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''Abies concolor'', the white fir, concolor fir, or Colorado fir, is a
conifer Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
ous tree in the pine family
Pinaceae The Pinaceae (), or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as Cedrus, cedars, firs, Tsuga, hemlocks, Pinyon_pine, piñons, larches, pines and spruces. The family is incl ...
. This tree is native to the mountains of western North America, including the
Sierra Nevada The Sierra Nevada ( ) is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primari ...
and southern
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in great-circle distance, straight-line distance from the northernmost part of Western Can ...
, and into the isolated mountain ranges of southern
Arizona Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
,
New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
, and
Northern Mexico Northern Mexico ( ), commonly referred as , is an informal term for the northern cultural and geographical area in Mexico. Depending on the source, it contains some or all of the states of Baja California, Baja California Sur, Chihuahua (state), ...
. It naturally occurs at elevations between . It is popular as an ornamental landscaping tree and as a
Christmas tree A Christmas tree is a decorated tree, usually an evergreen pinophyta, conifer, such as a spruce, pine or fir, associated with the celebration of Christmas. It may also consist of an artificial tree of similar appearance. The custom was deve ...
.


Description

This large evergreen conifer grows best in the central
Sierra Nevada The Sierra Nevada ( ) is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primari ...
of California, where the record specimen was recorded as tall and measured in
diameter at breast height Diameter at breast height, or DBH, is a standard method of expressing the diameter of the trunk or bole of a standing tree. DBH is one of the most common dendrometric measurements. Tree trunks are measured at the height of an adult's breast, ...
(dbh) in
Yosemite National Park Yosemite National Park ( ) is a List of national parks of the United States, national park of the United States in California. It is bordered on the southeast by Sierra National Forest and on the northwest by Stanislaus National Forest. The p ...
.American Forestry Association. 1978. National register of big trees. American Forests 84(4):19-47 The typical size of white fir ranges from tall and up to dbh. The largest specimens are found in the central Sierra Nevada, where the largest diameter recorded was found in Sierra National Forest at (1972); the west slope of the Sierra Nevada is also home to the tallest specimen on record, in height. ''Abies concolor'' subsp. ''concolor'' ( Rocky mountain white fir) rarely exceeds tall or dbh. Large (but not huge) trees in good soil range from tall and from dbh in California and southwestern Oregon and to tall and dbh in
Arizona Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
and New Mexico. The dead tree tops sometimes fork into new growth. The gray bark is usually at least thick, and brown-hued inside. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are needle-like, flattened, long and wide by thick, green to glaucous blue-green above, and with two glaucous blue-white bands of stomatal bloom below, and slightly notched to bluntly pointed at the tip. The leaf arrangement is spiral on the shoot, but with each leaf variably twisted at the base so they all lie in either two more-or-less flat ranks on either side of the shoot, or upswept across the top of the shoot but not below the shoot. The cones are long and broad, green or purple ripening pale brown, with about 100–150 scales; the scale bracts are short, and hidden in the closed cone. The winged
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s are released when the cones disintegrate at maturity about 6 months after pollination. White fir can live over 300 years.


Subspecies

As treated here, there are two
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
; these are also variously treated at either the lower rank of variety by some authors, or as distinct species by others:


Botanical collection

White fir was first collected by Augustus Fendler on his expedition to the Santa Fe area of
New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
in 1846–1847. Fendler's patron George Engelmann, a St. Louis area physician and botanist, then described the plant. This tree was first collected in California by William Lobb on his expedition to California of 1849–1853, after it was overlooked by David Douglas in his 1825–1827 expedition to the Pacific coast region. The specific epithet ''concolor'' means "all one color".


Distribution

This tree is native to the mountains of western North America from the southern Cascade Range in
Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...
, south throughout
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
and into the Sierra de San Pedro Mártir in northern
Baja California Baja California, officially the Free and Sovereign State of Baja California, is a state in Mexico. It is the northwesternmost of the 32 federal entities of Mexico. Before becoming a state in 1952, the area was known as the North Territory of B ...
, and south throughout the Colorado Plateau and southern
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in great-circle distance, straight-line distance from the northernmost part of Western Can ...
in
Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
and
Colorado Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...
, and into the isolated mountain ranges of southern
Arizona Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
, New Mexico, and
Northern Mexico Northern Mexico ( ), commonly referred as , is an informal term for the northern cultural and geographical area in Mexico. Depending on the source, it contains some or all of the states of Baja California, Baja California Sur, Chihuahua (state), ...
. It naturally occurs between above sea level.


Ecology

White fir is very closely related to grand fir (''Abies grandis''), and intergrades with it in central Oregon. It is also suggested that all of subspecies lowiana has experienced genetic introgression from grand fir. White fir occupies a different niche than grand fir, including dryer and higher elevation sites, being more drought tolerant and having thicker, fire-resistant bark. In Mexico, it is replaced by further close relatives, Durango fir (''A. durangensis'') and Mexican fir (''A. mexicana''). Like grand fir, white fir is more shade tolerant than
Douglas fir The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is the tallest tree in the Pinaceae family. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Or ...
, but less so than western hemlock and western redcedar. White fir is a climax species, which means the forest has reached complex maturity in forest succession in western coniferous forests of the U.S. White fir and yellow pine ( ponderosa pine/
Jeffrey pine ''Pinus jeffreyi'', also known as Jeffrey pine, Jeffrey's pine, yellow pine and black pine, is a North American pine tree. It is mainly found in California, but also in the westernmost part of Nevada, southwestern Oregon, and northern Baja Califo ...
) have co-existed for millennia in old growth forests throughout their range. In the presence of logging of large diameter trees and exclusion of cleansing wildfires, young trees have become abundant over the past two centuries. White fir had been regarded as a pest in the past by those in the lumber industry, but this opinion has changed. White fir is now one of the most important of all commercial softwoods according to the Western Wood Products Association. The white fir trait of retaining lower limbs creates an escape route for medium-to-small forest birds (such as spotted owl) from larger flying predators and provides a drip zone around the roots for collecting moisture. The retained limbs can become a fuel ladder that allows flames to climb up to the canopy. Limbing-up white fir, instead of removing medium to large diameter trees, in areas where the public is more likely to start fires can help keep other trees and specifically giant sequoia from experiencing canopy fire. Recent concern for sequoia groves has caused agencies to call for removal of white fir in the Sierra Nevada. While sequoia seedlings and young saplings are highly susceptible to mortality or serious injury by fire; mature sequoias are fire adapted with: fire-resistant bark, elevated canopies, self-pruning lower branches, latent buds, and serotinous cones. The sequoia ecosystem is incomplete without the mixed pine/fir and oak that make up the mid and understory. Giant Sequoia's cones release seeds when the heat of fire triggers them to open while the thick bark protects the inner
cambium A cambium (: cambiums or cambia), in plants, is a tissue layer that provides partially undifferentiated cells for plant growth. It is found in the area between xylem and phloem. A cambium can also be defined as a cellular plant tissue from whic ...
from fire damage. This tree is host to fir mistletoe (''Phoradendron pauciflorum''), a
parasitic plant A parasitic plant is a plant that derives some or all of its nutritional requirements from another living plant. They make up about 1% of angiosperms and are found in almost every biome. All Parasite, parasitic plants develop a specialized organ ...
. It is attacked by many types of insects, such as the fir engraver (''Scolytus ventralis'').


Dependent species

Mature white fir–yellow pine forests support
old-growth An old-growth forest or primary forest is a forest that has developed over a long period of time without Disturbance (ecology), disturbance. Due to this, old-growth forests exhibit unique ecological features. The Food and Agriculture Organizati ...
dependent wildlife species such as California spotted owl (''Strix occidentalis occidentalis''), Mount Pinos sooty grouse (''Dendragapus fuliginosus howardi''), and Pacific fisher (''Pekania pennanti''). The spotted owl and fisher utilize cavities in decadent large-diameter white fir for nesting and denning. The Mount Pinos sooty grouse requires large diameter trees for thermal cover and its winter diet consists of mostly white fir and yellow pine needles. This subspecies of sooty grouse has been extirpated along with a significant number of large diameter white fir from much of its range. Other subspecies of sooty grouse also utilize Douglas fir, which does not occur in the range of Mount Pinos sooty grouse. Squirrel also frequent the tree's branches. Deer browse the foliage of this species and porcupines chew the bark. Songbirds, grouse, and various mammals eat the seeds.


Uses

White fir and grand fir were used by Native Americans for medicinal use involving powdered bark or pitch to treat tuberculosis or skin ailments. The
Nlaka'pamux The Nlakaʼpamux or Nlakapamuk ( ; ), also previously known as the ''Thompson'', '' Thompson River Salish'', ''Thompson Salish'', ''Thompson River Indians'' or ''Thompson River people'', and historically as the ''Klackarpun'', ''Haukamaugh'', ''K ...
used the bark to cover lodges and make canoes. Branches were also used as bedding. White fir is a preferred construction species because of its nail-holding ability, lightness in weight, and resistance to split, twist, and pitch. It is straight-grained, non-resinous, fine-textured, stiff, and strong. It is popular as a
Christmas tree A Christmas tree is a decorated tree, usually an evergreen pinophyta, conifer, such as a spruce, pine or fir, associated with the celebration of Christmas. It may also consist of an artificial tree of similar appearance. The custom was deve ...
and for
Christmas Christmas is an annual festival commemorating Nativity of Jesus, the birth of Jesus Christ, observed primarily on December 25 as a Religion, religious and Culture, cultural celebration among billions of people Observance of Christmas by coun ...
decoration owing to its soft needles, generally excellent needle retention and abundance. It is often marketed as concolor or white fir.


Cultivation

White fir is widely planted as an
ornamental tree Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...
in parks and larger
garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...
s, particularly some
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
s of subsp. ''concolor'' selected for very bright glaucous blue foliage, such as cv. 'Violacea'. The dwarf cultivar 'Compacta', growing to a maximum height and spread of , has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. It includes the full range of cultivated p ...
.


Gallery

Image:Abies_concolor_8076.jpg, ''A. concolor'' subsp. ''lowiana'' foliage upperside Image:Abies_concolor_8065.jpg, ''A. concolor'' subsp. ''lowiana'' foliage underside Image:Baby whitefir.jpg, A young sapling of subsp. ''lowiana'' on Mount Whitney Image:White fir MN 2007.JPG, White fir in garden environment at Minnesota Landscape Arboretum Image:Young Abies concolor.jpg, Small stand of young white firs on Cuyamaca Peak, California Image:Toro Peak 537.jpg, White firs at Toro Peak, California File:Arceuthobium abietinum 1.jpg, Foliage, with '' Arceuthobium abietinum'' infestation


References


Further reading

* *


External links

*
Jepson eFlora Treatment - ''Abies concolor''

Gymnosperm Database: ''Abies concolor''
— ''(treated as varieties of one species)''.
Interactive Distribution Map of ''Abies concolor''

USDA Plants Profile for Abies concolor (white fir)

Conifers Around the World: Abies concolor - Rocky Mountain White Fir

Conifer Database
{{DEFAULTSORT:Abies Concolor concolor Flora of the Southwestern United States Flora of Baja California Flora of Chihuahua (state) Flora of Colorado Flora of Idaho Flora of Montana Flora of New Mexico Flora of Sonora Flora of Wyoming Natural history of the California Coast Ranges Natural history of the Peninsular Ranges Trees of Northern America