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Macronaria
Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located (hence ''macro''- meaning large, and –''naria'' meaning nose). Fossil evidence suggests that macronarian dinosaurs lived from the Middle Jurassic (Bathonian) through the Late Cretaceous (Maastrichtian). Macronarians have been found globally, including discoveries in Argentina, the United States, Portugal, China, and Tanzania. Like other sauropods, they are known to have inhabited primarily terrestrial areas, and little evidence exists to suggest that they spent much time in coastal environments. Macronarians are diagnosed through their distinct characters on their skulls, as well as appendicular and vertebral characters. Macronaria is composed of several subclades and families notably including Camarasauridae and Titanosauriformes, among severa ...
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Brachiosauridae
The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the leaves of tall trees that other sauropods would have been unable to reach. In addition, they possessed thick spoon-shaped teeth which helped them to consume tough plants more efficiently than other sauropods. They have also been characterized by a few unique traits or synapomorphies; dorsal vertebrae with 'rod-like' transverse processes and an ischium with an abbreviated pubic peduncle. ''Brachiosaurus'' is one of the best-known members of the Brachiosauridae, and was once thought to be the largest land animal to ever live. Brachiosaurids thrived in the regions which are now North and South America, Africa, Europe, and Asia. They first appear in the fossil record in the Late Jurassic Period (possibly even earlier in the Middle Jurassic) ...
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Camarasauridae
Camarasauridae is a family of sauropod dinosaurs. Among sauropods, camarasaurids are small to medium-sized, with relatively short necks. They are visually identifiable by a short skull with large nares, and broad, spatulate teeth filling a thick jaw.1. Weishampel, D. B., Dodson, P., & Osmólska, H. (Eds.). (1990). ''The dinosauria''. Univ of California Press. Based on cervical vertebrae and cervical rib biomechanics, camarasaurids most likely moved their necks in a vertical, rather than horizontal, sweeping motion, in contrast to most diplodocids. Taxonomy Camarasauridae was named by Edward Drinker Cope in 1877. Its type genus is ''Camarasaurus'', and it is defined as the clade containing all species more closely related to ''Camarasaurus supremus'' than ''Saltasaurus loricatus''. Phylogenetic relationships Camarasauridae is typically regarded as belonging to Macronaria, one of the two major branches of Neosauropoda. Within Macronaria, it occupies a basal position, outsi ...
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Haestasaurus
''Haestasaurus'' is a genus of herbivorous sauropod dinosaur, belonging to the Macronaria, that during the Early Cretaceous lived in the area of present-day England. The only species is ''Haestasaurus becklesii''.Upchurch P., Mannion P.D., Taylor M.P., 2015, "The Anatomy and Phylogenetic Relationships of “''Pelorosaurus''“ ''becklesii'' (Neosauropoda, Macronaria) from the Early Cretaceous of England", ''PLoS ONE'' 10(6): e0125819. Description As a sauropod, ''Haestasaurus'' would have been a large quadrupedal long-necked dinosaur. Little information is available about the specifics of its build because only a forelimb is known of the animal. An indication of the size of ''Haestasaurus'' is given by the length of the forelimb elements. The humerus is 599 millimetres long, the ulna 421 millimetres and the radius, situated next to the ulna in the lower arm, has a length of 404 millimetres. A 2015 study found several unique anatomical traits (autapomorphies) distinguishing ...
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Janenschia
''Janenschia'' (named after Werner Janensch) is a large herbivorous sauropod dinosaur from the Late Jurassic Tendaguru Formation of Lindi Region, Tanzania around 155 million years ago. Discovery and naming ''Janenschia'' has had a convoluted nomenclatural history. In 1907, Eberhard Fraas at "site P", nine hundred metres to the southeast of Tendaguru Hill, discovered two skeletons of gigantic sauropods. They were designated as "Skeleton A" and "Skeleton B". The fossils were transported to the collection of the '' Stuttgarter Naturaliensammlung'' in Stuttgart, Germany. Fraas in 1908 decided to name both skeletons as different species of one genus, ''Gigantosaurus''. Skeleton A became ''Gigantosaurus africanus'' and skeleton B became ''Gigantosaurus robustus''. The latter species was based on the holotype partial skeleton SMNS 12144, consisting of a right hindlimb. The specific name was inspired by the heavy build of the animal. While doing so, Fraas knew full well that the ...
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Pukyongosaurus
''Pukyongosaurus'' (meaning "Pukyong lizard", after the Pukyong National University) is a genus of titanosauriform dinosaur that lived in South Korea during the Early Cretaceous period ( Aptian - Albian). It may have been closely related to ''Euhelopus'', and is known from a series of vertebrae in the neck and back. The characteristics that were originally used to distinguish this genus have been criticized as being either widespread or too poorly preserved to evaluate, rendering the genus an indeterminate nomen dubium among titanosauriforms. The 2022 study noted that ''Pukyongosaurus'' is probably a somphospondylan. Discovery In 2001, several fragments of a sauropod skeleton were discovered in the Hasandong Formation in Hadong County, South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korean Peninsula and sharing a land border with North Korea. Its western border is formed by the Yellow Sea ...
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Jiutaisaurus
''Jiutaisaurus'' (meaning "Jiutai lizard") is a genus of sauropod dinosaur from the Quantou Formation of Jilin, China. ''Jiutaisaurus'' was a sauropod which lived during the Cretaceous. The type species, ''Jiutaisaurus xidiensis'', was described by Wu ''et al.'' in 2006, and is based on eighteen vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...e. It probably lived alongside '' Changchunsaurus'' and '' Helioceratops''. References Macronarians Cretaceous dinosaurs of Asia Fossil taxa described in 2006 {{Sauropodomorph-stub ...
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Sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', ' lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their body), and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include ''Brachiosaurus'', ''Diplodocus'', ''Apatosaurus'' and ''Brontosaurus''. The oldest known unequivocal sauropod dinosaurs are known from the Early Jurassic. '' Isanosaurus'' and '' Antetonitrus'' were originally described as Triassic sauropods, but their age, and in the case of ''Antetonitrus'' also its sauropod status, were subsequently questioned. Sauropod-like sauropodomorph tracks from the Fleming Fjord Formation (Greenland) might, however, indicate the occurrence of the group in the Late Triassic. By the Late Jurassic (150 mill ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, ar ...
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Somphospondyli
Somphospondyli is an extinct clade of titanosauriform sauropods that lived from the Late Jurassic until the end of the Late Cretaceous, comprising all titanosauriforms more closely related to Titanosauria proper than Brachiosauridae. The remains of somphospondylans have been discovered in all six continents. Classification The group has officially been defined under the PhyloCode as the largest clade containing ''Saltasaurus loricatus'', but not ''Giraffatitan brancai.'' Features found as diagnostic of this clade by Mannion ''et al.'' (2013) include the possession of at least 15 cervical vertebrae; a bevelled radius bone end; sacral vertebrae with camellate internal texture; convex posterior articular surfaces of middle to posterior caudal vertebrae; biconvex distal caudal vertebrae; humerus anterolateral corner "squared"; among multiple others. The following cladogram depicts the reference phylogeny used to defined Somphospondlyi under the PhyloCode The ''International Code ...
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Fukuititan
''Fukuititan'' (meaning " Fukui giant") is a genus of titanosauriform sauropod dinosaur that lived in the Early Cretaceous (either Barremian or Aptian age) in what is now Japan. It is known from FPDM-V8468, the associated partial skeleton of a single individual, recovered from the Kitadani Dinosaur Quarry (Kitadani Formation) of the Tetori Group, at Katsuyama City. The type species, ''Fukuititan nipponensis'', was described in 2010 by Japanese scientists Yoichi Azuma and Masateru Shibata of the Fukui Prefectural Dinosaur Museum The , located in Katsuyama, Fukui, Japan, is one of the leading dinosaur museums in Asia that is renowned for its exhibits of fossil specimens of dinosaurs and paleontological research. It is sited in the Nagaoyama Park ( Katsuyama Dinosaur Fore .... The discovery sheds light on Japanese titanosauriforms, which are poorly known in the region. References Early Cretaceous dinosaurs of Asia Fossil taxa described in 2010 Macronarians Fossils o ...
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Bashunosaurus
''Bashunosaurus'' is a genus of potentially macronarian sauropod dinosaur from the Middle Jurassic Shaximiao Formation of Kaijiang, China. The type and only species is ''Bashunosaurus kaijiangensis''. Discovery The holotype of ''Bashunosaurus'', KM 20100, was discovered in sediments of the lower Shaximiao Formation in Maanping, Sichuan Province, China. It consists of six cervical vertebrae, eight dorsal vertebrae, a partial left scapula, and a right humerus, ulna, ilium, femur, tibia, and fibula. An additional right ilium, specimen KM 20103, was assigned as a paratype. History The name "Bashunosaurus kaijiangensis" first appeared in Ouyang's description of '' Abrosaurus'' in 1989, although without a description or diagnosis, making it a '' nomen nudum'' (i.e. a nickname that is unavailable for use as an actual scientific name). Li ''et al.'' (1999), however, attribute the naming to "Kuang, 1996", still considering it a ''nomen nudum''. Although it was finally formally ...
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Marmarospondylus
''Marmarospondylus'' ("marble eference to the Forest Marble Formation">Forest_Marble_Formation.html" ;"title="eference to the Forest Marble Formation">eference to the Forest Marble Formationvertebra") is a nomen dubium, dubious genus of sauropod dinosaur from Middle Jurassic deposits in the English Midlands. The type species, ''Marmarospondylus robustus'', was described by Richard Owen as a species of the Late Jurassic genus '' Bothriospondylus'' in 1875. The holotype, NHMUK R.22428, a dorsal, was found in the Bathonian-age Forest Marble Formation The Forest Marble is a geological formation in England. Part of the Great Oolite Group, it dates to the late Bathonian stage of the Middle Jurassic.Weishampel, David B; et al. (2004). "Dinosaur distribution (Middle Jurassic, Europe)." In: Weisha ... at Bradford-on-Avon, Wiltshire. Owen himself in an addendum to the same publication coined ''Marmarospondylus'' for ''B. robustus'', presumably due to its being older than ''B. suffossu ...
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