Myriapod Anatomy
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Myriapod Anatomy
Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial animal, terrestrial. Although molecular evidence and similar fossils suggests a diversification in the Cambrian Period, the oldest known fossil record of myriapods dates between the Wenlock Epoch, Late Silurian and Early Devonian, with ''Pneumodesmus'' preserving the earliest known evidence of air-breathing on land. Other early myriapod fossil species around the similar time period include ''Kampecaris, Kampecaris obanensis'' and ''Archidesmus'' sp. The phylogenetic classification of myriapods is still debated. The scientific study of myriapods is myriapodology, and those who study myriapods are myriapodologists. Anatomy Myriapods have a single pair of antenna (biology), antennae and, in most cases, simple eyes. Exceptions are the two classes of symphylans and pauropods, the millipede order Polydesmida and ...
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Chilopoda
Centipedes (from Neo-Latin , "hundred", and Latin language, Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', "lip", and Neo-Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (Metamerism (biology), metameric) animals with one pair of legs per body segment. All centipedes are venomous and can inflict painful centipede bite, stings, injecting their venom through Pincer (biology), pincer-like appendages known as forcipules or toxicognaths, which are actually modified legs instead of fangs. Despite the name, no species of centipede has exactly 100 legs; the number of pairs of legs is an odd number that ranges from 15 pairs to 191 pairs. Centipedes are predominantly generalist (biology), generalist carnivore, carnivorous, hunting for a variety of prey items that can be overpowered. They have ...
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Early Devonian
The Early Devonian is the first of three Epoch (geology), epochs comprising the Devonian period, corresponding to the Lower Devonian Series (stratigraphy), series. It lasted from and began with the Lochkovian Stage , which was followed by the Pragian from and then by the Emsian, which lasted until the Middle Devonian began, . During this time, the first Ammonoidea, ammonoids appeared, descending from Bactritida, bactritoid Nautiloidea, nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatite, Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic Era. References

Early Devonian, Geological epochs Devonian geochronology, *01 {{Geochronology-stub ...
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Geophilomorpha
Geophilomorpha is an order of centipedes commonly known as soil centipedes. The name "Geophilomorpha" is from Ancient Greek roots meaning "formed to love the earth." This group is the most diverse centipede order, with 230 genera. These centipedes are found nearly worldwide but are absent in Antarctica and most Arctic regions. Description Centipedes in this order are epimorphic, hatching with a full complement of segments. These centipedes each have an odd number of leg-bearing segments ranging from 27 (in the genus '' Schendylops'') to 191 (in the species '' Gonibregmatus plurimipes''). They are eyeless and blind, with long and narrow bodies, ranging from yellow to brown in color and from about 1 cm to 22 cm in length. They bear spiracles on all leg-bearing segments except the first and the last. The antennae have 14 segments and are usually slightly attenuated. Suborders and families This order is a monophyletic group including two suborders: the monophyletic Placodesmata, ...
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Polydesmida
Polydesmida (from the Greek ''poly'' "many" and ''desmos'' "bond") is the largest order of millipedes, with more than 5,000 species, including all the millipedes reported to produce hydrogen cyanide (HCN). This order is also the most diverse of the millipede orders in terms of morphology. Millipedes in this order are found in all regions of the world other than Antarctica. Description Members of the order Polydesmida are also known as flat-backed millipedes, because on most species, each body segment has wide lateral keels known as paranota. These keels are produced by the posterior half (metazonite) of each body ring behind the collum. Polydesmids have no eyes, and vary in length from 1.4 to 134 mm. Many of the larger species show bright coloration patterns which warn predators of their toxic secretions. Adults usually have 20 segments, counting the collum as the first ring and the telson as the last ring. Juveniles have from 7 to 19 rings. In species with the usual 20 seg ...
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Pauropod
Pauropoda is a class of small, pale, millipede-like arthropods in the subphylum Myriapoda. More than 900 species in twelve families are found worldwide, living in soil and leaf mold. Pauropods look like centipedes or millipedes and may be a sister group of the latter, but a close relationship with Symphyla has also been posited. The name Pauropoda derives from the Greek ''pauros'' (meaning "small" or "few") and ''pous'', genitive ''podos'' (meaning "foot"), because most species in this class have only nine pairs of legs as adults, a smaller number than those found among adults in any other class of myriapods. Anatomy Pauropods are soft, cylindrical animals with bodies measuring only 0.3 to 2 mm in length. They have neither eyes nor hearts, although they do have sensory organs which can detect light. The body segments have ventral tracheal/spiracular pouches forming apodemes similar to those in millipedes and Symphyla, although the trachea usually connected to these structur ...
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Symphylan
Symphylans, also known as garden centipedes or pseudocentipedes, are soil-dwelling arthropods of the class (biology), class Symphyla in the subphylum Myriapoda. Symphylans resemble centipedes, but are very small, non-venomous, and Myriapoda#Myriapod relationships, may or may not form a clade with centipedes. More than 200 species are known worldwide. Symphyla are primarily herbivores and detritus feeders living deep in the soil, under stones, in decaying wood, and in other moist places. They are rapid runners, can move quickly through the pores between soil particles, and are typically found from the surface down to a depth of about . They detritivore, consume decaying vegetation, but can do considerable harm in an Agriculture, agricultural setting by consuming seeds, roots, and root hairs in cultivated soil. For example, the garden symphylan, ''Scutigerella immaculata'' can be a pest of crops. A species of ''Hanseniella (myriapod), Hanseniella'' has been recorded as a pest of su ...
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Antenna (biology)
An antenna (plural: antennae) is one of a pair of appendages used for Sensory system, sensing in arthropods. Antennae are sometimes referred to as ''feelers''. Antennae are connected to the first one or two Segmentation (biology), segments of the arthropod head. They vary widely in form but are always made of one or more jointed segments. While they are typically sensory organs, the exact nature of what they sense and how they sense it is not the same in all groups. Functions may variously include sensing tactition, touch, air motion, heat, vibration (sound), and especially insect olfaction, smell or gustation, taste. Antennae are sometimes modified for other purposes, such as mating, brooding, swimming, and even anchoring the arthropod to a substrate (biology), substrate. Larval arthropods have antennae that differ from those of the adult. Many crustaceans, for example, have free-swimming larvae that use their antennae for swimming. Antennae can also locate other group members i ...
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Houghton Mifflin Harcourt
Houghton Mifflin Harcourt Company ( ; HMH) is an American publisher of textbooks, instructional technology materials, assessments, and reference works. The company is based in the Financial District, Boston, Boston Financial District. It was formerly known as the Houghton Mifflin Company, but it changed its name following the 2007 acquisition of Harcourt (publisher), Harcourt Publishing. Prior to March 2010, it was a subsidiary of EMPG, Education Media and Publishing Group Limited, an Irish-owned holding company registered in the Cayman Islands and formerly known as Riverdeep. In 2022, it was acquired by Veritas Capital, a New York-based private-equity firm. Company history In 1832, William Ticknor and John Allen purchased a bookselling business in Boston and began to involve themselves in publishing; James T. Fields joined as a partner in 1843. Fields and Ticknor gradually gathered an impressive list of writers, including Ralph Waldo Emerson, Nathaniel Hawthorne, and Henry Dav ...
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Myriapodology
Myriapodology is the scientific study of myriapods which includes centipedes and millipedes. The field of myriapodology can also cover other myriapods such as pauropods and symphylans. Those who study myriapods are myriapodologists. Societies * International Society of Myriapodology Journals * ''International Journal of Myriapodology'' * ''Myriapodologica'' * ''Myriapod Memoranda'' Notable myriapodologists * Carl Attems (1868–1952), Austrian zoologist, described over 1,000 species * Stanley Graham Brade-Birks (1887-1982), English myriapodologist who with Hilda K Brade-Birks authored ''Notes on Myriapoda'': 23 papers jointly from 1916 to the 1920s; then twelve more solo until 1939 * Henry W. Brolemann (1860–1933), French myriapodologist, described around 500 species * Ralph Vary Chamberlin (1879–1967), American arachnologist and myriapodologist, described over 1,000 species * Orator F. Cook (1867–1949), American botanist and myriapodologist, co-described world's leggie ...
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Phylogenetic Classification
Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional method, by which taxon names are defined by a ''type'', which can be a specimen or a taxon of lower rank, and a description in words. Phylogenetic nomenclature is regulated currently by the '' International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature associates names with clades, groups consisting of an ancestor and all its descendants. Such groups are said to be monophyletic. There are slightly different methods of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the taxon named. Listing all these organisms (i.e. providing a full circumscription) requires the complete phylogenetic tree to be known. In practice, there ...
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Archidesmus
''Archidesmus'' is an extinct millipede genus from the Lower Devonian Old Red Sandstone Old Red Sandstone, abbreviated ORS, is an assemblage of rocks in the North Atlantic region largely of Devonian age. It extends in the east across Great Britain, Ireland and Norway, and in the west along the eastern seaboard of North America. It ... of the United Kingdom. It is the only member of the taxonomic family (biology), family Archidesmidae. Individuals were up to long, and had 60 to 80 body segments decorated with tubercles (bumps) on the upper surface, and most segments possessed wing-like keels (paranota) extending to the side. The type species ''Archidesmus macnicoli'' was described by British paleontologist Benjamin Peach in 1882. A second species, ''A. loganensis'' was also described by Peach but its status as a valid species- or even a myriapod- is debated. References

Millipede genera, †Archidesmus Devonian myriapods Fossils of Great Britain Fossil taxa describ ...
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