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Halteres
''Halteres'' (; singular ''halter'' or ''haltere'') (from grc, ἁλτῆρες, weights held in the hands to give an impetus in leaping) are a pair of small club-shaped organs on the body of two orders of flying insects that provide information about body rotations during flight. Insects of the large order Diptera (flies) have halteres which evolved from a pair of ancestral hindwings, while males of the much smaller order Strepsiptera (stylops)Merriam-Webster: stylops broadly: an insect of the order Strepsiptera/ref> have halteres which evolved from a pair of ancestral forewings. Halteres oscillate rapidly along with the wings and operate like vibrating structure gyroscopes: any rotation of the plane of oscillation causes a force on the vibrating halteres by the Coriolis effect. The insect detects this force with sensory organs called campaniform sensilla and chordotonal organs located at the base of the halteres and uses this information to interpret and correct its ...
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Diptera
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies and others, although only about 125,000 species have been described. Flies have a mobile head, with a pair of large compound eyes, and mouthparts designed for piercing and sucking (mosquitoes, black flies and robber flies), or for lapping and sucking in the other groups. Their wing arrangement gives them great maneuverability in flight, and claws and pads on their feet enable them to cling to smooth surfaces. Flies undergo complete metamorphosis; the eggs are often laid on the ...
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Coriolis Effect
In physics, the Coriolis force is an inertial or fictitious force that acts on objects in motion within a frame of reference that rotates with respect to an inertial frame. In a reference frame with clockwise rotation, the force acts to the left of the motion of the object. In one with anticlockwise (or counterclockwise) rotation, the force acts to the right. Deflection of an object due to the Coriolis force is called the Coriolis effect. Though recognized previously by others, the mathematical expression for the Coriolis force appeared in an 1835 paper by French scientist Gaspard-Gustave de Coriolis, in connection with the theory of water wheels. Early in the 20th century, the term ''Coriolis force'' began to be used in connection with meteorology. Newton's laws of motion describe the motion of an object in an inertial (non-accelerating) frame of reference. When Newton's laws are transformed to a rotating frame of reference, the Coriolis and centrifugal accelerations appea ...
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Coriolis Effect
In physics, the Coriolis force is an inertial or fictitious force that acts on objects in motion within a frame of reference that rotates with respect to an inertial frame. In a reference frame with clockwise rotation, the force acts to the left of the motion of the object. In one with anticlockwise (or counterclockwise) rotation, the force acts to the right. Deflection of an object due to the Coriolis force is called the Coriolis effect. Though recognized previously by others, the mathematical expression for the Coriolis force appeared in an 1835 paper by French scientist Gaspard-Gustave de Coriolis, in connection with the theory of water wheels. Early in the 20th century, the term ''Coriolis force'' began to be used in connection with meteorology. Newton's laws of motion describe the motion of an object in an inertial (non-accelerating) frame of reference. When Newton's laws are transformed to a rotating frame of reference, the Coriolis and centrifugal accelerations appea ...
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Campaniform Sensilla
Campaniform sensilla are a class of mechanoreceptors found in insects, which respond to local stress and strain within the animal's cuticle. Campaniform sensilla function as proprioceptors that detect mechanical load as resistance to muscle contraction, similar to mammalian Golgi tendon organs. Sensory feedback from campaniform sensilla is integrated in the control of posture and locomotion. Structure Each campaniform sensillum consists of a flexible dome, which is embedded in a spongy socket within the cuticle and innervated by the dendrites of a single bipolar sensory neuron (see schematic cross-section). Campaniform sensilla are often oval-shaped with long axes of about 5-10 µm (see SEM). Campaniform sensilla are distributed across the body surface of many insects. The fruit fly ''Drosophila melanogaster'', for example, has over 680 sensilla. Campaniform sensilla are located in regions where stress is likely to be high, including on the legs, antennae, wings, and ha ...
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Strepsiptera
The Strepsiptera are an order of insects with eleven extant families that include about 600 described species. They are endoparasites in other insects, such as bees, wasps, leafhoppers, silverfish, and cockroaches. Females of most species never emerge from the host after entering its body, finally dying inside it. The early-stage larvae do emerge because they must find an unoccupied living host, and the short-lived males must emerge to seek a receptive female in her host. They are believed to be most closely related to beetles, from which they diverged 300–350 million years ago, but do not appear in the fossil record until the mid-Cretaceous around 100 million years ago. The order is not well known to non-specialists, and the nearest they have to a common name is stylops. The name of the order translates to "twisted wing"', giving rise to other common names used for the order, twisted-wing insects and twisted-winged parasites. Adult males are rarely observed, although specimen ...
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Strepsiptera
The Strepsiptera are an order of insects with eleven extant families that include about 600 described species. They are endoparasites in other insects, such as bees, wasps, leafhoppers, silverfish, and cockroaches. Females of most species never emerge from the host after entering its body, finally dying inside it. The early-stage larvae do emerge because they must find an unoccupied living host, and the short-lived males must emerge to seek a receptive female in her host. They are believed to be most closely related to beetles, from which they diverged 300–350 million years ago, but do not appear in the fossil record until the mid-Cretaceous around 100 million years ago. The order is not well known to non-specialists, and the nearest they have to a common name is stylops. The name of the order translates to "twisted wing"', giving rise to other common names used for the order, twisted-wing insects and twisted-winged parasites. Adult males are rarely observed, although specimen ...
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Hox Gene
Hox genes, a subset of homeobox genes, are a group of related genes that specify regions of the body plan of an embryo along the head-tail axis of animals. Hox proteins encode and specify the characteristics of 'position', ensuring that the correct structures form in the correct places of the body. For example, Hox genes in insects specify which appendages form on a segment (for example, legs, antennae, and wings in fruit flies), and Hox genes in vertebrates specify the types and shape of vertebrae that will form. In segmented animals, Hox proteins thus confer segmental or positional identity, but do not form the actual segments themselves. Studies on Hox genes in ciliated larvae have shown they are only expressed in future adult tissues. In larvae with gradual metamorphosis the Hox genes are activated in tissues of the larval body, generally in the trunk region, that will be maintained through metamorphosis. In larvae with complete metamorphosis the Hox genes are mainly express ...
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Hindwing
Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments. The wings are strengthened by a number of longitudinal veins, which often have cross-connections that form closed "cells" in the membrane (extreme examples include the dragonflies and lacewings). The patterns resulting from the fusion and cross-connection of the wing veins are often diagnostic for different evolutionary lineages and can be used for identification to the family or even genus level in many orders of insects. Physically, some insects move their flight muscles directly, others indirectly. In insects with direct flight, the wing muscles directly attach to the wing base, so that a small downward movement of the wing base lifts the wing itself upward. Thos ...
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Forewing
Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments. The wings are strengthened by a number of longitudinal veins, which often have cross-connections that form closed "cells" in the membrane (extreme examples include the dragonflies and lacewings). The patterns resulting from the fusion and cross-connection of the wing veins are often diagnostic for different evolutionary lineages and can be used for identification to the family or even genus level in many orders of insects. Physically, some insects move their flight muscles directly, others indirectly. In insects with direct flight, the wing muscles directly attach to the wing base, so that a small downward movement of the wing base lifts the wing itself upward. Those ...
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Vibrating Structure Gyroscope
A vibrating structure gyroscope, defined by the IEEE as a Coriolis vibratory gyroscope (CVG), is a gyroscope that uses a vibrating structure to determine the rate of rotation. A vibrating structure gyroscope functions much like the halteres of flies (insects in the order Diptera). The underlying physical principle is that a vibrating object tends to continue vibrating in the same plane even if its support rotates. The Coriolis effect causes the object to exert a force on its support, and by measuring this force the rate of rotation can be determined. Vibrating structure gyroscopes are simpler and cheaper than conventional rotating gyroscopes of similar accuracy. Inexpensive vibrating structure gyroscopes manufactured with MEMS technology are widely used in smartphones, gaming devices, cameras and many other applications. Theory of operation Consider two proof masses vibrating in plane (as in the MEMS gyro) at frequency \omega_r. The Coriolis effect induces an acceleration ...
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Antenna (biology)
Antennae ( antenna), sometimes referred to as "feelers", are paired appendages used for Sensory system, sensing in arthropods. Antennae are connected to the first one or two Segmentation (biology), segments of the arthropod head. They vary widely in form but are always made of one or more jointed segments. While they are typically sensory organs, the exact nature of what they sense and how they sense it is not the same in all groups. Functions may variously include sensing tactition, touch, air motion, heat, vibration (sound), and especially insect olfaction, smell or gustation, taste. Antennae are sometimes modified for other purposes, such as mating, brooding, swimming, and even anchoring the arthropod to a substrate (biology), substrate. Larval arthropods have antennae that differ from those of the adult. Many crustaceans, for example, have free-swimming larvae that use their antennae for swimming. Antennae can also locate other group members if the insect lives in a group, lik ...
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Chordotonal Organ
Chordotonal organs are stretch receptor organs found only in insects and crustaceans. They are located at most joints and are made up of clusters of scolopidia that either directly or indirectly connect two joints and sense their movements relative to one another. They can have both extero- and proprioceptive functions, for example sensing auditory stimuli or leg movement. The word was coined by Vitus Graber in 1882, though he interpreted them as being stretched between two points like a string, sensing vibrations through resonance. Structure Chordotonal organs can be composed of a single scolopidium with only a single sensory, bipolar neuron (such as the tympanal ear of a notodontid moth), or up to several thousand scolopidia, each equipped with up to four sensory neurons (as in the mosquito Johnston's organ). The bipolar sensory neurons each have an apical dendritic structure with a cilium densely packed with microtubules and surrounded by two specialized cells, the sc ...
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