Atraphaxis is a
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s in the family
Polygonaceae
The Polygonaceae are a Family (biology), family of flowering plants known Common name, informally as the knotweed family or smartweed—buckwheat family in the United States. The Botanical name, name is Basionym, based on the genus ''Polygonum'', ...
with about 40
species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and bear the leaves and flowers. The leaves are simple and alternate, with very short stalks (almost sessile). The
ochrea
An ochrea (Latin ''ocrea'', greave or protective legging), also spelled ocrea, is a plant structure formed of stipules fused into a sheath surrounding the stem, and is typically found in the Polygonaceae
The Polygonaceae are a Family (biology ...
s are membranous and usually two-veined, more-or-less joined at the base. The
inflorescence
In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
is made up of several bundles (
fascicles) of one to three flowers. The flowers have persistent
tepal
A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s, either arranged in a narrow tube with unequal lobes or bell-shaped with equal segments. The fruits are wingless
achene
An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple fruit, simple dry fruits, dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and Dehi ...
s.
[
]
Taxonomy
The genus ''Atraphaxis'' was erected by Carl Linnaeus
Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1753. As with many other genera in the family Polygonaceae, the boundaries between the genera have been unclear, and some or all species placed in other genera. Molecular phylogenetic
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
studies have shown that ''Atraphaxis'' forms a distinct clade.[ The genus is placed in the tribe Polygoneae of the subfamily ]Polygonoideae
Polygonoideae is a subfamily of plants in the family Polygonaceae. It includes a number of plants that can be highly invasive, such as Japanese knotweed, '' Reynoutria japonica'', and its hybrid with '' R. sachalinensis'', ''R.'' × ''bohemica' ...
. Within the tribe, it is most closely related to the genera ''Duma
A duma () is a Russian assembly with advisory or legislative functions.
The term ''boyar duma'' is used to refer to advisory councils in Russia from the 10th to 17th centuries. Starting in the 18th century, city dumas were formed across Russia ...
'' and ''Polygonum
''Polygonum'' is a genus of about 130 species of flowering plants in the buckwheat and knotweed family Polygonaceae. Common names include knotweed and knotgrass (though the common names may refer more broadly to plants from Polygonaceae). In the ...
'', forming the so-called "DAP clade".[
]
Species
, Plants of the World Online
Plants of the World Online (POWO) is an online taxonomic database published by the Royal Botanic Gardens, Kew.
History
Following the Convention on Biological Diversity, the Royal Botanic Gardens in Kew launched Plants of the World Online i ...
accepted 41 species:[
*'' Atraphaxis angustifolia'' Jaub. & Spach
*'']Atraphaxis ariana
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' (Grigorj.) T.M.Schust. & Reveal
*'' Atraphaxis arida'' (Boiss. & Hausskn.) S.Tavakkoli & Kaz.Osaloo
*'' Atraphaxis atraphaxiformis'' (Botsch.) T.M.Schust. & Reveal
*''Atraphaxis aucheri
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' Jaub. & Spach
*''Atraphaxis avenia
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' Botsch.
*''Atraphaxis badghysi
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' Kult.
*''Atraphaxis billardierei
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' Jaub. & Spach
*''Atraphaxis binaludensis
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' S.Tavakkoli, Mozaff. & Kaz.Osaloo
*''Atraphaxis botuliformis
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' (Mozaff.) S.Tavakkoli & Mozaff.
*''Atraphaxis bracteata
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' Losinsk.
*''Atraphaxis canescens
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' Bunge
*''Atraphaxis caucasica
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' (Hoffm.) Pavlov
*''Atraphaxis compacta
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' Ledeb.
*'' Atraphaxis daghestanica'' (Lovelius) Lovelius
*'' Atraphaxis decipiens'' Jaub. & Spach
*'' Atraphaxis dumosa'' (Boiss.) S.Tavakkoli & Kaz.Osaloo
*'' Atraphaxis frutescens'' (L.) K.Koch
*'' Atraphaxis grandiflora'' Willd.
*'' Atraphaxis intricata'' Mozaff.
*'' Atraphaxis irtyschensis'' Chang Y.Yang & Y.L.Han
*'' Atraphaxis kamelinii'' Yurtseva
*'' Atraphaxis karataviensis'' Pavlov & Lipsch.
*'' Atraphaxis kermanica'' S.Tavakkoli & Kaz.Osaloo
*'' Atraphaxis khajeh-jamali'' (Khosravi & Poormahdi) S.Tavakkoli & Kaz.Osaloo
*'' Atraphaxis kopetdagensis'' Kovalevsk.
*'' Atraphaxis laetevirens'' (Ledeb.) Jaub. & Spach
*'' Atraphaxis macrocarpa'' Rech.f. & Schiman-Czeika
*'' Atraphaxis manshurica'' Kitag.
*'' Atraphaxis muschketowii'' Krasn.
*'' Atraphaxis popovii'' (Borodina) Yurtseva
*'' Atraphaxis pungens'' (M.Bieb.) Jaub. & Spach
*''Atraphaxis pyrifolia
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' Bunge
*''Atraphaxis radkanensis
Atraphaxis is a genus of flowering plants in the family Polygonaceae with about 40 species.
Description
Species of ''Atraphaxis'' are much branched woody plants, forming shrubs or shrubby tufts. The current year's branchlets are herbaceous and b ...
'' S.Tavakkoli, Kaz.Osaloo & Mozaff.
*'' Atraphaxis rodinii'' Botsch.
*'' Atraphaxis salicornioides'' (Jaub. & Spach) S.Tavakkoli & Kaz.Osaloo
*'' Atraphaxis seravschanica'' Pavlov
*'' Atraphaxis spinosa'' L.
*'' Atraphaxis suaedifolia'' Jaub. & Spach
*'' Atraphaxis teretifolia'' (Popov) Kom.
*'' Atraphaxis toktogulica'' (Lazkov) T.M.Schust. & Reveal
References
{{Taxonbar, from=Q889292
Polygonoideae
Polygonaceae genera