Hyaenodont
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Hyaenodont
Hyaenodonta ("hyena teeth") is an extinct order of hypercarnivorous placental pan-carnivoran mammals from mirorder Ferae. Hyaenodonts were important mammalian predators that arose during the early Paleocene in Europe and persisted well into the late Miocene. Characteristics Hyaenodonts are characterized by long skulls, slender jaws, slim bodies and a plantigrade stance. They generally ranged in size from 30 to 140 cm at the shoulder. While '' Simbakubwa kutokaafrika'' may have been up to (surpassing the modern polar bear in size) and ''Hyaenodon gigas'' (the largest species from genus ''Hyaenodon'') was as much as 1.4 m high at the shoulder, 3.0 m long and weighed about 330 kg, most of hyaenodonts were in the 5–15 kg range, equivalent to a mid-sized dog. The anatomy of their skulls show that they had a particularly acute sense of smell, while their teeth were adapted for shearing, rather than crushing. Because of their size range, it is probable that diffe ...
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Hyaenodon Horridus Skull
''Hyaenodon'' ("hyena-tooth") is an extinct genus of carnivorous hyaenodont mammals from tribe Hyaenodontini, within subfamily Hyaenodontinae in family Hyaenodontidae,Malcolm C. McKenna, Susan K. Bell (1997."Classification of Mammals: Above the Species Level" Columbia University Press, New York, 631 pages. that lived in Eurasia and North America during the middle Eocene to early Miocene, existing for about 25.1 million years. Description Typical of early carnivorous mammals, individuals of ''Hyaenodon'' had a very massive skull, but only a small brain. The skull is long with a narrow snout - much larger in relation to the length of the skull than in canine carnivores, for instance. The neck was shorter than the skull, while the body was long and robust and terminated in a long tail. Compared to the generally larger (but not closely related) ''Hyainailouros'', the dentition of ''Hyaenodon'' was geared more towards shearing meat and less towards bone crushing. Some species of ...
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Hyaenodon
''Hyaenodon'' (" hyena-tooth") is an extinct genus of carnivorous hyaenodont mammals from tribe Hyaenodontini, within subfamily Hyaenodontinae in family Hyaenodontidae,Malcolm C. McKenna, Susan K. Bell (1997."Classification of Mammals: Above the Species Level" Columbia University Press, New York, 631 pages. that lived in Eurasia and North America during the middle Eocene to early Miocene, existing for about 25.1 million years. Description Typical of early carnivorous mammals, individuals of ''Hyaenodon'' had a very massive skull, but only a small brain. The skull is long with a narrow snout - much larger in relation to the length of the skull than in canine carnivores, for instance. The neck was shorter than the skull, while the body was long and robust and terminated in a long tail. Compared to the generally larger (but not closely related) ''Hyainailouros'', the dentition of ''Hyaenodon'' was geared more towards shearing meat and less towards bone crushing. Some specie ...
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Megistotherium
''Megistotherium'' ("greatest beast") is an extinct genus of hyainailourid hyaenodonts from paraphyletic subfamily Hyainailourinae that lived in Africa. Etymology The name of this genus comes and . The name of species ''Megistotherium osteothlastes'' comes and (with ''-es'' being an agent noun: 'bone-crusher'). Description ''Megistotherium osteothlastes'' is the only known species from this genus, and it was a large hyainailourid that lived during the early Miocene. Its remains have been found in the Ngorora and Muruyur Formations of Kenya, Egypt,Morlo, M., Miller, E.R., and El-Barkooky, A.N. 2007. Creodonta and Carnivora from Wadi Moghra, Egypt. Journal of Vertebrate Paleontology 27: 145–159. Namibia, UgandaMorales, J. and M. Pickford. (2008). "Creodonts and carnivores from the Middle Miocene Muruyur Formation at Kipsaraman and Cheparawa, Baringo District, Kenya." ''Comptes Rendus Palevol'' 7 (8): 487-497 and Libya. Named by Robert Savage in 1973, ''Megistotherium'' ...
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Apterodontinae
Apterodontinae ("without winged tooth") is an extinct subfamily of hyainailourid hyaenodonts that lived in Africa and Europe during the late Eocene to middle Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the .... Classification and phylogeny Taxonomy * Subfamily: †Apterodontinae ** Genus: †'' Apterodon'' *** †''Apterodon altidens'' *** †''Apterodon gaudryi'' *** †''Apterodon langebadreae'' *** †''Apterodon macrognathus'' *** †''Apterodon rauenbergensis'' *** †''Apterodon saghensis'' *** †''Apterodon'' sp. ur At-Talah escarpment, Libya ** Genus: †'' Quasiapterodon'' *** †''Quasiapterodon minutus'' Phylogeny The phylogenetic relationships of subfamily Apterodontinae are shown in the following cladogram: References {{Taxonbar, from=Q34514 ...
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Exiguodon
''Exiguodon'' ("strict tooth") is an extinct genus of hyainailourid hyaenodont mammal of the polyphyletic tribe Hyainailourini within paraphyletic subfamily Hyainailourinae. Remains are known from early Miocene deposits in Kenya and Uganda, in East Africa. Description ''Exiguodon'' is distinguished from other hyainailourines by the following features: diminutive dimensions, lower molars (m3 – m2) with greatly reduced talonid; protoconid and paraconid similar in size. Paraconid of the molars lingually oriented. M2 and M1 close in size and morphology. Occlusal outline sub-triangular, with greatly reduced protocone, which appears like an antero-lingual cingulum which extends anteriorly and buccally. Presence of a strong parastyle in an antero-buccal position, united to the apex of the paracone by a well defined crista. The buccal cingulum borders a wide buccal platform, particularly large in the M2. Paracone tall and narrow, elongated blade-like metastyle. P4 broadened, with redu ...
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Hyainailouros
''Hyainailouros'' ("hyena-cat") is an extinct polyphyletic genus of hyainailourid hyaenodont mammal of the polyphyletic tribe Hyainailourini within paraphyletic subfamily Hyainailourinae, that lived during the early to middle Miocene, of which there were at least three species spread across Europe, Africa, and Asia. Closely related to other large African hyaenodonts such as ''Simbakubwa'' and ''Megistotherium'', ''Hyainailouros'' walked with a semi-digitigrade stance and was probably capable of large, leaping bounds. Alongside its African relatives and the last members of the genus ''Hyaenodon ''Hyaenodon'' (" hyena-tooth") is an extinct genus of carnivorous hyaenodont mammals from tribe Hyaenodontini, within subfamily Hyaenodontinae in family Hyaenodontidae,Malcolm C. McKenna, Susan K. Bell (1997."Classification of Mammals: Above t ...'' from Asia, ''Hyainailouros'' was among the largest hyaenodonts that existed. Phylogeny The phylogenetic relationships of genus ''Hyai ...
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Simbakubwa
''Simbakubwa'' ("great lion") is an extinct genus of hyainailourid hyaenodonts from paraphyletic subfamily Hyainailourinae that lived in Kenya during the early Miocene. Etymology The name of this genus comes from Swahili language for "great lion". Species name ''Simbakubwa kutokaafrika'' means "great lion of Africa". Description Different regression models produce a wide range of body mass estimates for ''Simbakubwa kutokaafrika'': from a low estimate of , comparable to the largest lions, to an upper estimate possibly reaching up to , which would surpass the modern polar bear in size. The fossils of ''Simbakubwa'' were first discovered by rural Kenyans at Meswa Bridge, Western Kenya. Thereafter Matthew Borths and Nancy Stevens published the findings after examining the fossils which had been stored at the Nairobi National Museum in Kenya for decades. The type specimen consists of a mandible from the lower jaw, a right upper maxilla and some postcranial remains. The light wear ...
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Sectisodon
''Sectisodon'' ("cutting tooth") is an extinct genus of hyainailourid hyaenodont mammal of the polyphyletic tribe Hyainailourini within paraphyletic subfamily Hyainailourinae from early Oligocene to early Miocene deposits in Egypt and Uganda. Description Morales and Pickford (2017, p. 346) diagnose ''Sectisodon'' as follows: "Hyainailourinae of small dimensions, M1 and M2 of similar length. M2 with paracone and metacone fused together, tall and pointed. Protocone very reduced, flattened and extending basally beyond the base of the paracone. Anterior cingulum with a well-defined parastyle with a moderate buccal cingulum. M2 and M1 with the metastyle quite a bit longer than the paracone-metacone. P3 short, with tall main cusp, posterior cusplet moderate, and much reduced protocone. Basal lingual cingulum strong, weaker on the buccal side. Lower molars with protoconid bigger than the metaconid, m2 with talonid present, much reduced in the m3." Classification and phylogeny Ta ...
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Pan-Carnivora
Ferae ( , , "wild beasts") is a mirorder of placental mammalsMalcolm C. McKenna, Susan K. Bell: ''Classification of Mammals: Above the Species Level'' in Columbia University Press, New York (1997), 631 Seiten. that groups together clades Pan-Carnivora and Pholidotamorpha. The Ferae is a sister group to the clade Pan-Euungulata and together they make grandorder Ferungulata. Classification and phylogeny Taxonomy : Phylogeny The phylogenetic relationships of mirorder Ferae are shown in the following cladogram, reconstructed from mitochondrial and nuclear DNA and protein characters, as well as the fossil record.O’Leary, M. A., Bloch JI, Flynn, J. J., Gaudin, T. J., Giallombardo, A., Giannini, N. P., Goldber, S. L, Kraatz, B. P., Luo, Z-X, Jin Meng, Xijun Ni, Novacek, M. J., Perini, F. A., Randall, Z. S., Rougier, G. W., Sargis, E. J., Silcox, M. T., Simmons, N. B., Spaulding, M. Velazco, P. M., Weksler, M., Wible, J. R. Cirranello, A. L. (2013."The Placental Mammal Ancestor an ...
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Ferae
Ferae ( , , "wild beasts") is a mirorder of placental mammalsMalcolm C. McKenna, Susan K. Bell: ''Classification of Mammals: Above the Species Level'' in Columbia University Press, New York (1997), 631 Seiten. that groups together clades Pan-Carnivora and Pholidotamorpha. The Ferae is a sister group to the clade Pan-Euungulata and together they make grandorder Ferungulata. Classification and phylogeny Taxonomy : Phylogeny The phylogenetic relationships of mirorder Ferae are shown in the following cladogram, reconstructed from mitochondrial and nuclear DNA and protein characters, as well as the fossil record.O’Leary, M. A., Bloch JI, Flynn, J. J., Gaudin, T. J., Giallombardo, A., Giannini, N. P., Goldber, S. L, Kraatz, B. P., Luo, Z-X, Jin Meng, Xijun Ni, Novacek, M. J., Perini, F. A., Randall, Z. S., Rougier, G. W., Sargis, E. J., Silcox, M. T., Simmons, N. B., Spaulding, M. Velazco, P. M., Weksler, M., Wible, J. R. Cirranello, A. L. (2013."The Placental Mammal Ancestor an ...
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Carnivora
Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are formally referred to as carnivorans, and have evolved to specialize in eating flesh. The order is the fifth largest order of mammals, comprising at least 279 species. Carnivorans live on every major landmass and in a variety of habitats, ranging from the cold polar regions to the hyper-arid region of the Sahara Desert to the open seas. They come in a very large array of different body plans in contrasting shapes and sizes. Carnivora can be divided into two subclades: the cat-like Feliformia and the dog-like Caniformia, which are differentiated based on the structure of their ear bones and cranial features. The feliforms include families such as the felidae, cats, the hyenas, the mongooses and the viverridae, civets. The majority of felifor ...
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Carnassial
Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified fourth upper premolar and the first lower molar, however this may vary in family. These teeth are also referred to as sectorial teeth. Taxonomy The name carnivoran is applied to a member of the order Carnivora. Carnivorans possess a common arrangement of teeth called carnassials, in which the first lower molar and the last upper premolar possess blade-like enamel crowns that act similar to a pair of shears for cutting meat. This dental arrangement has been modified by adaptation over the past 60 million years for diets composed of meat, for crushing vegetation, or for the loss of the carnassial function altogether as in seals, sea lions, and walruses. Carnassial dentition Carnassial teeth are modified molars (and in the case of carn ...
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