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Paraneoptera
Paraneoptera or Acercaria is a Order (biology), superorder of insects which includes Psocodea, lice (Psocoptera, bark lice and Louse, true lice), thrips, and hemipterans, the true bugs. It also includes the extinct order Permopsocida, known from fossils dating from the Cisuralian, Early Permian to the mid-Cretaceous. All of the insects classified here exhibit various “reductions” or “simplifications” from the primitive body-plan found in typical polyneopterans. Cercus, Cerci, for example, are entirely absent in all living paraneopterans (Acercaria meaning without cerci). Other “reductions” occur in wing venation, in the number of tarsal segments (no more than three), only four Malpighian tubules, and only one complex of abdominal ganglia. The mouthparts of the Paraneoptera reflect diverse feeding habits. Basal groups are microbial surface feeders, whereas more advanced groups feed on plant or animal fluids. Phylogeny Paraneoptera consists of Psocodea (lice), along ...
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Psocodea
Psocodea is a taxonomic group of insects comprising the bark lice, book lice and parasitic lice. It was formerly considered a superorder, but is now generally considered by entomologists as an order. Despite the greatly differing appearance of parasitic lice (Phthiraptera), they are believed to have evolved from within the former order Psocoptera, which contained the bark lice and book lice, now found to be paraphyletic. They are often regarded as the most primitive of the hemipteroids. Psocodea contains around 11,000 species, divided among four suborders and more than 70 families. They range in size from 1–10 millimetres (0.04–0.4 in) in length. The species known as booklice received their common name because they are commonly found amongst old books—they feed upon the paste used in binding. The barklice are found on trees, feeding on algae and lichen. Anatomy and biology Psocids are small, scavenging insects with a relatively generalized body plan. They feed ...
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Neoptera
Neoptera (Ancient Greek ''néos'' ("new") + ''pterón'' ("wing")) is a classification group that includes most orders of the winged insects, specifically those that can flex their wings over their abdomens. This is in contrast with the more basal orders of winged insects (the " Palaeoptera" assemblage), which are unable to flex their wings in this way. Classification The taxon Neoptera was proposed by А.М. Martynov in 1923 and 1924, in the following classification: Pterygota *division Palaeoptera **order Odonata **order Agnatha (correct name: Ephemeroptera) **†order Dictyoneuridea **†order Megasecoptera **†order Meganisoptera **†order Protephemeroidea *division Neoptera **superorder Endopterygota ***order Coleoptera ***order Strepsiptera ***order Neuroptera ***order Raphidioptera ***order Megaloptera ***order Diptera ***order Mecoptera ***order Trichoptera ***order Lepidoptera ***order Hymenoptera **subdivision Polyneoptera ***superorder Orthopteroid ...
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Permopsocida
Permopsocida is an extinct order of insects known from the Carboniferous ( Moscovian) to the Mid-Cretaceous. It is part of Paraneoptera, alongside bark lice (including lice), bugs and thrips. Within Paraneoptera Paraneoptera or Acercaria is a Order (biology), superorder of insects which includes Psocodea, lice (Psocoptera, bark lice and Louse, true lice), thrips, and hemipterans, the true bugs. It also includes the extinct order Permopsocida, known from ... it is considered to be closer to the clade containing bugs and thrips rather than bark lice, with an estimated divergence during the Late Carboniferous. The group was first named as a suborder by Robert John Tillyard in 1926, and was raised to a full order by Huang et al. in 2016. It is currently divided up into three families, Psocidiidae which is known from the Late Carboniferous to Liassic. Permopsocidae which is only known from the Permian, and Archipsyllidae, which is known from the Late Triassic to mid-Cretaceous (C ...
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Hemiptera
Hemiptera (; ) is an order of insects, commonly called true bugs, comprising more than 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from to around , and share a common arrangement of piercing-sucking mouthparts. The name "true bugs" is sometimes limited to the suborder Heteroptera. Entomologists reserve the term ''bug'' for Hemiptera or Heteroptera,Gilbert Waldbauer. ''The Handy Bug Answer Book.'' Visible Ink, 1998p. 1. which does not include other arthropods or insects of other orders such as ants, bees, beetles, or butterflies. In some varieties of English, all terrestrial arthropods (including non-insect arachnids and myriapods) also fall under the colloquial understanding of ''bug''. Many insects with "bug" in their common name, especially in American English, belong to other orders; for example, the lovebug is a fly and the Maybug and ladybug are beetles. ...
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Insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, Thorax (insect anatomy), thorax and abdomen (insect anatomy), abdomen), three pairs of jointed Arthropod leg, legs, compound eyes, and a pair of antenna (biology), antennae. Insects are the most diverse group of animals, with more than a million described species; they represent more than half of all animal species. The insect nervous system consists of a insect brain, brain and a ventral nerve cord. Most insects reproduce Oviparous, by laying eggs. Insects Respiratory system of insects, breathe air through a system of Spiracle (arthropods), paired openings along their sides, connected to Trachea#Invertebrates, small tubes that take air directly to the tissues. The blood therefore does not carry oxygen; it is only partly contained in ves ...
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Eumetabola
Eumetabola is an unranked clade of Neoptera. Two large unities known as the Eumetabola and Paurometabola are probably from the adelphotaxa of the Neoptera after exclusion of the Plecoptera. The monophyly of these unities appears to be weakly justified. According to a phylogenetic analysis, the Eumetabola clade originated 390–350 million years ago, in the Late Devonian. Phylogeny The phylogeny of Eumetabola is shown in the cladogram, using the molecular phylogeny of Wipfler et al. 2019 for the Polyneoptera, Johnson et al 2018 for the Paraneoptera (where Psocomorpha contains Phthiraptera), and Kjer et al 2016 for the Holometabola Holometabola (from Ancient Greek "complete" + "change"), also known as Endopterygota (from "inner" + "wing" + Neo-Latin "-having"), is a supra-order (biology), ordinal clade of insects within the infraclass Neoptera that go through distincti .... References External links Eumetabola Hennig 1953''insecta.bio.spbu.ru'' * Grimaldi, D.; En ...
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Condylognatha
Condylognatha or Panhemiptera is a monophyletic grouping (superorder) that contains Hemiptera (true bugs) and Thysanoptera (thrips). Condylognatha belongs to Paraneoptera, which include its sister group, lice (Psocodea). Taxonomy Hemiptera and Thysanoptera are both the sister-groups based on morphological characters, and jointly known as Condylognatha. Hemiptera Hemiptera is an order of insects most often known as the ''true bugs'' (''cf.'' bug), comprising around 50,000–80,000 species of cicadas, aphids, planthoppers, leafhoppers, shield bugs, bed bugs and others. They range in size from to around , and share a common arrangement of sucking mouthparts. Thrips Order Thysanoptera includes 5,500 species classified into two suborders distinguished by the ovipositor. Terebrantia have a well-developed conical ovipositor, while the Tubulifera Phlaeothripidae is a family of thrips with hundreds of genera. They are the only extant family of the suborder Tubulifera, alo ...
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Polyneoptera
The cohort Polyneoptera is one of the major groups of winged insects, comprising the Orthoptera (grasshoppers, crickets, etc.) and all other neopteran insects believed to be more closely related to Orthoptera than to any other insect orders. They were formerly grouped together with the Palaeoptera and Paraneoptera as the Hemimetabola or Exopterygota on the grounds that they have no pupa, the wings gradually developing externally throughout the nymphal stages. Many members of the group have leathery forewings ( tegmina) and hindwings with an enlarged anal field (vannus). When Carl Linnaeus started applying binomial names to animals in the 10th edition of his ''Systema Naturae'' in 1758, there were few animals included in the scheme, and consequently few groups. As more and more new species were discovered and differences recognised, the original groups proposed by Linnaeus were split up. Originally all polyneopteran insects were in the genus ''Gryllus'', this genus now con ...
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Psocoptera
Psocoptera () are a paraphyletic group of insects that are commonly known as booklice, barklice or barkflies. The name Psocoptera has been replaced with Psocodea in recent literature, with the inclusion of the former order Phthiraptera into Psocodea (as part of the suborder Troctomorpha). They are often regarded as the most primitive of the paraneopterans. There are more than 5,500 species in 41 families in three suborders. Many of these species have only been described in the early twenty-first century. They range in size from in length. The species known as booklice received their common name because they are commonly found amongst old books—they feed upon the paste used in binding. The barklice are found on trees, feeding on algae and lichen. Etymology Their name originates from the Greek word ψῶχος (''psokhos''), meaning " gnawed" or " rubbed" and πτερά (''ptera''), meaning " wings". Classification In the 2000s, morphological and molecular phylogenetic evid ...
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Sister Clade
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomic ...
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Phthiraptera
Louse (: lice) is the common name for any member of the infraorder Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera was previously recognized as an order, until a 2021 genetic study determined that they are a highly modified lineage of the order Psocodea, whose members are commonly known as booklice, barklice or barkflies. Lice are obligate parasites, living externally on warm-blooded hosts, which include every species of bird and mammal, except for monotremes, pangolins, and bats. Chewing lice live among the hairs or feathers of their host and feed on skin and debris, whereas sucking lice pierce the host's skin and feed on blood and other secretions. They usually spend their whole life on a single host, cementing their eggs, called nits, to hairs or feathers. The eggs hatch into nymphs, which moult three times before becoming fully grown, a process that takes about four weeks. Humans host two species of louse—the head louse an ...
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