Blastoids (class Blastoidea) are an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

type of stemmed

echinoderm An echinoderm () is any animal of the phylum Echinodermata (), which includes starfish, brittle stars, sea urchins, sand dollars and sea cucumbers, as well as the sessile sea lilies or "stone lilies". While bilaterally symmetrical as ...

, often referred to as sea buds. They first appear, along with many other echinoderm classes, in the

Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era, and the second of twelve periods of the Phanerozoic Eon (geology), Eon. The Ordovician spans 41.6 million years f ...

period, and reached their greatest diversity in the Mississippian subperiod of the

Carboniferous The Carboniferous ( ) is a Geologic time scale, geologic period and System (stratigraphy), system of the Paleozoic era (geology), era that spans 60 million years, from the end of the Devonian Period Ma (million years ago) to the beginning of the ...

period. However, blastoids may have originated in the

Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordov ...

. Blastoids persisted until their

extinction Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

at the end of

Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the s ...

, about 250 million years ago. Although never as diverse as their contemporary relatives, the

crinoid Crinoids are marine invertebrates that make up the class Crinoidea. Crinoids that remain attached to the sea floor by a stalk in their adult form are commonly called sea lilies, while the unstalked forms, called feather stars or comatulids, are ...

s, blastoids are common fossils, especially in many Mississippian-age rocks.

Description

Like most echinoderms, blastoids were protected by a set of interlocking plates of

calcium carbonate Calcium carbonate is a chemical compound with the chemical formula . It is a common substance found in Rock (geology), rocks as the minerals calcite and aragonite, most notably in chalk and limestone, eggshells, gastropod shells, shellfish skel ...

, which formed the main body, or ''

theca In biology, a theca (: thecae) is a sheath or a covering. Botany In botany, the theca is related to plant's flower anatomy. The theca of an angiosperm consists of a pair of microsporangia that are adjacent to each other and share a common ar ...

''. In life, the theca of a typical blastoid was attached to a stalk or column made up of stacked disc-shaped plates. The other end of the column was attached to the ocean floor by a holdfast, very much like stalked

s. The stalk was usually relatively short, and in some species, was absent, with the holdfast being attached directly to the base of the theca. The mouth was at the summit of the theca. Radiating like flower petals from the center were five food grooves, or '' ambulacra''. Each ambulacrum had many long thin fine structures called ''brachioles'', which were used to trap food particles and bring them to the mouth. Brachioles were delicate structures, and in fossils are not usually preserved in place. A series of five spiracle plates surrounded the star-shaped mouth, which included the

anus In mammals, invertebrates and most fish, the anus (: anuses or ani; from Latin, 'ring' or 'circle') is the external body orifice at the ''exit'' end of the digestive tract (bowel), i.e. the opposite end from the mouth. Its function is to facil ...

, mouth and entrances to a set of five complex, folded respiratory organs known as ''hydrospires''. These spiracles prevented mixing of the various fluids. Waste elimination was through the ''anispiracle'', an opening formed by the fusing of anus and adjacent spiracles. Pentremites Glen Dean Fm KY aboral

Like crinoids, blastoids were high-level stalked suspension feeders (feeding mainly on

plankton Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are ca ...

ic organisms) that inhabited clear-to-silty, moderately agitated ocean waters from shelf to basin. The food gathering system of blastoids consisted of several types of ambulacra. Food entered the brachiolar ambulacra, was transferred to the side ambulacra through the brachiolar pit, then transferred to the main (median) ambulacra, and finally entered the mouth. Each of these ambulacra was roofed by cover plates. The cover plates of the brachiolar groove were movable and could open, allowing food to enter, or close as needed. Other cover plates may also have been movable.

Taxonomy

Blastoids are assumed to have evolved from the

Cystoids Cystoidea was defined as a class (biology), class of extinct paleozoic blastozoan echinoderms established to encompass stalked taxa that were neither crinoids nor blastoids. It was shown to be polyphyletic in the late 1960s but continues to be u ...

. Blastoids are subdivided into two orders: Fissiculata, which are characterized by direct entrance to the individual hydrospires by way of slits; and Spiraculata, which are characterized by indirect entrance to the hydrospires through canals by way of pores. The earliest blastoid yet found, ''Macurdablastus'' from the Middle Ordovician of

Tennessee Tennessee (, ), officially the State of Tennessee, is a landlocked U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders Kentucky to the north, Virginia to the northeast, North Carolina t ...

, cannot be classified as either order.

References

External links

Blastoids at UC Berkeley Museum of Paleontology
*Drawings and color reconstruction of ''Pentremites godoni'' blastoid a
www.emilydamstra.comThree Dimensional Structure and Fluid Flow though the Hydrospires of the Blastoid Pentremites rusticus
{{Authority control Blastozoa Ordovician echinoderms Devonian echinoderms Permian echinoderms Ordovician first appearances Permian extinctions Lopingian extinctions