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Maniraptora
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Dromaeosauridae, Troodontidae, Oviraptorosauria, and Therizinosauria. ''Ornitholestes'' and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period (see ''Eshanosaurus''), and origin of birds#Phylogeny, survive today as living birds. Description Maniraptorans are characterized by long arms and three-fingered hands (though reduced or fused in some lineages), as well as a "half-moon shaped" (semilunar bone, semi-lunate) bone in the wrist (Carpal bones, carpus). In 2004, Tom Holtz and Halszka Osmólska pointed out six other maniraptoran characteristics relating to specific de ...
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Therizinosauria
Therizinosaurs (; once called segnosaurs) are an extinct group of large herbivorous Theropoda, theropod dinosaurs whose fossils have been mainly discovered from Cretaceous deposits in Asia and North America. Potential fragmentary remains have also been found in Jurassic deposits of Asia and Europe. Various features of the forelimbs, skull and pelvis unite these finds as both theropods and maniraptorans, making them relatives of birds. The name of the representative genus, ''Therizinosaurus'', is derived from the Ancient Greek , Greek (, 'to reap' or 'scythe')Translated paper
and (, 'lizard'). The older representative, ''Segnosaurus'', is derived from the Latin ('slow') and the Greek .


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Nothronychus
''Nothronychus'' (meaning "slothful claw") is a genus of therizinosaurid theropod dinosaurs that lived in North America during the Late Cretaceous period. The type species, ''Nothronychus mckinleyi'', was described by James Kirkland and Douglas G. Wolfe in 2001. It was recovered near New Mexico's border with Arizona, in an area known as the Zuni Basin, from rocks assigned to the Moreno Hill Formation, dating to the Turonian stage of the Late Cretaceous. A second specimen, described in 2009 as a second species, ''Nothronychus graffami'', was found in the Tropic Shale of Utah, dating between one million and a half million years older than ''N. mckinleyi''. ''Nothronychus'' were bulky herbivorous theropods with wide, sloth-like hip (resembling that of the non-related ornithischians), four-toed feet with all four toes facing forward, elongated necks and prominent arms with sharp claws. Both species were similar in dimensions, but ''N. graffami'' was slightly more robust than ''N. ...
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Maniraptoriformes
Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrannosauroids, ornithomimosaurs, maniraptorans, and over the recent years, megaraptorans (Although position within the clade is unclear). Maniraptora includes birds, the only known dinosaur group alive today. In the past, Coelurosauria was used to refer to all small theropods, but this classification has since been amended. Most feathered dinosaurs discovered so far have been coelurosaurs. Philip J. Currie had considered it likely and probable that all coelurosaurs were feathered. However, several skin impressions found for some members of this group show pebbly, scaly skin, indicating that feathers did not completely replace scales in all taxa. Anatomy Bodyplan The studying of anatomical traits in coelurosaurs indicates that the last c ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutionary history, evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, Evolution of birds, having evolved from earlier Theropoda, theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomy (biology), taxonomic, ...
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Coelurosauria
Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrannosauroids, ornithomimosaurs, maniraptorans, and over the recent years, megaraptorans (Although position within the clade is unclear). Maniraptora includes birds, the only known dinosaur group alive today. In the past, Coelurosauria was used to refer to all small theropods, but this classification has since been amended. Most feathered dinosaurs discovered so far have been coelurosaurs. Philip J. Currie had considered it likely and probable that all coelurosaurs were feathered. However, several skin impressions found for some members of this group show pebbly, scaly skin, indicating that feathers did not completely replace scales in all taxa. Anatomy Bodyplan The studying of anatomical traits in coelurosaurs indicates that the last c ...
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Fukuivenator
''Fukuivenator'' ("hunter of Fukui Prefecture") is an extinct genus of maniraptoran theropod dinosaur from the Early Cretaceous of Japan. Discovery and naming The type species ''Fukuivenator paradoxus'' was named and described in 2016 by Yoichi Azuma, Xu Xing (paleontologist), Xu Xing, Masateru Shibata, Soichiro Kawabe, Kazunori Miyata and Takuya Imai. The generic name combines a reference to the Fukui prefecture with Latin ''venator'', "hunter". The specific name (zoology), specific name refers to the paradoxical combination of traits shown by the species. The rocks in which the skeleton of ''Fukuivenator'', holotype FPDM-V8461, was found in August 2007 belong to the Kitadani Formation, which is probably of Barremian or Aptian age. Radiometric dating of nearby rock units has given this formation an estimated age of somewhere between 127 and 115 million years old. The holotype consists of a partial skeleton with skull. The skeleton of ''F. paradoxus'' is currently the most c ...
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Bird
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a four-chambered heart, and a strong yet lightweight Bird skeleton, skeleton. Birds live worldwide and range in size from the bee hummingbird to the common ostrich. There are over 11,000 living species and they are split into 44 Order (biology), orders. More than half are passerine or "perching" birds. Birds have Bird wing, wings whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the Flightless bird, loss of flight in some birds, including ratites, penguins, and diverse endemism, endemic island species. The digestive and respiratory systems of birds are also uniquely a ...
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Paraves
Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosauridae, dromaeosaurids, troodontidae, troodontids, Anchiornithidae, anchiornithids, and possibly the scansoriopterygidae, scansoriopterygids, the group also contains the Avialae, avialans, which include diverse extinct taxa as well as the over 11,000 species of living birds. Basal members of Paraves are well known for the possession of an enlarged claw on the second digit of the foot, which was held off the ground when walking in some species. A number of differing scientific interpretations of the relationships between paravian taxa exist. New fossil discoveries and analyses make the classification of Paraves an active subject of research. Description Like other theropods, all paravians are bipedal, walking on their two hind legs. The teeth of basal paravians were curved and serrated, but not blade-like except in some specialized species, suc ...
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Alvarezsauroidea
Alvarezsauroidea (from the Argentine historian, writer and physician Gregorio Álvarez) is a group of small maniraptoran dinosaurs. The group was first formally proposed by Choiniere and colleagues in 2010, to contain the family Alvarezsauridae and non-alvarezsaurid alvarezsauroids, such as '' Haplocheirus'',Choiniere, J.N., Xu, X., Clark, J.M., Forster, C.A., Guo, Y. and Han, F. (2010). "A basal alvarezsauroid theropod from the early Late Jurassic of Xinjiang, China." ''Science'', 327: 571-574. which is the basalmost of the Alvarezsauroidea (from the Late Jurassic, Asia). The discovery of '' Haplocheirus'' extended the stratigraphic evidence for the group Alvarezsauroidea about 63 million years further in the past. The division of Alvarezsauroidea into the Alvarezsauridae and the non-alvarezsaurid alvarezsauroids is based on differences in their morphology, especially in their hand morphology. Description Fossils of alvarezsauroids were described since the 1990s, with many a ...
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Ornitholestes
''Ornitholestes'' (meaning "bird robber") is a small theropod dinosaur of the late Jurassic ( Brushy Basin Member of the Morrison Formation, middle Kimmeridgian age, about 154 million years agoTurner, C.E. and Peterson, F., (1999). "Biostratigraphy of dinosaurs in the Upper Jurassic Morrison Formation of the Western Interior, U.S.A." Pp. 77–114 in Gillette, D.D. (ed.), ''Vertebrate Paleontology in Utah''. Utah Geological Survey Miscellaneous Publication 99-1.) of Western Laurasia (the area that was to become North America). To date, ''Ornitholestes'' is known only from a single partial skeleton with a badly crushed skull found at the Bone Cabin Quarry near Medicine Bow, Wyoming, in 1900. It was described by Henry Fairfield Osborn in 1903. An incomplete hand was later attributed to ''Ornitholestes'', although it now appears to belong to '' Tanycolagreus''. The type (and only known) species is ''O. hermanni''. The specific name honors the American Museum of Natural History prepa ...
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Yaverlandia
''Yaverlandia'' (meaning "of Yaverland Point/Yaverland Battery") is a genus of maniraptoran dinosaur. Known from a partial fossil skull (MIWG 1530) found in Lower Cretaceous strata of the Wessex Formation ( Upper Silty Bed; Vectis Formation) on the Isle of Wight. it was described as the earliest known member of the pachycephalosaurid family, but research by Darren Naish shows it to have actually been a theropod, seemingly a maniraptoran. The type species is ''Y. bitholus''.Galton, P.M. (1971) "A primitive dome-headed dinosaur (Ornithischia: Pachycephalosauridae) from the Lower Cretaceous of England and the function of the dome of pachycephalosaurids". ''Journal of Paleontology'', Vol. 45, No. 1, Pages 40–47. Discovery and naming MIWG 1530, the holotype skull, was discovered in 1930, in England and were commented upon by Watson (1930). It was referred to as an iguanodontid of the genus ''Vectisaurus'' in 1936. When Steel (1969) followed Hulke (1879) in listing ''Vectisaurus'' ...
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Migmanychion
''Migmanychion'' (meaning "claw mixture") is an extinct genus of maniraptoran theropod dinosaur from the Early Cretaceous Longjiang Formation of China. The genus contains a single species, ''M. laiyang'', known from an incomplete left forelimb. Discovery and naming The ''Migmanychion'' holotype specimen, LY 2022JZ3001, was discovered in sediments of the Longjiang Formation (Pigeon Hill locality), near Baoshan in Morin Dawa Daur Autonomous Banner, Inner Mongolia, China. The specimen consists of rib fragments and a partial left arm with a complete hand, preserved on a slab and counterslab. In 2023, Wang ''et al''. described ''Migmanychion laiyang'' as a new genus and species of maniraptoran theropod based on these fossil remains. The generic name, "''Migmanychion''", combines the Greek words "μείγμα νυχιών", meaning "claw mixture", referencing the difference in the morphology of the manual unguals. The specific name, "''laiyang''", references the holotype's ...
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