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Plasmodesmata
Plasmodesmata (singular: plasmodesma) are microscopic channels which traverse the cell walls of plant cells and some algal cells, enabling transport and communication between them. Plasmodesmata evolved independently in several lineages, and species that have these structures include members of the Charophyceae, Charales, Coleochaetales and Phaeophyceae (which are all algae), as well as all embryophytes, better known as land plants. Unlike animal cells, almost every plant cell is surrounded by a polysaccharide cell wall. Neighbouring plant cells are therefore separated by a pair of cell walls and the intervening middle lamella, forming an extracellular domain known as the apoplast. Although cell walls are permeable to small soluble proteins and other solutes, plasmodesmata enable direct, regulated, symplastic transport of substances between cells. There are two forms of plasmodesmata: primary plasmodesmata, which are formed during cell division, and secondary plasmodesma ...
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Plasmodesmata Structure
Plasmodesmata (singular: plasmodesma) are microscopic channels which traverse the cell walls of plant cells and some algal cells, enabling transport and communication between them. Plasmodesmata evolved independently in several lineages, and species that have these structures include members of the Charophyceae, Charales, Coleochaetales and Phaeophyceae (which are all algae), as well as all embryophytes, better known as land plants. Unlike animal cells, almost every plant cell is surrounded by a polysaccharide cell wall. Neighbouring plant cells are therefore separated by a pair of cell walls and the intervening middle lamella, forming an extracellular domain known as the apoplast. Although cell walls are permeable to small soluble proteins and other solutes, plasmodesmata enable direct, regulated, symplastic transport of substances between cells. There are two forms of plasmodesmata: primary plasmodesmata, which are formed during cell division, and secondary plasmodesmata, which c ...
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Pit (botany)
Pits are relatively thinner portions of the cell wall that adjacent cell (biology), cells can Cell communication, communicate or exchange fluid through. Pits are characteristic of cell walls with Cell wall#Rigidity of cell walls, secondary layers. Generally each pit has a complementary pit opposite of it in the neighboring cell. These complementary pits are called "pit pairs". Pits are composed of three parts: the pit chamber, the pit aperture, and the pit membrane. The pit chamber is the hollow area where the secondary layers of the cell wall are absent. The pit aperture is the opening at either end of the pit chamber. The pit membrane is the primary cell wall and middle lamella, or the membrane between adjacent cell walls, at the middle of the pit chamber. The primary cell wall at the pit membrane may also have depressions similar to the pit depressions of the secondary layers. These depressions are primary pit-fields, or primary pits. In the primary pit, the primordial pit pro ...
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Biological Sciences
Biology is the scientific study of life and living organisms. It is a broad natural science that encompasses a wide range of fields and unifying principles that explain the structure, function, growth, origin, evolution, and distribution of life. Central to biology are five fundamental themes: the cell as the basic unit of life, genes and heredity as the basis of inheritance, evolution as the driver of biological diversity, energy transformation for sustaining life processes, and the maintenance of internal stability (homeostasis). Biology examines life across multiple levels of organization, from molecules and cells to organisms, populations, and ecosystems. Subdisciplines include molecular biology, physiology, ecology, evolutionary biology, developmental biology, and systematics, among others. Each of these fields applies a range of methods to investigate biological phenomena, including observation, experimentation, and mathematical modeling. Modern biology is grounded in ...
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Cytosol
The cytosol, also known as cytoplasmic matrix or groundplasm, is one of the liquids found inside cells ( intracellular fluid (ICF)). It is separated into compartments by membranes. For example, the mitochondrial matrix separates the mitochondrion into many compartments. In the eukaryotic cell, the cytosol is surrounded by the cell membrane and is part of the cytoplasm, which also comprises the mitochondria, plastids, and other organelles (but not their internal fluids and structures); the cell nucleus is separate. The cytosol is thus a liquid matrix around the organelles. In prokaryotes, most of the chemical reactions of metabolism take place in the cytosol, while a few take place in membranes or in the periplasmic space. In eukaryotes, while many metabolic pathways still occur in the cytosol, others take place within organelles. The cytosol is a complex mixture of substances dissolved in water. Although water forms the large majority of the cytosol, its structure and proper ...
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Phospholipid Bilayer
The lipid bilayer (or phospholipid bilayer) is a thin polar membrane made of two layers of lipid molecules. These membranes form a continuous barrier around all cells. The cell membranes of almost all organisms and many viruses are made of a lipid bilayer, as are the nuclear membrane surrounding the cell nucleus, and membranes of the membrane-bound organelles in the cell. The lipid bilayer is the barrier that keeps ions, proteins and other molecules where they are needed and prevents them from diffusing into areas where they should not be. Lipid bilayers are ideally suited to this role, even though they are only a few nanometers in width, because they are impermeable to most water-soluble ( hydrophilic) molecules. Bilayers are particularly impermeable to ions, which allows cells to regulate salt concentrations and pH by transporting ions across their membranes using proteins called ion pumps. Biological bilayers are usually composed of amphiphilic phospholipids that hav ...
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Plasmalemma
The cell membrane (also known as the plasma membrane or cytoplasmic membrane, and historically referred to as the plasmalemma) is a biological membrane that separates and protects the cytoplasm, interior of a Cell (biology), cell from the extracellular space, outside environment (the extracellular space). The cell membrane consists of a lipid bilayer, made up of two layers of phospholipids with cholesterols (a lipid component) interspersed between them, maintaining appropriate membrane Membrane fluidity, fluidity at various temperatures. The membrane also contains membrane proteins, including integral proteins that span the membrane and serve as membrane transporters, and peripheral proteins that loosely attach to the outer (peripheral) side of the cell membrane, acting as enzymes to facilitate interaction with the cell's environment. Glycolipids embedded in the outer lipid layer serve a similar purpose. The cell membrane membrane transport, controls the movement of substances in ...
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Plasma Membrane
The cell membrane (also known as the plasma membrane or cytoplasmic membrane, and historically referred to as the plasmalemma) is a biological membrane that separates and protects the interior of a cell from the outside environment (the extracellular space). The cell membrane consists of a lipid bilayer, made up of two layers of phospholipids with cholesterols (a lipid component) interspersed between them, maintaining appropriate membrane fluidity at various temperatures. The membrane also contains membrane proteins, including integral proteins that span the membrane and serve as membrane transporters, and peripheral proteins that loosely attach to the outer (peripheral) side of the cell membrane, acting as enzymes to facilitate interaction with the cell's environment. Glycolipids embedded in the outer lipid layer serve a similar purpose. The cell membrane controls the movement of substances in and out of a cell, being selectively permeable to ions and organic molecu ...
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Membrane Transportation
A membrane is a selective barrier; it allows some things to pass through but stops others. Such things may be molecules, ions, or other small particles. Membranes can be generally classified into synthetic membranes and biological membranes. Biological membranes include cell membranes (outer coverings of cells or organelles that allow passage of certain constituents); nuclear membranes, which cover a cell nucleus; and tissue membranes, such as mucous membrane, mucosae and serous membrane, serosae. Synthetic membranes are made by humans for use in laboratory, laboratories and industry (such as chemical plants). This concept of a membrane has been known since the eighteenth century but was used little outside of the laboratory until the end of World War II. Drinking water supplies in Europe had been compromised by The War and membrane filters were used to test for water safety. However, due to the lack of reliability, slow operation, reduced selectivity and elevated costs, membran ...
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New Phytologist
''New Phytologist'' is a peer-reviewed scientific journal published on behalf of the New Phytologist Foundation by Wiley-Blackwell. It covers all aspects of botany and was established in 1902 by Arthur Tansley, who served as editor until 1931. Maarja Öpik took up the position of Editor-in-Chief of ''New Phytologist'' in January 2025. The previous Editor-in-Chief was Alistair M. Hetherington. Article categories The journal publishes articles in the following categories: * Original research articles * Priority reports * Research reviews * Commentaries * Letters * Meeting reports * Modelling/Theory and Methods papers * Tansley reviews * Tansley insights * Viewpoints * Community resources Abstracting and indexing The journal is abstracted and indexed in the Science Citation Index Expanded, Current Contents/Agriculture, Biology & Environmental Sciences, and Scopus. According to the ''Journal Citation Reports'', the journal has a 2023 impact factor The impact factor (IF) or ...
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Endoplasmic Reticulum
The endoplasmic reticulum (ER) is a part of a transportation system of the eukaryote, eukaryotic cell, and has many other important functions such as protein folding. The word endoplasmic means "within the cytoplasm", and reticulum is Latin for "little net". It is a type of organelle made up of two subunits – rough endoplasmic reticulum (RER), and smooth endoplasmic reticulum (SER). The endoplasmic reticulum is found in most eukaryotic cells and forms an interconnected network of flattened, membrane-enclosed sacs known as cisternae (in the RER), and tubular structures in the SER. The membranes of the ER are continuous with the outer nuclear membrane. The endoplasmic reticulum is not found in red blood cells, or spermatozoa. There are two types of ER that share many of the same proteins and engage in certain common activities such as the synthesis of certain lipids and cholesterol. Different types of Cell (biology), cells contain different ratios of the two types of ER dependin ...
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Plastid
A plastid is a membrane-bound organelle found in the Cell (biology), cells of plants, algae, and some other eukaryotic organisms. Plastids are considered to be intracellular endosymbiotic cyanobacteria. Examples of plastids include chloroplasts (used for photosynthesis); chromoplasts (used for synthesis and storage of pigments); leucoplasts (non-pigmented plastids, some of which can cellular differentiation, differentiate); and apicoplasts (non-photosynthetic plastids of apicomplexa derived from secondary endosymbiosis). A permanent primary endosymbiosis event occurred about 1.5 billion years ago in the Archaeplastida cladeEmbryophyte, land plants, red algae, green algae and glaucophytesprobably with a cyanobiont, a symbiotic cyanobacteria related to the genus ''Gloeomargarita lithophora, Gloeomargarita''. Another primary endosymbiosis event occurred later, between 140 and 90 million years ago, in the photosynthetic plastids ''Paulinella'' amoeboids of the cyanobacteria genera '' ...
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Stromule
A stromule is a microscopic structure found in plant cells. Stromules ( stroma-filled tubules) are highly dynamic structures extending from the surface of all plastid types, including proplastids, chloroplasts, etioplasts, leucoplasts, amyloplasts, and chromoplasts. Protrusions from and interconnections between plastids were observed in 1888 ( Gottlieb Haberlandt) and 1908 ( Gustav Senn) and have been described sporadically in the literature since then. Stromules were recently rediscovered in 1997 and have since been reported to exist in a number of angiosperm species including ''Arabidopsis thaliana'', wheat, rice and tomato, but their role is not yet fully understood. This highly dynamic nature is caused by the close relationship between plastid stromules and actin microfilaments, which are anchored to the stromule extensions, either in a longitudinal fashion to pull from the stromule and guide the plastid in a given direction or in a hinge fashion allowing the plastid to rest ...
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