The Assam macaque (''Macaca assamensis'') or Assamese macaque is a

macaque The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and Europe (in Gibraltar). Macaques are principally f ...

of the

Old World monkey Old World monkeys are primates in the family Cercopithecidae (). Twenty-four genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons (genus '' Papio''), red colobus (genus '' Piliocolob ...

family native to

South South is one of the cardinal directions or compass points. The direction is the opposite of north and is perpendicular to both west and east. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Proto-Germanic ''*sunþa ...

and

Southeast Asia Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...

. Since 2008, it has been listed as

Near Threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...

on the

IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...

, as it is experiencing significant declines due to

poaching Poaching is the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set against the huntin ...

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

degradation, and fragmentation.

Characteristics

The Assam macaque has a yellowish-grey to dark brown pelage. The facial skin is dark brownish to purplish. The head has a dark fringe of hair on the cheeks directed backwards to the ears. The hair on the crown is parted in the middle. The shoulders, head and arms tend to be paler than the hindquarters, which are greyish. The tail is well-haired and short. Head-to-body-length measures , and the tail is long. Adult weight is .

Distribution and habitat

In Nepal, the Assam macaque occurs east of the Kali Gandaki river at elevations of . In northeastern India, it lives in tropical and subtropical semi evergreen forests, dry

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

and

montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...

s up to elevations of . In Laos and Vietnam, it prefers elevations above . It usually inhabits hilly areas above , but in the wetter east it also occurs in the lowlands below this elevation. In

limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...

karst Karst () is a topography formed from the dissolution of soluble carbonate rocks such as limestone and Dolomite (rock), dolomite. It is characterized by features like poljes above and drainage systems with sinkholes and caves underground. Ther ...

forests, it also occurs at lower elevations.

Ecology and behaviour

Assam macaques are diurnal, and at times both

arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...

and terrestrial. They are

omnivorous An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize ...

and feed on

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

s, leaves,

invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...

s and

cereal A cereal is a grass cultivated for its edible grain. Cereals are the world's largest crops, and are therefore staple foods. They include rice, wheat, rye, oats, barley, millet, and maize ( Corn). Edible grains from other plant families, ...

s. In

Namdapha National Park Namdapha National Park is a large national park in Arunachal Pradesh of Northeast India. The park was established in 1983. With more than 1,000 floral and about 1,400 faunal species, it is a biodiversity hotspot in the Eastern Himalayas. It ha ...

Arunachal Pradesh Arunachal Pradesh (; ) is a States and union territories of India, state in northeast India. It was formed from the North-East Frontier Agency (NEFA) region, and India declared it as a state on 20 February 1987. Itanagar is its capital and la ...

, 15 groups were recorded in 2002 comprising 209 individuals. The population had a group density of 1.11 individuals per , and an average group size of 13.93 individuals.Chetry, D., Medhi, R., Biswas, J., Das, D. and Bhattacharjee, P. C. (2003)
''Nonhuman primates in the Namdapha National Park, Arunachal Pradesh, India''
International Journal of Primatology 24(2): 383–388. During a survey in Nepal's Langtang National Park in 2007, a total of 213 Assamese macaques were encountered in 9 groups in the study area of . Troop sizes varied between 13 and 35 individuals, with a mean troop size of 23.66 individuals, and comprised 31% adult females, 16% adult males, and their young of various ages. They preferred

maize Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...

kernals, followed by

potato The potato () is a starchy tuberous vegetable native to the Americas that is consumed as a staple food in many parts of the world. Potatoes are underground stem tubers of the plant ''Solanum tuberosum'', a perennial in the nightshade famil ...

tubers, but also raided fields with

wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...

buckwheat Buckwheat (''Fagopyrum esculentum'') or common buckwheat is a flowering plant in the knotweed family Polygonaceae cultivated for its grain-like seeds and as a cover crop. Buckwheat originated around the 6th millennium BCE in the region of what ...

, and

millet Millets () are a highly varied group of small-seeded grasses, widely grown around the world as cereal crops or grains for fodder and human food. Most millets belong to the tribe Paniceae. Millets are important crops in the Semi-arid climate, ...

Threats

The threats to this species' habitat include selective

logging Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidder, skidding, on-site processing, and loading of trees or trunk (botany), logs onto logging truck, truckspet trade. Additionally, hybridization with adjacent species poses a threat to some populations.Molur, S., Brandon-Jones, D., Dittus, W., Eudey, A., Kumar, A., Singh, M., Feeroz, M. M., Chalise, M., Priya, P. and Walker, S. (2003)
''Status of South Asian Primates: Conservation Assessment and Management Plan Report''
. Workshop Report, 2003. Zoo Outreach Organization/CBSG-South Asia, Coimbatore, India.

Conservation

''Macaca assamensis'' is listed in

CITES Appendix II CITES (shorter acronym for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of inte ...

. It is legally protected in all countries of occurrence. For the populations in India, the species is listed under Schedule II, part I of the Indian Wildlife Act.

References

External links

ARKive: Assam macaque (''Macaca assamensis'')
{{Taxonbar, from=Q738925 Assam macaque Primates of Asia Mammals of Bangladesh Mammals of Bhutan Mammals of Myanmar Mammals of China Mammals of India Mammals of Laos Mammals of Nepal Mammals of Thailand Mammals of Vietnam Fauna of Yunnan Fauna of Assam Assam macaque Taxa named by John McClelland (doctor)