Acacia Harpophylla
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''Acacia harpophylla'', commonly known as brigalow, brigalow spearwood or orkor, is an
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
tree of Australia. The
Aboriginal Australian Aboriginal Australians are the various Indigenous peoples of the Australian mainland and many of its islands, such as Tasmania, Fraser Island, Hinchinbrook Island, the Tiwi Islands, and Groote Eylandt, but excluding the Torres Strait Islands ...
group the
Gamilaraay The Gamilaraay, also known as Gomeroi, Kamilaroi, Kamillaroi and other variations, are an Aboriginal Australian people whose lands extend from New South Wales to southern Queensland. They form one of the four largest Indigenous nations in Aust ...
peoples know the tree as Barranbaa or Burrii. It is found in central and coastal
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
to northern
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
. It can reach up to tall and forms extensive open-forest communities on clay soils.


Description

The tree is root-suckering and has hard, furrowed and almost black coloured bark. The
glabrous Glabrousness (from the Latin '' glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...
or hairy branchlets are angular at extremities. Like most species of ''Acacia'' it has
phyllode Phyllodes are modified petioles or leaf stems, which are leaf-like in appearance and function. In some plants, these become flattened and widened, while the leaf itself becomes reduced or vanishes altogether. Thus the phyllode comes to serve the ...
s rather than true leaves. The coriaceous, sericeous and evergreen phyllodes have a
falcate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular ...
shape with a length of and a width of and have many closely parallel nerves with three to seven of the nerves are more prominent than the others. When it blooms, between July and October, it produces condensed
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s in groups of two to eight on racemes, usually appearing as axillary clusters. The spherical flower-heads have a diameter of and contain 15 to 35 golden coloured flowers. After flowering crustaceous and glabrous
seed pod This page provides a glossary of plant morphology. Botanists and other biologists who study plant morphology use a number of different terms to classify and identify plant organs and parts that can be observed using no more than a handheld magnify ...
s form that are subterete and straight to slightly curved. The pods are raised over and constricted between seeds and have a length of up to and a width of with longitudinal nerves. The soft, dull, brown coloured seeds within the pods are arranged longitudinally and have an oblong or broadly elliptic shape and are flattened but thick with a length of and have a filiform funicle.


Distribution and habitat

Two species, brigalow (''A. harpophylla'') and gidgee ( ''A. cambagei'') form open woodlands on flat and gently undulating terrain on heavy and relatively fertile clay and clay-loam soils primarily in the 300-700mm annual rainfall region of Eastern Australia. These woodlands extend from a northern extreme of 20° S into northern New South Wales. Brigalow and gidgee occur as mixed communities in some regions and are commonly associated with several other woody species, including overstorey species such as '' Eucalyptus coolabah'', ''E. cambageana'', ''
Casuarina cristata ''Casuarina cristata'' is an Australian tree of the sheoak family Casuarinaceae known as belah. It is native to a band across inland eastern Australia. Taxonomy The Dutch botanist Friedrich Anton Wilhelm Miquel described the belah in 1848, and ...
'', and a range of understorey species.Anderson, E. and P. Back (1990). Fire in brigalow lands. Fire in the management of northern Australian pastoral lands. T. C. Grice and S. M. Slatter. St. Lucia, Australia, Tropical Grassland Society of Australia. ''A. tephrina'', ''A. georginae'' and ''A. argyrodendron'' also occupy similar habitats and have similar habits and growth forms, but are less widespread, while a number of other ''Acacia'' species also form structurally similar communities.Johnson, R. W. and W. H. Burrows (1994). Acacia open forest, woodlands and shrublands. Australian Vegetation. R. H. Groves. Cambridge, Cambridge University Press. Brigalow occurs from coastal regions receiving in excess of rainfall per year through to the semi arid rainfall region although it is primarily a semi-arid zone species. Gidgee (''A. cambagei'') replaces brigalow as rainfall drops in western regions and extends from . Gidgee, with a maximum height of approximately is somewhat smaller than brigalow which can attain heights of . In the north-western regions Black gidgee (''A. argyrodendron'') replaces brigalow in many areas, while in Central-Western districts Boree (''A. tephrina'') forms woodlands and shrublands, frequently on cracking clay soils and often in association with ''A. cambagei''. Georgina gidgee (''A. georginae'') woodlands are found in more arid regions in the rainfall belt. In New South Wales it is found from around
Roto ''Roto'', f. ''rota'', (literally "torn" or "broken") is a term used to refer to Chilean people and in particular to the common Chilean. In Chile from the start of the 20th century it was applied with a negative classist connotation to poor ci ...
in the south, to around
Hungerford Hungerford is a historic market town and civil parish in Berkshire, England, west of Newbury, east of Marlborough, northeast of Salisbury and 60 miles (97 km) west of London. The Kennet and Avon Canal passes through the town alongside the ...
in the east and
Willow Tree Willows are a genus of trees. Willow Tree may refer to: Places * Willow Tree, New South Wales, a village in Australia * Willow Tree railway station, in Australia * Willow Tree (LIRR station), a railway station in New York Entertainment * "Wil ...
in the west along the
Great Dividing Range The Great Dividing Range, also known as the East Australian Cordillera or the Eastern Highlands, is a cordillera system in eastern Australia consisting of an expansive collection of mountain ranges, plateaus and rolling hills, that runs rough ...
. In Queensland it is found as far north as
Townsville Townsville is a city on the north-eastern coast of Queensland, Australia. With a population of 180,820 as of June 2018, it is the largest settlement in North Queensland; it is unofficially considered its capital. Estimated resident population, 3 ...
.


Taxonomy

The species was first formally described by the botanist
George Bentham George Bentham (22 September 1800 – 10 September 1884) was an English botanist, described by the weed botanist Duane Isely as "the premier systematic botanist of the nineteenth century". Born into a distinguished family, he initially studi ...
in 1864 as part of the work ''
Flora Australiensis ''Flora Australiensis: a description of the plants of the Australian Territory'', more commonly referred to as ''Flora Australiensis'', and also known by its standard abbreviation ''Fl. Austral.'', is a seven-volume flora of Australia published be ...
''. It was reclassified as ''Racosperma harpophyllum'' by
Leslie Pedley Leslie Pedley (19 May 1930 – 27 November 2018)IPNILeslie Pedley/ref> was an Australian botanist who specialised in the genus ''Acacia''. He is notable for bringing into use the generic name ''Racosperma'', creating a split in the genus, which r ...
and transferred back the genus ''Acacia'' in 2001. The
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
was collected from around
Rockhampton Rockhampton is a city in the Rockhampton Region of Central Queensland, Australia. The population of Rockhampton in June 2021 was 79,967, Estimated resident population, 30 June 2018. making it the fourth-largest city in the state outside of the ...
. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
is in reference to the falcate shape of the phyllodes on the tree.


Response to fire

Species associated with these brigalow communities generally have a good capacity for re-sprouting following fire, and brigalow itself sprouts freely from the butt, roots and living stems in response to fire damage. Both gidgee and blackwood, in contrast, have a limited capacity to resprout following fire damage. A notable exception to the fire tolerance of brigalow communities occurs in what are referred to as softwood scrubs, which are dense communities of brigalow and a range of particularly fire-sensitive species. Fire in any brigalow or gidgee woodland would be a rare event under natural circumstances, since pasture is at best sparse in these communities, consisting of '' Chloris'', ''
Paspalidium ''Paspalidium'' (watercrown grass) is a genus of tropical and subtropical plants in the grass family. ''Paspalidium'' includes about 40 species native to tropical and subtropical regions of Asia, Africa, Australia, and the Americas Th ...
'', '' Dicanthium'', ''
Sporobolus ''Sporobolus'' is a nearly cosmopolitan genus of plants in the grass family. The name ''Sporobolus'' means "seed-thrower", and is derived from Ancient Greek word (), meaning "seed", and the root of () "to throw", referring to the dispersion of ...
'' and ''
Eragrostis ''Eragrostis'' is a large and widespread genus of plants in the grass family, found in many countries on all inhabited continents and many islands. ''Eragrostis'' is commonly known as lovegrass or canegrass. The name of the genus is derived fr ...
'' species.Weston, E. J. (1988). Native Pasture Communities. Native pastures in Queensland their resources and management. W. H. Burrows, J. C. Scanlan and M. T. Rutherford. Brisbane, Department of Primary Industries.


See also

* * * List of ''Acacia'' species


References

{{Taxonbar, from=Q2707674 harpophylla Flora of Queensland Flora of New South Wales Fabales of Australia Drought-tolerant trees Trees of Australia Bushfood Plants described in 1864 Taxa named by Ferdinand von Mueller