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Fish Jaw
Most bony fishes have two sets of jaws made mainly of bone. The primary oral jaws open and close the mouth, and a second set of pharyngeal jaws are positioned at the back of the throat. The oral jaws are used to capture and manipulate prey by biting and crushing. The pharyngeal jaws, so-called because they are positioned within the pharynx, are used to further process the food and move it from the mouth to the stomach. Cartilaginous fishes, such as sharks and Batoidea, rays, have one set of oral jaws made mainly of cartilage. They do not have pharyngeal jaws. Generally jaws are Articulation (anatomy), articulated and oppose vertically, comprising an upper jaw and a lower jaw and can bear numerous ordered teeth. Cartilaginous fishes grow multiple sets ''(polyphyodont)'' and replace teeth as they wear by moving new teeth laterally from the medial jaw surface in a conveyor-belt fashion. Teeth are replaced multiple times also in most bony fishes, but unlike cartilaginous fishes, the ...
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Skull
The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent parts: the neurocranium and the facial skeleton, which evolved from the first pharyngeal arch. The skull forms the frontmost portion of the axial skeleton and is a product of cephalization and vesicular enlargement of the brain, with several special senses structures such as the eyes, ears, nose, tongue and, in fish, specialized tactile organs such as barbels near the mouth. The skull is composed of three types of bone: cranial bones, facial bones and ossicles, which is made up of a number of fused flat and irregular bones. The cranial bones are joined at firm fibrous junctions called sutures and contains many foramina, fossae, processes, and sinuses. In zoology, the openings in the skull are called fenestrae, the most ...
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Pharyngeal Arch
The pharyngeal arches, also known as visceral arches'','' are transient structures seen in the Animal embryonic development, embryonic development of humans and other vertebrates, that are recognisable precursors for many structures. In fish, the arches support the Fish gill, gills and are known as the branchial arches, or gill arches. In the human embryo, the arches are first seen during the fourth week of human embryonic development, development. They appear as a series of outpouchings of mesoderm on both sides of the developing pharynx. The vasculature of the pharyngeal arches are the aortic arches that arise from the aortic sac. Structure In humans and other vertebrates, the pharyngeal arches are derived from all three germ layers (the primary layers of cells that form during embryonic development). Neural crest cells enter these arches where they contribute to features of the skull and facial skeleton such as bone and cartilage. However, the existence of pharyngeal structu ...
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Four-bar Linkage
In the study of Mechanism (engineering), mechanisms, a four-bar linkage, also called a four-bar, is the simplest closed-Kinematic chain, chain movable linkage (mechanical), linkage. It consists of four Rigid body, bodies, called ''bars'' or ''links'', connected in a loop by four Kinematic pair, joints. Generally, the joints are configured so the links move in parallel planes, and the assembly is called a ''planar four-bar linkage''. Spherical and spatial four-bar linkages also exist and are used in practice. Planar four-bar linkage Planar four-bar linkages are constructed from four links connected in a loop by four one-degrees of freedom (mechanics), degree-of-freedom joints. A joint may be either a revolute joint – also known as a pin joint or hinged joint – denoted by R, or a prismatic joint – also known as a sliding pair – denoted by P. A link that is fixed in place relative to the viewer is called a ''ground link.'' A link connecting to the ground by a revolute joint ...
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Aquatic Predation
Aquatic feeding mechanisms face a special difficulty as compared to feeding on land, because the density of water is about the same as that of the prey, so the prey tends to be pushed away when the mouth is closed. This problem was first identified by Robert McNeill Alexander. As a result, underwater predators, especially bony fish, have evolved a number of specialized feeding mechanisms, such as filter feeding, ram feeding, suction feeding, protrusion, and pivot feeding. Most underwater predators combine more than one of these basic principles. For example, a typical generalized predator, such as the cod, combines suction with some amount of protrusion and pivot feeding. Suction feeding Suction feeding is a method of ingesting a prey item in fluids by sucking the prey into the predator's mouth. It is a highly coordinated behavior achieved by the dorsal rotation of the dermatocranium, lateral expansion of the suspensorium, and the depression of the lower jaw and hyoid. Suct ...
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Evolution
Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, resulting in certain characteristics becoming more or less common within a population over successive generations. The process of evolution has given rise to biodiversity at every level of biological organisation. The scientific theory of evolution by natural selection was conceived independently by two British naturalists, Charles Darwin and Alfred Russel Wallace, in the mid-19th century as an explanation for why organisms are adapted to their physical and biological environments. The theory was first set out in detail in Darwin's book ''On the Origin of Species''. Evolution by natural selection is established by observable facts about living organisms: (1) more offspring are often produced than can possibly survive; (2) phenotypic variatio ...
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Wrasses
The wrasses are a family, Labridae, of marine ray-finned fish, many of which are brightly colored. The family is large and diverse, with over 600 species in 81 genera, which are divided into nine subgroups or tribes. They are typically small, most of them less than long, although the largest, the humphead wrasse, can measure up to . They are efficient carnivores, feeding on a wide range of small invertebrates. Many smaller wrasses follow the feeding trails of larger fish, picking up invertebrates disturbed by their passing. Juveniles of some representatives of the genera ''Bodianus'', '' Epibulus'', '' Cirrhilabrus'', '' Oxycheilinus'', and '' Paracheilinus'' hide among the tentacles of the free-living mushroom corals and '' Heliofungia actiniformis''. Etymology The word "wrasse" comes from the Cornish word ''wragh'', a lenited form of ''gwragh'', meaning an old woman or hag, via Cornish dialect ''wrath''. It is related to the Welsh ''gwrach'' and Breton ''gwrac'h''. T ...
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Bony Fishes
Osteichthyes ( ; ), also known as osteichthyans or commonly referred to as the bony fish, is a diverse clade of vertebrate animals that have endoskeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes (cartilaginous fish) and the extinct placoderms and acanthodians, which have endoskeletons primarily composed of cartilage. The vast majority of extant fish are members of Osteichthyes, being an extremely diverse and abundant group consisting of 45 orders, over 435 families and 28,000 species. The group is divided into two main clades, the ray-finned fish (Actinopterygii, which makes up the vast majority of extant fish) and the lobe-finned fish (Sarcopterygii, which gave rise to all land vertebrates, i.e. tetrapods). The oldest known fossils of bony fish are about 425 million years old from the late Silurian, which are also transitional fossils showing a tooth pattern that is in between the tooth rows of sharks and true bony fishes. Desp ...
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Linkage (mechanical)
A mechanical linkage is an assembly of systems connected so as to manage forces and Motion, movement. The movement of a body, or link, is studied using geometry so the link is considered to be Rigid body, rigid. The connections between links are modeled as providing ideal movement, pure rotation or Sliding (motion), sliding for example, and are called joints. A linkage modeled as a network of rigid links and ideal joints is called a kinematic chain. Linkages may be constructed from open chains, closed chains, or a combination of open and closed chains. Each link in a chain is connected by a joint to one or more other links. Thus, a kinematic chain can be modeled as a graph in which the links are paths and the joints are vertices, which is called a linkage graph. The movement of an ideal joint is generally associated with a subgroup of the group of Euclidean geometry, Euclidean displacements. The number of parameters in the subgroup is called the degree of freedom (mecha ...
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Agnatha
Agnatha (; ) or jawless fish is a paraphyletic infraphylum of animals in the subphylum Vertebrata of the phylum Chordata, characterized by the lack of jaws. The group consists of both extant taxon, living (Cyclostomi, cyclostomes such as hagfishes and lampreys) and Extinction, extinct clades (e.g. conodonts and Cephalaspidomorphi, cephalaspidomorphs, among others). They are sister taxon, sister to vertebrates with jaws known as gnathostomes, who evolution, evolved from jawless ancestors during the early Silurian by developing folding joint, articulations in the first pairs of gill arches. Sequencing, Molecular data, both from rRNA and from mtDNA as well as Embryology, embryological data, strongly supports the hypothesis that both groups of living agnathans, hagfishes and lampreys, are more closely related to each other than to Gnathostomata, jawed fish, forming the Class (biology), superclass Cyclostomi. The oldest fossil agnathans appeared in the Cambrian. Living jawless fish c ...
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Buccal Pump
Buccal pumping (/ˈbʌk.əl/...) is "breathing with one's cheeks": a method of ventilation used in respiration in which the animal moves the floor of its mouth in a rhythmic manner that is externally apparent.Brainerd, E. L. (1999). New perspectives on the evolution of lung ventilation mechanisms in vertebrates. Experimental Biology Online 4, 11-28. http://www.brown.edu/Departments/EEB/brainerd_lab/pdf/Brainerd-1999-EBO.pdf It is the sole means of inflating the lungs in amphibians. There are two methods of buccal pumping, defined by the number of movements of the floor of the mouth needed to complete both inspiration and expiration. Four stroke Four-stroke buccal pumping is used by some basal ray-finned fish and aquatic amphibians such as ''Xenopus'' and '' Amphiuma''. This method has several stages. These will be described for an animal starting with lungs in a deflated state: First, the glottis (opening to the lungs) is closed, and the nostrils are opened. The floor of the mo ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) ...
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Spiny Shark
Acanthodii or acanthodians is an extinct class of gnathostomes (jawed fishes). They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans (bony fish) and chondrichthyans (cartilaginous fish). In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians ( gars, bowfins). The popular name "spiny sharks" is because they were superficially shark-shaped, with a streamlined body, paired fins, a strongly upturned tail, and stout, largely immovable bony spines supporting all the fins except the tail—hence, "spiny sharks". However, acanthodians are not true sharks; their close relation to modern cartilaginous fish can lead them to be considered "stem-sharks". Acanthodians had a cartilaginous skeleton, but their f ...
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