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Priapulid
Priapulida (priapulid worms, from Gr. πριάπος, ''priāpos'' 'Priapus' + Lat. ''-ul-'', diminutive), sometimes referred to as penis worms, is a phylum of unsegmented marine worms. The name of the phylum relates to the Greek god of fertility, because their general shape and their extensible spiny introvert (eversible) proboscis may resemble the shape of a human penis. They live in the mud and in comparatively shallow waters up to deep. Some species show a remarkable tolerance for hydrogen sulfide and anoxia. They can be quite abundant in some areas. In an Alaskan bay as many as 85 adult individuals of ''Priapulus caudatus'' per square meter has been recorded, while the density of its larvae can be as high as 58,000 per square meter. Together with Echiura and Sipuncula, they were once placed in the taxon Gephyrea, but consistent morphological and molecular evidence supports their belonging to Ecdysozoa, which also includes arthropods and nematodes. Fossil findings show th ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Stem-group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown ...
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Priapulus Caudatus
''Priapulus caudatus'' known as the cactus worm, is a marine invertebrate belonging to the phylum Priapulida. It is a cylindrical, unsegmented worm which burrows in soft sediment on the seabed. It has a circumpolar distribution. Taxonomy and evolution ''Priapulus caudatus'' is one of only nineteen known species in the phylum Priapulida. French naturalist Jean-Baptiste Lamarck first described it in 1816. Phylogenetic studies have indicated that scalidophorans, to which priapulids belong, are a basal clade of ecdysozoans (animals that grow by shedding their exoskeleton), and thus a sister group to all other ecdysozoans, an assortment including nematodes and arthropods. Priapulids were abundant and widespread in the Early Cambrian period and their tunnelling activities in soft sediment created many trace fossils. ''P. caudatus'' is likely to be very similar to the animals existing at that time. Description A cylindrical unsegmented worm, ''P. caudatus'' grows to a length of . The ...
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Echiura
The Echiura, or spoon worms, are a small group of marine animals. Once treated as a separate phylum, they are now considered to belong to Annelida. Annelids typically have their bodies divided into segments, but echiurans have secondarily lost their segmentation. The majority of echiurans live in burrows in soft sediment in shallow water, but some live in rock crevices or under boulders, and there are also deep sea forms. More than 230 species have been described. Spoon worms are cylindrical, soft-bodied animals usually possessing a non-retractable proboscis which can be rolled into a scoop-shape to feed. In some species the proboscis is ribbon-like, longer than the trunk and may have a forked tip. Spoon worms vary in size from less than a centimetre in length to more than a metre. Most are deposit feeders, collecting detritus from the sea floor. Fossils of these worms are seldom found and the earliest known fossil specimen is from the Upper Carboniferous (Pennsylvanian). T ...
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Respiratory Pigment
A respiratory pigment is a metalloprotein that serves a variety of important functions, its main being O2 transport. Other functions performed include O2 storage, CO2 transport, and transportation of substances other than respiratory gases. There are four major classifications of respiratory pigment: hemoglobin, hemocyanin, erythrocruorin– chlorocruorin, and hemerythrin. The heme-containing globin is the most commonly-occurring respiratory pigment, occurring in at least 9 different phyla of animals. Comparing Respiratory Pigments Hemoglobin, erythrocruorin, and chlorocruorin are all globins, iron-heme proteins with a common core. Their color comes from the absorption spectra of heme with Fe2+. Erythrocruorin and chlorocruorin are closely related giant globins found used by some invertebrates. Chlorocruorin has a special heme group, giving it different colors. Globins The globin is thought to be a very ancient molecule, even acting as a molecular clock of sorts. It has ...
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Scalidophora
Scalidophora is a group of marine pseudocoelomate protostomes that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous cuticle that is moulted, rings of scalids on the introvert, flosculi, and two rings of introvert retracts.Heiner, I., Kristensen, R.H. 2005. Two new species of the genus ''Pliciloricus'' (Loricifera, Pliciloricidae) from the Faroe Bank, North Atlantic. Zoologischer Anzeiger. 243: 121–138. However, the monophyly of the Scalidophora was not supported by two molecular studies, where the position of the Loricifera was uncertain or as sister to the Panarthropoda. Both studies supported a reduced Scalidophora comprising the Kinorhyncha and Priapulida as sister phyla. Their closest relatives are the Panarthropoda, Nematoda and Nematomorpha; thus, they are placed in the group Ecdysozoa. The two species in the genus '' Markuelia'', known fro ...
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Ecdysozoa
Ecdysozoa () is a group of protostome animals, including Arthropoda (insects, chelicerata, crustaceans, and myriapods), Nematoda, and several smaller phyla. They were first defined by Aguinaldo ''et al.'' in 1997, based mainly on phylogenetic trees constructed using 18S ribosomal RNA genes. A large study in 2008 by Dunn ''et al.'' strongly supported the Ecdysozoa as a clade, that is, a group consisting of a common ancestor and all its descendants. The group is also supported by morphological characters, and includes all animals that grow by ecdysis, moulting their cuticle. The group was initially contested by a significant minority of biologists. Some argued for groupings based on more traditional taxonomic techniques, while others contested the interpretation of the molecular data. Etymology The name ''Ecdysozoa'' stems etymologically . Characteristics The most notable characteristic shared by ecdysozoans is a three-layered cuticle (four in Tardigrada) composed ...
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Haemerythrin
Hemerythrin (also spelled haemerythrin; grc, αἷμα, haîma, blood, grc, ἐρυθρός, erythrós, red) is an oligomeric protein responsible for oxygen (O2) transport in the marine invertebrate phyla of sipunculids, priapulids, brachiopods, and in a single annelid worm genus, ''Magelona''. Myohemerythrin is a monomeric O2-binding protein found in the muscles of marine invertebrates. Hemerythrin and myohemerythrin are essentially colorless when deoxygenated, but turn a violet-pink in the oxygenated state. Hemerythrin does not, as the name might suggest, contain a heme. The names of the blood oxygen transporters hemoglobin, hemocyanin, hemerythrin, do not refer to the heme group (only found in globins), instead these names are derived from the Greek word for blood. Hemerythrin may also contribute to innate immunity and anterior tissue regeneration in certain worms. O2 binding mechanism The mechanism of dioxygen binding is unusual. Most O2 carriers operate via formation o ...
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Worm
Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete worms (bristle worms); for the African giant earthworm, '' Microchaetus rappi''; and for the marine nemertean worm (bootlace worm), '' Lineus longissimus''. Various types of worm occupy a small variety of parasitic niches, living inside the bodies of other animals. Free-living worm species do not live on land but instead live in marine or freshwater environments or underground by burrowing. In biology, "worm" refers to an obsolete taxon, '' vermes'', used by Carolus Linnaeus and Jean-Baptiste Lamarck for all non-arthropod invertebrate animals, now seen to be paraphyletic. The name stems from the Old English word '' wyrm''. Most animals called "worms" are invertebrates, but the term is also used for the amphibian caecilians and the slow ...
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Hydrogen Sulfide
Hydrogen sulfide is a chemical compound with the formula . It is a colorless chalcogen-hydride gas, and is poisonous, corrosive, and flammable, with trace amounts in ambient atmosphere having a characteristic foul odor of rotten eggs. The underground mine gas term for foul-smelling hydrogen sulfide-rich gas mixtures is ''stinkdamp''. Swedish chemist Carl Wilhelm Scheele is credited with having discovered the chemical composition of purified hydrogen sulfide in 1777. The British English spelling of this compound is hydrogen sulphide, a spelling no longer recommended by the Royal Society of Chemistry or the International Union of Pure and Applied Chemistry. Hydrogen sulfide is toxic to humans and most other animals by inhibiting cellular respiration in a manner similar to hydrogen cyanide. When it is inhaled or it or its salts are ingested in high amounts, damage to organs occurs rapidly with symptoms ranging from breathing difficulties to convulsions and death. Despite this, ...
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Polychaete
Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made of chitin. More than 10,000 species are described in this class. Common representatives include the lugworm (''Arenicola marina'') and the sandworm or clam worm ''Alitta''. Polychaetes as a class are robust and widespread, with species that live in the coldest ocean temperatures of the abyssal plain, to forms which tolerate the extremely high temperatures near hydrothermal vents. Polychaetes occur throughout the Earth's oceans at all depths, from forms that live as plankton near the surface, to a 2- to 3-cm specimen (still unclassified) observed by the robot ocean probe ''Nereus'' at the bottom of the Challenger Deep, the deepest known spot in the Earth's oceans. Only 168 species (less than 2% of all polychaetes) are known from ...
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Classification
Classification is a process related to categorization, the process in which ideas and objects are recognized, differentiated and understood. Classification is the grouping of related facts into classes. It may also refer to: Business, organizations, and economics * Classification of customers, for marketing (as in Master data management) or for profitability (e.g. by Activity-based costing) * Classified information, as in legal or government documentation * Job classification, as in job analysis * Standard Industrial Classification, economic activities Mathematics * Attribute-value system, a basic knowledge representation framework * Classification theorems in mathematics * Mathematical classification, grouping mathematical objects based on a property that all those objects share * Statistical classification, identifying to which of a set of categories a new observation belongs, on the basis of a training set of data Media * Classification (literature), a figure of spee ...
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