An adaptive system is a set of interacting or interdependent entities, real or abstract, forming an integrated whole that together are able to respond to environmental changes or changes in the interacting parts, in a way analogous to either continuous physiological
homeostasis
In biology, homeostasis (British English, British also homoeostasis) Help:IPA/English, (/hɒmɪə(ʊ)ˈsteɪsɪs/) is the state of steady internal, physics, physical, and chemistry, chemical conditions maintained by organism, living systems. Thi ...
or evolutionary
adaptation in
biology
Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditar ...
.
Feedback loops represent a key feature of adaptive systems, such as
ecosystems
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
and individual
organisms
In biology, an organism () is any living system that functions as an individual entity. All organisms are composed of cells ( cell theory). Organisms are classified by taxonomy into groups such as multicellular animals, plants, and f ...
; or in the human world,
communities,
organizations
An organization or organisation (Commonwealth English; see spelling differences), is an entity—such as a company, an institution, or an association—comprising one or more people and having a particular purpose.
The word is derived fr ...
, and
families. Adaptive systems can be organized into a hierarchy.
Artificial adaptive systems include
robots with
control system
A control system manages, commands, directs, or regulates the behavior of other devices or systems using control loops. It can range from a single home heating controller using a thermostat controlling a domestic boiler to large industrial ...
s that utilize
negative feedback
Negative feedback (or balancing feedback) occurs when some function of the output of a system, process, or mechanism is fed back in a manner that tends to reduce the fluctuations in the output, whether caused by changes in the input or by othe ...
to maintain desired states.
The law of adaptation
The law of adaptation may be stated informally as:
Formally, the law can be defined as follows:
Given a system
, we say that a physical event
is a stimulus for the system
if and only if the probability
that the system suffers a change or be perturbed (in its elements or in its processes) when the event
occurs is strictly greater than the prior probability that
suffers a change independently of
:
:
''Let
be an arbitrary system subject to changes in time
and let
be an arbitrary event that is a stimulus for the system
: we say that
is an adaptive system if and only if when t tends to infinity
the probability that the system
change its behavior
in a time step
given the event
is equal to the probability that the system change its behavior independently of the occurrence of the event
. In mathematical terms:''
#-
#-
Thus, for each instant
will exist a temporal interval
such that:
:
Hierarchy of adaptations: Practopoiesis

How do various types of adaptations interact in a living system? Practopoiesis, a term due to its originator Danko Nikolić, is a reference to a hierarchy of adaptation mechanisms answering this question. The adaptive hierarchy forms a kind of a self-adjusting system in which
autopoiesis
The term autopoiesis () refers to a system capable of producing and maintaining itself by creating its own parts.
The term was introduced in the 1972 publication '' Autopoiesis and Cognition: The Realization of the Living'' by Chilean biologists ...
of the entire ''organism'' or a ''cell'' occurs through a hierarchy of
allopoietic interactions among ''components''.
This is possible because the components are organized into a
poietic hierarchy: adaptive actions of one component result in creation of another component. The theory proposes that living systems exhibit a hierarchy of a total of four such adaptive poietic operations:
''
evolution
Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
'' (i) → ''
gene expression
Gene expression is the process by which information from a gene is used in the synthesis of a functional gene product that enables it to produce end products, protein or non-coding RNA, and ultimately affect a phenotype, as the final effect. ...
'' (ii) → ''non gene-involving
homeostatic mechanisms (anapoiesis)'' (iii) → ''final cell function'' (iv)
As the hierarchy evolves towards higher levels of organization, the speed of adaptation increases. Evolution is the slowest; gene expression is faster; and so on. The final cell function is the fastest. Ultimately, practopoiesis challenges current neuroscience doctrine by asserting that mental operations primarily occur at the homeostatic, anapoietic level (iii) — i.e., that minds and thought emerge from fast homeostatic mechanisms poietically controlling the cell function. This contrasts the widespread assumption that
thinking
In their most common sense, the terms thought and thinking refer to conscious cognitive processes that can happen independently of sensory stimulation. Their most paradigmatic forms are judging, reasoning, concept formation, problem solving, an ...
is synonymous with computations executed at the level of
neural activity
Neurotransmission (Latin: ''transmissio'' "passage, crossing" from ''transmittere'' "send, let through") is the process by which signaling molecules called neurotransmitters are released by the axon terminal of a neuron (the presynaptic neuron), ...
(i.e., with the 'final cell function' at level iv).
Sharov proposed that only
Eukaryote
Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bact ...
cells can achieve all four levels of organization.
Each slower level contains knowledge that is more general than the faster level; for example, genes contain more general knowledge than anapoietic mechanisms, which in turn contain more general knowledge than cell functions. This hierarchy of knowledge enables the anapoietic level to implement
concept
Concepts are defined as abstract ideas. They are understood to be the fundamental building blocks of the concept behind principles, thoughts and beliefs.
They play an important role in all aspects of cognition. As such, concepts are studied by s ...
s, which are the most fundamental ingredients of a mind. Activation of concepts through anapoiesis is suggested to underlie
ideasthesia
Ideasthesia (alternative spelling ideaesthesia) is a neuropsychological phenomenon in which activations of concepts (inducers) evoke perception-like sensory experiences (concurrents). The name comes from the Ancient Greek () and (), meaning 'se ...
. Practopoiesis also has implications for understanding the limitations of
Deep Learning. It has been proposed that the biological underpinnings of anapoietic mechanisms are
metabotropic receptors
A metabotropic receptor, also referred to by the broader term G-protein-coupled receptor, is a type of membrane receptor that initiates a number of metabolic steps to modulate cell activity. The nervous system utilizes two types of receptors: me ...
and
G protein-gated ion channels.
Empirical tests of practopoiesis require learning on double-loop tasks: One needs to assess how the learning capability adapts over time, i.e., how the system learns to learn (adapts its adapting skills).
Benefit of self-adjusting systems
In an adaptive system, a parameter changes slowly and has no preferred value. In a self-adjusting system though, the parameter value “depends on the history of the system dynamics”. One of the most important qualities of ''self-adjusting systems'' is its “
adaptation to the edge of chaos” or ability to avoid
chaos. Practically speaking, by heading to the
edge of chaos without going further, a leader may act spontaneously yet without disaster. A March/April 2009 Complexity article further explains the self-adjusting systems used and the realistic implications. Physicists have shown that
adaptation to the
edge of chaos occurs in almost all systems with
feedback
Feedback occurs when outputs of a system are routed back as inputs as part of a chain of cause-and-effect that forms a circuit or loop. The system can then be said to ''feed back'' into itself. The notion of cause-and-effect has to be handled ...
.
See also
*
Adaptive immune system
The adaptive immune system, also known as the acquired immune system, is a subsystem of the immune system that is composed of specialized, systemic cells and processes that eliminate pathogens or prevent their growth. The acquired immune system ...
*
Artificial neural network
Artificial neural networks (ANNs), usually simply called neural networks (NNs) or neural nets, are computing systems inspired by the biological neural networks that constitute animal brains.
An ANN is based on a collection of connected units ...
*
Complex adaptive system
A complex adaptive system is a system that is '' complex'' in that it is a dynamic network of interactions, but the behavior of the ensemble may not be predictable according to the behavior of the components. It is '' adaptive'' in that the indi ...
*
Diffusion of innovations
Diffusion of innovations is a theory that seeks to explain how, why, and at what rate new ideas and technology spread. Everett Rogers, a professor of communication studies, popularized the theory in his book ''Diffusion of Innovations''; the boo ...
*
Ecosystems
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
*
Gaia hypothesis
The Gaia hypothesis (), also known as the Gaia theory, Gaia paradigm, or the Gaia principle, proposes that living organisms interact with their inorganic surroundings on Earth to form a synergistic and self-regulating, complex system that helps ...
*
Gene expression programming
*
Genetic algorithms
In computer science and operations research, a genetic algorithm (GA) is a metaheuristic inspired by the process of natural selection that belongs to the larger class of evolutionary algorithms (EA). Genetic algorithms are commonly used to gene ...
*
Learning
*
Neural adaptation
Notes
References
*
External links
{{Wiktionary , practopoiesis
* Funn
animated videoexplaining the theory of practopoiesis, made by Mind & Brain.
* Practopoiesis offers solutions t
nine long-standing problems in neuroscience and philosophy of mind
Blog series on practopoiesis
Control engineering
Cybernetics
Systems theory