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Tunicates
Tunicates are marine invertebrates belonging to the subphylum Tunicata ( ). This grouping is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates). The subphylum was at one time called Urochordata, and the term urochordates is still sometimes used for these animals. Despite their simple appearance and very different adult form, their close relationship to the vertebrates is certain. Both groups are chordates, as evidenced by the fact that during their mobile larval stage, tunicates possess a notochord, a hollow dorsal nerve cord, pharyngeal slits, post-anal tail, and an endostyle. They resemble a tadpole. Tunicates are the only chordates that have lost their myomeric segmentation, with the possible exception of the seriation of the gill slits. However, doliolids still display segmentation of the muscle bands. Some tunicates live as solitary individuals, but others replicate by budding and become colonies, e ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Chordata
A chordate ( ) is a bilaterian animal belonging to the phylum Chordata ( ). All chordates possess, at some point during their larval or adult stages, five distinctive physical characteristics (Apomorphy and synapomorphy, synapomorphies) that distinguish them from other Taxon, taxa. These five synapomorphies are a notochord, a neural tube, hollow dorsal nerve cord, an endostyle or thyroid, pharyngeal slits, and a post-anus, anal tail. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and inner mitochondrial membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cephalochordates. These CSIs provide molecular means to reliably distinguish chordates from all other animals. Chordates are divided into three phylum, subphyla: Vertebrata (fish, amphibians, reptiles, birds and mammals), whose notochor ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Thaliacea
Thaliacea is a class of marine chordates within the subphylum Tunicate, Tunicata, comprising the salps, pyrosomes and doliolids. Unlike their benthic relatives the Ascidiacea, ascidians, from which they are believed to have emerged, thaliaceans are free-floating (pelagic) for their entire lifespan. The group includes species with complex life cycles, with both solitary and Colony (biology), colonial forms. Anatomy The three orders of thaliaceans are filter feeders. Pyrosoma, Pyrosomes are colonial animals, with multiple tiny ascidian-like zooids arranged in a cylinder closed at one end. All of the atrial siphons point inwards, emptying into a single, common cloaca in the centre of the cylinder. As the water exhaled by the zooids exits through a common opening, the water movement slowly propels the pyrosome through the sea. Salps and doliolids have a transparent barrel-shaped body through which they pump water, propelling them through the sea, and from which they extract food. The ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Filter Feeder
Filter feeders are aquatic animals that acquire nutrients by feeding on organic matters, food particles or smaller organisms (bacteria, microalgae and zooplanktons) suspended in water, typically by having the water pass over or through a specialized filtering organ that sieves out and/or traps solids. Filter feeders can play an important role in condensing biomass and removing excess nutrients (such as nitrogen and phosphate) from the local waterbody, and are therefore considered water-cleaning ecosystem engineers. They are also important in bioaccumulation and, as a result, as indicator organisms. Filter feeders can be sessile, planktonic, nektonic or even neustonic (in the case of the buoy barnacle) depending on the species and the niches they have evolved to occupy. Extant species that rely on such method of feeding encompass numerous phyla, including poriferans ( sponges), cnidarians (jellyfish, sea pens and corals), arthropods ( krill, mysids and barna ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Vertebrates
Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebrata with some 65,000 species, by far the largest ranked grouping in the phylum Chordata. The vertebrates include mammals, birds, amphibians, and various classes of fish and reptiles. The fish include the jawless Agnatha, and the jawed Gnathostomata. The jawed fish include both the Chondrichthyes, cartilaginous fish and the Osteichthyes, bony fish. Bony fish include the Sarcopterygii, lobe-finned fish, which gave rise to the tetrapods, the animals with four limbs. Despite their success, vertebrates still only make up less than five percent of all described animal species. The first vertebrates appeared in the Cambrian explosion some 518 million years ago. Jawed vertebrates evolved in the Ordovician, followed by bony fishes in the Devonian. T ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Larvacean
Larvaceans, copelates or appendicularians, class Appendicularia, are solitary, free-swimming tunicates found throughout the world's oceans. While larvaceans are filter feeders like most other tunicates, they keep their tadpole-like shape as adults, with the notochord running through the tail. They can be found in the pelagic zone, specifically in the photic zone, or sometimes deeper. They are transparent planktonic animals, usually ranging from to in body length including the tail, although giant larvaceans can reach up to in length. Larvaceans are known for the large houses they build around their bodies to assist in filter-feeding. Secreted from mucus and cellulose, these structures often comprise several layers of filters and can reach up to ten times their body length. In some genera like '' Oikopleura'', houses are built and discarded every few hours, with sinking houses playing a key role in the oceanic carbon cycle. History The study of larvaceans began with the d ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Marine Invertebrates
Marine invertebrates are invertebrate animals that live in marine habitats, and make up most of the macroscopic life in the oceans. It is a polyphyletic blanket term that contains all marine animals except the marine vertebrates, including the non-vertebrate members of the phylum Chordata such as lancelets, sea squirts and salps. As the name suggests, marine invertebrates lack any mineralized tissue, mineralized axial skeleton, axial endoskeleton, i.e. the vertebral column, and some have evolved a rigid seashell, shell, test (biology), test or exoskeleton for protection and/or aquatic locomotion, locomotion, while others rely on hydroskeleton, internal fluid pressure to support their bodies. Marine invertebrates have a large variety of body plans, and have been categorized into over 30 phyla. Evolution The earliest animals were marine invertebrates, that is, vertebrates came later. Animals are multicellular eukaryotes, and are distinguished from plants, algae, and fungi by l ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Polycarpa Aurata
''Polycarpa aurata'', also known as the ox heart ascidian, the gold-mouth sea squirt or the ink-spot sea squirt, is a species of tunicate in the family Styelidae. Description ''Polycarpa aurata'' grows to a height of . It has an urn-shaped, hollow body with two siphons, one at the top and the other on the side. The body colour of this tunicate is white with purple and orange patches and purple lines. The inside is yellow or orange and this is visible around the rim of the siphons. Distribution This species is found in the tropical eastern Indian Ocean and the western Pacific Ocean. Its range includes the Philippines, Indonesia and northern Australia. Its depth range is . Biology Tunicates feed by drawing water in through the branchial siphon at the top, filtering out phytoplankton, bacteria and other food particles, before expelling the water through the atrial siphon at the side. Hydroids and algae may grow on the outside of the tunicate and nudibranch Nudibranchs () ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Shankouclava
''Shankouclava'' is an extinct genus of tunicates. It is one of two candidates for the oldest member of this group, dating to . It has been found in the Lower Cambrian Maotianshan Shale at Shankou village, Anning, near Kunming (South China). Each of the eight specimens found and used for description were isolated, suggesting that the genus was solitary and not colonial. Morphology ''Shankouclava'' had a soft, sac-like body that was elongated and pointed proximally. The body lengths of individuals vary from 2 cm (0.8 in) to 4 cm (1.6 in). Etymology The generic name is composed of the fossil locality, Shankou, and the Latin word ''clava'' (club-shaped). Classification A stem group tunicate affinity for ''Shankouclava'' has been broadly accepted, though some authors have expressed reservations that the evidence in favour of this affinity is not conclusive. Doubts regarding ''Shankouclavas tunicate affinities arose in the context of a long-running debate tha ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Pharyngeal Slit
Pharyngeal slits are Filter feeder, filter-feeding organs found among deuterostomes. Pharyngeal slits are repeated openings that appear along the pharynx caudal to the mouth. With this position, they allow for the movement of water in the mouth and out the pharyngeal slits. It is postulated that this is how pharyngeal slits first assisted in filter-feeding, and later, with the addition of gills along their walls, aided in respiration of aquatic chordates. These repeated segments are controlled by similar developmental mechanisms. Some hemichordate species can have as many as 200 gill slits. Pharynx, Pharyngeal clefts resembling gill slits are transiently present during the embryonic stages of tetrapod development. The presence of pharyngeal arches and clefts in the neck of the developing human embryo famously led Ernst Haeckel to postulate that "ontogeny recapitulates phylogeny"; this hypothesis, while false, contains elements of truth, as explored by Stephen Jay Gould in ''Onto ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Cambrian Stage 3
Cambrian Stage 3 is the still unnamed third stage of the Cambrian. It succeeds Cambrian Stage 2 and precedes Cambrian Stage 4, although neither its base nor top have been formally defined. The plan is for its lower boundary to correspond approximately to the first appearance of trilobites, about million years ago, though the globally asynchronous appearance of trilobites warrants the use of a separate, globally synchronous marker to define the base. The upper boundary and beginning of Cambrian Stage 4 is informally defined as the first appearance of the trilobite genera ''Olenellus'' or ''Redlichia'' around million years ago. Naming The International Commission on Stratigraphy has not officially named the 3rd stage of the Cambrian. The stage approximately corresponds to the "Atdabanian", which is used by geologists working in Siberia. Biostratigraphy The oldest trilobite known is ''Lemdadella'' which appears at the beginning of the ''Fallotaspis'' zone. The Cambrian radiation of ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Endostyle
The endostyle is an organ found in invertebrate chordate species of tunicates, lancelets, and in the larval stage of vertebrate lampreys. It assists in filter-feeding. It has evolved into the thyroid in vertebrate chordates. Since the endostyle is found in all three chordate lineages, it is presumed to have arisen in the common ancestor of these taxa, along with a shift to internal feeding for extracting suspended food particles from the water. When feeding, food particles suspended in the water adhere to the mucus the endostyle produces. The filtered water is then expelled through the gill slits, while the food and mucus are swept into the esophagus by movements of the cilia that coat the endostyle. The endostyle of larval lampreys ( ammocoetes) metamorphoses into the thyroid gland in adults, and is regarded as being homologous to the thyroid in other vertebrates due to its iodine-concentrating activity. One early hypothesis for the function of the endostyle, developed ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Notochord
The notochord is an elastic, rod-like structure found in chordates. In vertebrates the notochord is an embryonic structure that disintegrates, as the vertebrae develop, to become the nucleus pulposus in the intervertebral discs of the vertebral column. In non-vertebrate chordates a notochord persists. The notochord is derived from the embryonic mesoderm and consists of an inner core of vacuolated cells filled with glycoproteins, covered by two helical collagen-elastin sheaths. It lies longitudinally along the rostral-caudal (head to tail) axis of the body, dorsal to the gut tube, and ventral to the dorsal nerve cord. Some chordate invertebrates, such as tunicates, develop a notochord during the larval stage but lose it through subsequent stages into adulthood. The notochord is important for signaling the dorso-ventral patterning of cells coming from the mesodermal progenitors. This helps form the precursors needed for certain organs and the embryo to develop ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
