An adaptive system is a set of interacting or interdependent entities, real or abstract, forming an integrated whole that together are able to respond to environmental changes or changes in the interacting parts, in a way analogous to either continuous physiological
homeostasis or evolutionary
adaptation
In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...
in
biology.
Feedback loops
Feedback occurs when outputs of a system are routed back as inputs as part of a chain of cause-and-effect that forms a circuit or loop. The system can then be said to ''feed back'' into itself. The notion of cause-and-effect has to be handled ...
represent a key feature of adaptive systems, such as
ecosystems and individual
organisms; or in the human world,
communities
A community is a Level of analysis, social unit (a group of living things) with commonality such as place (geography), place, Norm (social), norms, religion, values, Convention (norm), customs, or Identity (social science), identity. Communiti ...
,
organizations, and
families. Adaptive systems can be organized into a hierarchy.
Artificial adaptive systems include
robots with
control system
A control system manages, commands, directs, or regulates the behavior of other devices or systems using control loops. It can range from a single home heating controller using a thermostat controlling a domestic boiler to large industrial c ...
s that utilize
negative feedback
Negative feedback (or balancing feedback) occurs when some function (Mathematics), function of the output of a system, process, or mechanism is feedback, fed back in a manner that tends to reduce the fluctuations in the output, whether caused by ...
to maintain desired states.
The law of adaptation
The law of adaptation may be stated informally as:
Formally, the law can be defined as follows:
Given a system
, we say that a physical event
is a stimulus for the system
if and only if the probability
that the system suffers a change or be perturbed (in its elements or in its processes) when the event
occurs is strictly greater than the prior probability that
suffers a change independently of
:
:
''Let
be an arbitrary system subject to changes in time
and let
be an arbitrary event that is a stimulus for the system
: we say that
is an adaptive system if and only if when t tends to infinity
the probability that the system
change its behavior
in a time step
given the event
is equal to the probability that the system change its behavior independently of the occurrence of the event
. In mathematical terms:''
#-
#-
Thus, for each instant
will exist a temporal interval
such that:
:
Hierarchy of adaptations: Practopoiesis

How do various types of adaptations interact in a living system? Practopoiesis, a term due to its originator Danko Nikolić, is a reference to a hierarchy of adaptation mechanisms answering this question. The adaptive hierarchy forms a kind of a self-adjusting system in which
autopoiesis of the entire ''organism'' or a ''cell'' occurs through a hierarchy of
allopoietic interactions among ''components''.
This is possible because the components are organized into a
poietic hierarchy: adaptive actions of one component result in creation of another component. The theory proposes that living systems exhibit a hierarchy of a total of four such adaptive poietic operations:
''
evolution'' (i) → ''
gene expression
Gene expression is the process by which information from a gene is used in the synthesis of a functional gene product that enables it to produce end products, protein or non-coding RNA, and ultimately affect a phenotype, as the final effect. The ...
'' (ii) → ''non gene-involving
homeostatic mechanisms (anapoiesis)'' (iii) → ''final cell function'' (iv)
As the hierarchy evolves towards higher levels of organization, the speed of adaptation increases. Evolution is the slowest; gene expression is faster; and so on. The final cell function is the fastest. Ultimately, practopoiesis challenges current neuroscience doctrine by asserting that mental operations primarily occur at the homeostatic, anapoietic level (iii) — i.e., that minds and thought emerge from fast homeostatic mechanisms poietically controlling the cell function. This contrasts the widespread assumption that
thinking is synonymous with computations executed at the level of
neural activity
Neurotransmission (Latin: ''transmissio'' "passage, crossing" from ''transmittere'' "send, let through") is the process by which signaling molecules called neurotransmitters are released by the axon terminal of a neuron (the presynaptic neuron), ...
(i.e., with the 'final cell function' at level iv).
Sharov proposed that only
Eukaryote
Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacte ...
cells can achieve all four levels of organization.
Each slower level contains knowledge that is more general than the faster level; for example, genes contain more general knowledge than anapoietic mechanisms, which in turn contain more general knowledge than cell functions. This hierarchy of knowledge enables the anapoietic level to implement
concepts, which are the most fundamental ingredients of a mind. Activation of concepts through anapoiesis is suggested to underlie
ideasthesia. Practopoiesis also has implications for understanding the limitations of
Deep Learning
Deep learning (also known as deep structured learning) is part of a broader family of machine learning methods based on artificial neural networks with representation learning. Learning can be supervised, semi-supervised or unsupervised.
De ...
. It has been proposed that the biological underpinnings of anapoietic mechanisms are
metabotropic receptors
A metabotropic receptor, also referred to by the broader term G-protein-coupled receptor, is a type of membrane receptor that initiates a number of metabolic steps to modulate cell activity. The nervous system utilizes two types of receptors: met ...
and
G protein-gated ion channel
G protein-gated ion channels are a family of transmembrane ion channels in neurons and atrial myocytes that are directly gated by G proteins.
Overview of mechanisms and function
Generally, G protein-gated ion channels are specific ion channels ...
s.
Empirical tests of practopoiesis require learning on double-loop tasks: One needs to assess how the learning capability adapts over time, i.e., how the system learns to learn (adapts its adapting skills).
Benefit of self-adjusting systems
In an adaptive system, a parameter changes slowly and has no preferred value. In a self-adjusting system though, the parameter value “depends on the history of the system dynamics”. One of the most important qualities of ''self-adjusting systems'' is its “
adaptation to the edge of chaos” or ability to avoid
chaos. Practically speaking, by heading to the
edge of chaos without going further, a leader may act spontaneously yet without disaster. A March/April 2009 Complexity article further explains the self-adjusting systems used and the realistic implications. Physicists have shown that
adaptation
In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...
to the
edge of chaos occurs in almost all systems with
feedback
Feedback occurs when outputs of a system are routed back as inputs as part of a chain of cause-and-effect that forms a circuit or loop. The system can then be said to ''feed back'' into itself. The notion of cause-and-effect has to be handled ...
.
See also
*
Adaptive immune system
*
Artificial neural network
*
Complex adaptive system
*
Diffusion of innovations
*
Ecosystems
*
Gaia hypothesis
The Gaia hypothesis (), also known as the Gaia theory, Gaia paradigm, or the Gaia principle, proposes that living organisms interact with their inorganic surroundings on Earth to form a synergistic and self-regulating, complex system that help ...
*
Gene expression programming
In computer programming, gene expression programming (GEP) is an evolutionary algorithm that creates computer programs or models. These computer programs are complex tree structures that learn and adapt by changing their sizes, shapes, and compos ...
*
Genetic algorithms
*
Learning
Learning is the process of acquiring new understanding, knowledge, behaviors, skills, value (personal and cultural), values, attitudes, and preferences. The ability to learn is possessed by humans, animals, and some machine learning, machines ...
*
Neural adaptation
Notes
References
*
External links
{{Wiktionary , practopoiesis
* Funn
animated videoexplaining the theory of practopoiesis, made by Mind & Brain.
* Practopoiesis offers solutions t
nine long-standing problems in neuroscience and philosophy of mind
Blog series on practopoiesis
Control engineering
Cybernetics
Systems theory