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Lophophorate
The Lophophorata are a Lophotrochozoan clade consisting of the Brachiozoa and the Bryozoa. They have a lophophore. Molecular phylogenetic analyses suggest that lophophorates are protostomes Protostomia () is the clade of animals once thought to be characterized by the formation of the organism's mouth before its anus during embryonic development. This nature has since been discovered to be extremely variable among Protostomia's me ..., but on morphological grounds they have been assessed as deuterostomes. Fossil finds of a segmented worm named Wufengella suggest that they evolved from a worm close to annelids. References {{Protostome-stub ...
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Lophophore
The lophophore () is a characteristic feeding organ possessed by four major groups of animals: the Brachiopoda, Bryozoa, Hyolitha, and Phoronida, which collectively constitute the protostome group Lophophorata.Introduction to the Lophotrochozoa
– Retrieved 3 May 2010
All lophophores are found in aquatic organisms.


Etymology

''Lophophore'' is derived from the Greek ''lophos'' (crest, tuft) and ''-phore'', ''-phoros'' (φορος) (bearing), a derivative of ''phérein'' (φέρειν) (to bear); thus crest-bearing.


Characteristics

The lophophore can most easily be described as a ring of ciliated tentacles surrounding the mouth, but it is often horseshoe-shaped or coiled. Phoronids have thei ...
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Bryozoa
Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869living species are known. At least two genera are solitary (''Aethozooides'' and ''Monobryozoon''); the rest are colonial. The terms Polyzoa and Bryozoa were introduced in 1830 and 1831, respectively. Soon after it was named, another group of animals was discovered whose filtering mechanism looked similar, so it was included in Bryozoa until 1869, when the two groups wer ...
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Ectoprocta
Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869living species are known. At least two genera are solitary (''Aethozooides'' and ''Monobryozoon''); the rest are colonial. The terms Polyzoa and Bryozoa were introduced in 1830 and 1831, respectively. Soon after it was named, another group of animals was discovered whose filtering mechanism looked similar, so it was included in Bryozoa until 1869, when the two groups we ...
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Tommotiida
Tommotiids are an extinct group of Cambrian invertebrates thought to be early lophophorates (the group containing Bryozoa, Brachiopoda, and Phoronida). The majority of tommotiids are mineralised with calcium phosphate rather than calcium carbonate. although silicified examples hint that some species bore carbonate or carbonaceous sclerites. ''Micrina'' and '' Paterimitra'' possess bivalved shells in their larval phases, which preserve characters that might position them in the Linguliformea and Rhynchonelliformea stem lineages respectively. This would indicate that the brachiopod shell represents the retention of a larval character. For a long part of their history, the tommotiids were only known from disarticulated shells - a complete organism had not been found. The 2008 discovery of ''Eccentrotheca'' offered the first insight into a complete organism, and permitted a reconstruction of the animal as a sessile, tube-like animal made up of a spiral of overlapping plates. ...
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Wufengella
''Wufengella'' is a genus of extinct camenellan " tommotiid" that lived during the Early Cambrian ( Stage 3). Described in 2022, the only species ''Wufengella bengtsonii'' was discovered from the Maotianshan Shales of Chiungchussu (Qiongzhusi) Formation in Yunnan, China. The fossil indicates that the animal was an armoured worm that close to the common ancestry of the phyla Phonorida, Brachiozoa and Bryozoa, which are collectively grouped into a clade called Lophophorata. Discovery ''Wufengella'' is known from a single specimen. The fossil was discovered by Chinese palaeontologists Jin Guo and Peiyun Cong at the Yunnan University. An almost complete fossil, parts of the anterior end are missing. The location of the specimen, Chiungchussu Formation at Haikou, Kunming, Southwest China, is member of the Chengjian Lagerstätte that is established to belong to Cambrian Stage 3 (between 521 and 514 million year ago). The same fossil deposit had yielded worm-like lobopod ''Facive ...
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Lophotrochozoa
Lophotrochozoa (, "crest/wheel animals") is a clade of protostome animals within the Spiralia. The taxon was established as a monophyletic group based on molecular evidence. The clade includes animals like annelids, molluscs, bryozoans, brachiopods, and platyhelminthes. Groups Lophotrochozoa was defined in 1995 as the "last common ancestor of the three traditional lophophorate taxa (brachiopods, bryozoans, and phoronid worms), the mollusks and the annelids, and all of the descendants of that common ancestor". It is a cladistic definition (a node-based name), so the affiliation to Lophotrochozoa of spiralian groups not mentioned directly in the definition depends on the topology of the spiralian tree of life, and in some phylogenetic hypotheses, Lophotrochozoa may even be synonymous to Spiralia. Nemertea and Orthonectida (if not directly considered as part of Annelida) are probably lophotrochozoan phyla; Dicyemida, Gastrotricha, and Platyhelminthes may be lophotrochozoa ...
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Brachiozoa
Brachiozoa is a grouping of lophophorate animals including Brachiopoda and Phoronida. It also includes their ancestors, the extinct tommotiids. References Lophophorata Protostome unranked clades {{Protostome-stub ...
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Deuterostome
Deuterostomia (; in Greek) are animals typically characterized by their anus forming before their mouth during embryonic development. The group's sister clade is Protostomia, animals whose digestive tract development is more varied. Some examples of deuterostomes include vertebrates (and thus humans), sea stars, and crinoids. In deuterostomy, the developing embryo's first opening (the blastopore) becomes the anus, while the mouth is formed at a different site later on. This was initially the group's distinguishing characteristic, but deuterostomy has since been discovered among protostomes as well. This group is also known as enterocoelomates, because their coelom develops through enterocoely. The three major clades of deuterostomes are Chordata (e.g. vertebrates), Echinodermata (e.g. starfish), and Hemichordata (e.g. acorn worms). Together with Protostomia and their out-group Xenacoelomorpha, these compose the Bilateria, animals with bilateral symmetry and th ...
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Entoprocta
Entoprocta (), or Kamptozoa , is a phylum of mostly sessile aquatic animals, ranging from long. Mature individuals are goblet-shaped, on relatively long stalks. They have a "crown" of solid tentacles whose cilia generate water currents that draw food particles towards the mouth, and both the mouth and anus lie inside the "crown". The superficially similar Bryozoa (Ectoprocta) have the anus outside a "crown" of hollow tentacles. Most families of entoprocts are colonial, and all but 2 of the 150 species are marine. A few solitary species can move slowly. Some species eject unfertilized ova into the water, while others keep their ova in brood chambers until they hatch, and some of these species use placenta-like organs to nourish the developing eggs. After hatching, the larvae swim for a short time and then settle on a surface. There they metamorphose, and the larval gut rotates by up to 180°, so that the mouth and anus face upwards. Both colonial and solitary species a ...
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Cycliophora
''Symbion'' is a genus of commensal aquatic animals, less than 0.5 mm wide, found living attached to the mouthparts of cold-water lobsters. They have sac-like bodies, and three distinctly different forms in different parts of their two-stage life-cycle. They appear so different from other animals that they were assigned their own, new phylum Cycliophora shortly after they were discovered in 1995. This was the first new phylum of multicelled organism to be discovered since the Loricifera in 1983. Taxonomy ''Symbion'' was discovered in 1995 by Reinhardt Kristensen and Peter Funch on the mouthparts of the Norway lobster (''Nephrops norvegicus''). Other, related, species have since been discovered on: * the American lobster (''Homarus americanus'', host to ''Symbion americanus'') * the European lobster (''Homarus gammarus'', host to an as yet unnamed species of ''Symbion'') The genus is so named because of its commensal relationship with the lobster (a form of symbio ...
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