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Ectomycorrhizal Extramatrical Mycelium
Ectomycorrhizal extramatrical mycelium (also known as extraradical mycelium) is the collection of filamentous fungal hyphae emanating from ectomycorrhizas. It may be composed of fine, hydrophilic hypha which branches frequently to explore and exploit the soil matrix or may aggregate to form rhizomorphs; highly differentiated, hydrophobic, enduring, transport structures. Characteristics Overview Apart from mycorrhizas, extramatrical mycelium is the primary vegetative body of ectomycorrhizal fungi. It is the location of mineral acquisition, enzyme production, and a key means of colonizing new root tips. Extramatrical mycelium facilitates the movement of carbon into the rhizosphere, moves carbon and nutrients between hosts and is an important food source for invertebrates. Exploration type The mycelial growth pattern, extent of biomass accumulation, and the presence or absence of rhizomorphs are used to classify fungi by exploration type. Agerer first proposed the desi ...
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Ascomycetes
Ascomycota is a phylum of the kingdom Fungi that, together with the Basidiomycota, forms the subkingdom Dikarya. Its members are commonly known as the sac fungi or ascomycetes. It is the largest phylum of Fungi, with over 64,000 species. The defining feature of this fungal group is the "ascus" (), a microscopic sexual structure in which nonmotile spores, called ascospores, are formed. However, some species of the Ascomycota are asexual, meaning that they do not have a sexual cycle and thus do not form asci or ascospores. Familiar examples of sac fungi include morels, truffles, brewers' and bakers' yeast, dead man's fingers, and cup fungi. The fungal symbionts in the majority of lichens (loosely termed "ascolichens") such as '' Cladonia'' belong to the Ascomycota. Ascomycota is a monophyletic group (it contains all descendants of one common ancestor). Previously placed in the Deuteromycota along with asexual species from other fungal taxa, asexual (or anamorphic) ascomycetes ...
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Xylem
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from the Ancient Greek word (''xylon''), meaning "wood"; the best-known xylem tissue is wood, though it is found throughout a plant. The term was introduced by Carl Nägeli in 1858. Structure The most distinctive xylem cells are the long tracheary elements that transport water. Tracheids and vessel elements are distinguished by their shape; vessel elements are shorter, and are connected together into long tubes that are called ''vessels''. Xylem also contains two other type of cells: parenchyma and fibers. Xylem can be found: * in vascular bundles, present in non-woody plants and non-woody parts of woody plants * in secondary xylem, laid down by a meristem called the vascular cambium in woody plants * as part of a stelar arrangement ...
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Mycelial Cords
Mycelial cords are linear aggregations of parallel-oriented hyphae. The mature cords are composed of wide, empty vessel hyphae surrounded by narrower sheathing hyphae. Cords may look similar to plant roots, and also frequently have similar functions; hence they are also called rhizomorphs (literally, "root-forms"). As well as growing underground or on the surface of trees and other plants, some fungi make mycelial cords which hang in the air from vegetation. Mycelial cords are capable of conducting nutrients over long distances. For instance, they can transfer nutrients to a developing fruiting body, or enable wood-rotting fungi to grow through soil from an established food base in search of new food sources. For parasitic fungi, they can help spread infection by growing from established clusters to uninfected parts. The cords of some wood-rotting fungi (like ''Serpula lacrymans'') may be capable of penetrating masonry. The mechanism of the cord formation is not yet precisely u ...
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Surface Area
The surface area of a solid object is a measure of the total area that the surface of the object occupies. The mathematical definition of surface area in the presence of curved surfaces is considerably more involved than the definition of arc length of one-dimensional curves, or of the surface area for polyhedra (i.e., objects with flat polygonal faces), for which the surface area is the sum of the areas of its faces. Smooth surfaces, such as a sphere, are assigned surface area using their representation as parametric surfaces. This definition of surface area is based on methods of infinitesimal calculus and involves partial derivatives and double integration. A general definition of surface area was sought by Henri Lebesgue and Hermann Minkowski at the turn of the twentieth century. Their work led to the development of geometric measure theory, which studies various notions of surface area for irregular objects of any dimension. An important example is the Minkows ...
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Niche Separation
In ecology, niche differentiation (also known as niche segregation, niche separation and niche partitioning) refers to the process by which competing species use the environment differently in a way that helps them to coexist. The competitive exclusion principle states that if two species with identical niches (ecological roles) compete, then one will inevitably drive the other to extinction. This rule also states that two species cannot occupy the same exact niche in a habitat and coexist together, at least in a stable manner. When two species differentiate their niches, they tend to compete less strongly, and are thus more likely to coexist. Species can differentiate their niches in many ways, such as by consuming different foods, or using different areas of the environment. As an example of niche partitioning, several anole lizards in the Caribbean islands share common diets—mainly insects. They avoid competition by occupying different physical locations. Although these li ...
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Leaf Litter
Plant litter (also leaf litter, tree litter, soil litter, litterfall or duff) is dead plant material (such as leaves, bark, needles, twigs, and cladodes) that have fallen to the ground. This detritus or dead organic material and its constituent nutrients are added to the top layer of soil, commonly known as the litter layer or O horizon ("O" for "organic"). Litter is an important factor in ecosystem dynamics, as it is indicative of ecological productivity and may be useful in predicting regional nutrient cycling and soil fertility. Characteristics and variability Litterfall is characterized as fresh, undecomposed, and easily recognizable (by species and type) plant debris. This can be anything from leaves, cones, needles, twigs, bark, seeds/nuts, logs, or reproductive organs (e.g. the stamen of flowering plants). Items larger than 2 cm diameter are referred to as coarse litter, while anything smaller is referred to as fine litter or litter. The type of litterfall is mos ...
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Organic Material
Organic matter, organic material, or natural organic matter refers to the large source of carbon-based compounds found within natural and engineered, terrestrial, and aquatic environments. It is matter composed of organic compounds that have come from the feces and remains of organisms such as plants and animals. Organic molecules can also be made by chemical reactions that do not involve life. Basic structures are created from cellulose, tannin, cutin, and lignin, along with other various proteins, lipids, and carbohydrates. Organic matter is very important in the movement of nutrients in the environment and plays a role in water retention on the surface of the planet. Formation Living organisms are composed of organic compounds. In life, they secrete or excrete organic material into their environment, shed body parts such as leaves and roots and after organisms die, their bodies are broken down by bacterial and fungal action. Larger molecules of organic matter can be formed ...
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Mineral Soil
Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former term specifically to displaced soil. Soil consists of a solid phase of minerals and organic matter (the soil matrix), as well as a porous phase that holds gases (the soil atmosphere) and water (the soil solution). Accordingly, soil is a three-state system of solids, liquids, and gases. Soil is a product of several factors: the influence of climate, relief (elevation, orientation, and slope of terrain), organisms, and the soil's parent materials (original minerals) interacting over time. It continually undergoes development by way of numerous physical, chemical and biological processes, which include weathering with associated erosion. Given its complexity and strong internal connectedness, soil ecologists regard soil as an ecosystem. Most ...
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Soil Horizons
A soil horizon is a layer parallel to the soil surface whose physical, chemical and biological characteristics differ from the layers above and beneath. Horizons are defined in many cases by obvious physical features, mainly colour and texture. These may be described both in absolute terms (particle size distribution for texture, for instance) and in terms relative to the surrounding material, i.e. 'coarser' or 'sandier' than the horizons above and below. The identified horizons are indicated with symbols, which are mostly used in a hierarchical way. Master horizons (main horizons) are indicated by capital letters. Suffixes, in form of lowercase letters and figures, further differentiate the master horizons. There are many different systems of horizon symbols in the world. No one system is more correct—as artificial constructs, their utility lies in their ability to accurately describe local conditions in a consistent manner. Due to the different definitions of the horizon symb ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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