OH, are re ...
s,
hormones
A hormone (from the Greek participle , "setting in motion") is a class of signaling molecules in multicellular organisms that are sent to distant organs or tissues by complex biological processes to regulate physiology and behavior. Hormones a ...
,
mineral
In geology and mineralogy, a mineral or mineral species is, broadly speaking, a solid substance with a fairly well-defined chemical composition and a specific crystal structure that occurs naturally in pure form.John P. Rafferty, ed. (2011): Mi ...
s and sugars, up from the
root system
In mathematics, a root system is a configuration of vector space, vectors in a Euclidean space satisfying certain geometrical properties. The concept is fundamental in the theory of Lie groups and Lie algebras, especially the classification and ...
of the vine and it is expelled from the cuts (or "wounds") left over from pruning the vine. During this period a single vine can "bleed" up to of water.
[J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 82 Oxford University Press 2006 ]
Tiny buds on the vine start to swell and eventually
shoots begin to grow from the buds. Buds are the small part of the vine that rest between the vine's
stem and the
petiole (leaf stem). Inside the buds contain usually three primordial shoots. These buds appear in the summer of previous growth cycle green and covered in scales. During winter dormancy they turn brown until the spring when the vine begins the process of bud break and the first sign of green in the vineyard emerges in the form of tiny shoots.
[J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 107 Oxford University Press 2006 ] The energy to facilitate this growth comes from reserves of
carbohydrate
A carbohydrate () is a biomolecule composed of carbon (C), hydrogen (H), and oxygen (O) atoms. The typical hydrogen-to-oxygen atomic ratio is 2:1, analogous to that of water, and is represented by the empirical formula (where ''m'' and ''n'' ...
stored in roots and wood of the vine from the last growth cycle. Eventually the shoots sprout tiny leaves that can begin the process of
photosynthesis
Photosynthesis ( ) is a system of biological processes by which photosynthetic organisms, such as most plants, algae, and cyanobacteria, convert light energy, typically from sunlight, into the chemical energy necessary to fuel their metabo ...
, producing the energy to accelerate growth. In warm climates, after about 4 weeks the growth of the shoots starts to rapidly accelerate with the shoots growing in length an average of a day.
In
temperate climate
In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ra ...
s, where temperatures can reach above in mid-winter, some early budding varieties (such as
Chardonnay
Chardonnay (, ; ) is a green-skinned grape variety used in the production of white wine. The variety originated in the Burgundy wine region of eastern France, but is now grown wherever wine is produced, from England to New Zealand. For new a ...
) can be at risk of premature bud break. This is a potential viticultural hazard in places like the
Margaret River region of Western Australia where warm currents from the Indian Ocean can coax Chardonnay vines to prematurely bud in the mid-winter month of July. After bud break, the young shoots are very vulnerable to
frost
Frost is a thin layer of ice on a solid surface, which forms from water vapor that deposits onto a freezing surface. Frost forms when the air contains more water vapor than it can normally hold at a specific temperature. The process is simila ...
damage with vineyard managers going to great lengths to protect the fragile shoots should temperatures dramatically drop below freezing. This can include setting up heaters or wind circulators in the vineyard to keep cold air from settling on the vines.
Flowering

Depending on temperatures, 40–80 days after bud break the process of flowering begins with small flower clusters resembling buttons appearing on the tips of the young shoots. Flowering occurs when average daily temperatures stay between , which in the Northern Hemisphere wine regions is generally around May and for the Southern Hemisphere regions around November. A few weeks after the initial clusters appear, the flowers start to grow in size with individual flowers becoming observable.
It is during this stage of flowering that the
pollination
Pollination is the transfer of pollen from an anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds. Pollinating agents can be animals such as insects, for example bees, beetles or bu ...
and
fertilization
Fertilisation or fertilization (see American and British English spelling differences#-ise, -ize (-isation, -ization), spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give ...
of the grapevine takes place with the resulting product being a grape berry, containing 1–4
seeds.
[J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 276 Oxford University Press 2006 ]

Most cultivated ''
Vitis vinifera
''Vitis vinifera'', the common grape vine, is a species of flowering plant, native to the Mediterranean Basin, Mediterranean region, Central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern ...
'' grape vines are
hermaphroditic, with both male
stamen
The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10
Morphology and terminology
A stamen typically consists of a stalk called the filament ...
s and female
ovaries
The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are endocr ...
, while many wild grapes are either male, producing pollen but no fruit, or female, producing fruit only if a pollinator is nearby.
Hermaphroditic vines are preferred for cultivation because each vine is more likely to
self-pollinate and produce fruit.
At the beginning of the flowering process the only part that is visible is the fused cap of petals known as the
calyptra. Shortly after the calyptra is shed, liberating the
pollen
Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
from the
anther
The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10
Morphology and terminology
A stamen typically consists of a stalk called the filament ...
s of the stamen. Wind and insects generally play only a small role in aiding pollination, with the process being mostly self-contained within the vine. But
cross-pollination between vines of different varieties is possible:
Cabernet Sauvignon is a cross of
Cabernet Franc
Cabernet Franc is one of the major black grape varieties worldwide. It is principally grown for blending with Cabernet Sauvignon and Merlot in the Bordeaux (wine), Bordeaux style, but can also be vinified alone, as in the Loire (wine), Loire's C ...
and
Sauvignon blanc;
Petite Sirah is a cross of
Syrah
Syrah (), also known as Shiraz, is a dark-skinned grape variety grown throughout the world and used primarily to produce red wine. In 1999, Syrah was found to be the offspring of two obscure grapes from southeastern France, Dureza and Mondeuse ...
and
Peloursin. During the process of fertilization, the pollen fertilizes the ovary which produces seeds as the flower begins the transformation into a grape berry, encapsulating the seed. Detrimental weather (cold, wind & rain) can severely affect the flowering process, causing many flowers not to be fertilized and produce a group.
It is during this time that the buds that will become next year's crops begin to form.
Fruit set

The stage of fruit set follows flowering almost immediately, when the fertilized flower begins to develop a seed and grape berry to protect the seed. In the Northern Hemisphere, this normally takes place in May and in the Southern Hemisphere in November.
This stage is very critical for wine production since it determines the potential crop
yield. Not every flower on the vine gets fertilized, with the unfertilized flowers eventually falling off the vine. The percentage of fertilized flowers averages around 30 but can get as high as 60 or be much lower. Climate and the health of the vine play an important role with low
humidity
Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation (meteorology), precipitation, dew, or fog t ...
, high temperatures and water stress having the potential of severely reducing the percentage of flowers that get fertilized.
Coulure
Coulure (pronounced coo-LYUR) is a viticultural hazard that is the result of metabolic reactions to weather conditions that causes a failure of grapes to develop after flowering. In English the word ''shatter'' is sometimes used. Coulure is tri ...
occurs when there is an imbalance of carbohydrate levels in the vine tissues and some berries fail to set or simply fall off the bunch. Varieties like
Grenache and
Malbec are prone to this abnormal fruit set.
Millerandage occurs when some fertilized flowers do not form seeds but only small berry clusters. Grape berry size depends on the number of seeds so berries with no seeds will be significantly smaller than berries containing seeds. On one cluster there may be berries of various sizes which can create problems during winemaking due to the varying "skin to pulp" ratio among the grapes.
[J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 291 Oxford University Press 2006 ] This can be caused by vine disease, such as
fanleaf, or by a
boron
Boron is a chemical element; it has symbol B and atomic number 5. In its crystalline form it is a brittle, dark, lustrous metalloid; in its amorphous form it is a brown powder. As the lightest element of the boron group it has three ...
deficiency in the vine.
Gewürztraminer
Gewürztraminer () is an aromatic wine grape variety, used in white wines, and which performs best in cooler climates. In English, it is sometimes referred to colloquially as Gewürz ( ; although this is never the case in German, because mean ...
and the Chardonnay
clones IA and Mendoza are both prone to millerandage.
[J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 443 Oxford University Press 2006 ]
Veraison
Following fruit set, the grape berries are green and hard to the touch. They have very little
sugar
Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose
Glucose is a sugar with the Chemical formula#Molecular formula, molecul ...
and are high in organic acids. They begin to grow to about half their final size when they enter the stage of veraison. This stage signals the beginning of the
ripening
Ripening is a process in fruits that causes them to become more palatable. In general, fruit becomes sweeter, less green, and softer as it ripens. Even though the acidity of fruit increases as it ripens, the higher acidity level does not make t ...
process and normally takes place around 40–50 days after fruit set. In the Northern Hemisphere this will be around the end of July and into August and between the end of January into February for the Southern Hemisphere.
During this stage the colors of the grape take form—red/black or yellow/green depending on the grape varieties. This color changing is due to the
chlorophyll
Chlorophyll is any of several related green pigments found in cyanobacteria and in the chloroplasts of algae and plants. Its name is derived from the Greek words (, "pale green") and (, "leaf"). Chlorophyll allows plants to absorb energy ...
in the berry skin being replaced by
anthocyanins
Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart named a chemical compound that gives flow ...
(red wine grapes) and
carotenoids
Carotenoids () are yellow, orange, and red organic compound, organic pigments that are produced by plants and algae, as well as several bacteria, archaea, and Fungus, fungi. Carotenoids give the characteristic color to pumpkins, carrots, parsnips ...
(white wine grapes). In a process known as
engustment, the berries start to soften as they build up
sugars
Sugar is the generic name for Sweetness, sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose, fructose, and galactose. Compound sugars, also called disaccharides ...
. Within six days of the start of veraison, the berries begin to grow dramatically as they accumulate
glucose
Glucose is a sugar with the Chemical formula#Molecular formula, molecular formula , which is often abbreviated as Glc. It is overall the most abundant monosaccharide, a subcategory of carbohydrates. It is mainly made by plants and most algae d ...
and
fructose
Fructose (), or fruit sugar, is a Ketose, ketonic monosaccharide, simple sugar found in many plants, where it is often bonded to glucose to form the disaccharide sucrose. It is one of the three dietary monosaccharides, along with glucose and gal ...
and acids begin to fall.
[J. Robinson (ed) ''"The Oxford Companion to Wine"'' Third Edition pg 730 Oxford University Press 2006 ]
The onset of veraison does not occur uniformly among all berries. Typically the berries and clusters that are most exposed to warmth, on the outer extents of the
canopy, undergo veraison first with the berries and clusters closer to the trunk and under the canopy shade undergoing it last. There are some factors in the vineyards that can control the onset of veraison, limited
water stress and canopy management that creates a high "fruit to leaf" ratio can encourage veraison. This is because the vine is biologically programmed to channel all its energies and resources into the berries, which houses its seedling offspring, to provide them a better chance of survival. Conversely, very vigorous vines with plentiful leaf shading for photosynthesis and water supply will delay the start of veraison due to the vines energies being directed towards continued shoot growth of new buds. For the production of high-quality wine, it is considered ideal to have an earlier veraison. During this period the cane of the vine starts to ripen as well changing from green and springing to brown and hard. The vines begins to divert some of its energy production into its reserves in preparation for its next growth cycle.
After harvest

In the vineyard, the antepenultimate event is the
harvest
Harvesting is the process of collecting plants, animals, or fish (as well as fungi) as food, especially the process of gathering mature crops, and "the harvest" also refers to the collected crops. Reaping is the cutting of grain or pulses fo ...
in which the grapes are removed from the vine and transported to the winery to begin the wine making process. In the Northern Hemisphere this is generally between September and October while in the Southern Hemisphere it is generally between February and April. The time of harvest depends on a variety of factors – most notably the subjective determination of
ripeness. As the grape ripens on the vines, sugars and
pH increase as acids (such as
malic acid
Malic acid is an organic compound with the molecular formula . It is a dicarboxylic acid that is made by all living organisms, contributes to the sour taste of fruits, and is used as a food additive. Malic acid has two stereoisomeric forms ( ...
) decrease.
Tannin
Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and Precipitation (chemistry), precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' is widel ...
s and other
phenolics also develop, which can affect the flavors and aromas in the resulting wine. The threat of detrimental weather and vine diseases (such as
grey rot) can also play a role in the time table. The balance of all these factors contributes to when a winemaker or vineyard manager decides that it is time to harvest.
Following the harvest, the vines continue the process of photosynthesis, creating carbohydrate reserves to store in the vine's roots and trunks. It will continue doing this until an appropriate level of reserves have been stored. At that point the chlorophyll in the leaves begins to break down and the leaves change color from green to yellow. Following the first frost the leaves begin to fall as the vine starts to enter its winter dormancy period. The following spring, the cycle begins again.
See also
*
BBCH-scale (grape)
References
External links
Annual Cycle of the Grapevine
Stages of Grape Berry Development, information from Cooperative Extension
{{Viticulture
Viticulture