Marrellomorpha
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Marrellomorpha
Marrellomorpha are an extinct group of arthropods known from the Cambrian to the Early Devonian. They lacked mineralised hard parts, so are only known from areas of exceptional preservation, limiting their fossil distribution. The best known member is '' Marrella'', with thousands of specimens found in the Cambrian aged Burgess Shale of Canada. Taxonomy The group is divided up into two major orders, Marrellida and Acercostraca. Marrellida is recognised by the possession of head shields with two or three pairs of elongate spine-like projections, and three attached pairs of uniramous (single-branched) appendages, while Acercostraca generally have large ovoid carapaces that cover the entire upper half of the body, and five pairs of uniramous appendages attached to the head section. Both groups have unbranched antennae and a segmented trunk with biramous (divided into two branches) appendages. Recent research has suggested the previously enigmatic Cambrian trilobite-like arthrop ...
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Marrella
''Marrella'' is an extinct genus of marrellomorph arthropod known from the Middle Cambrian of North America and Asia. It is the most common animal represented in the Burgess Shale of British Columbia, Canada, with tens of thousands of specimens collected. Much rarer remains are also known from deposits in China. History ''Marrella'' was the first fossil collected by Charles Doolittle Walcott from the Burgess Shale, in 1909. Walcott described ''Marrella'' informally as a "lace crab" and described it more formally as an odd trilobite. It was later reassigned to the now defunct class Trilobitoidea in the ''Treatise on Invertebrate Paleontology''. In 1971, Whittington undertook a thorough redescription of the animal and, on the basis of its legs, gills and head appendages, concluded that it was neither a trilobite, nor a chelicerate, nor a crustacean. ''Marrella'' is one of several unique arthropod-like organisms found in the Burgess Shale. Other examples are '' Opabinia'' and ' ...
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Artiopoda
Artiopoda is a clade of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomorpha. Trilobites, in part due to abundance of findings owing to their mineralized exoskeletons, are by far the best recorded, diverse, and long lived members of the clade. Other members, which lack mineralised exoskeletons, are known mostly from Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordov ... deposits. Description According to Stein and Selden (2012) artiopods are recognised by the possession of filiform antennulae, limbs with bilobate exopods (upper branches), with the proximal (closest to base of the limb) lobe being elong ...
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Arthropod
Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metamerism (biology), metameric) Segmentation (biology), segments, and paired jointed appendages. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. They form an extremely diverse group of up to ten million species. Haemolymph is the analogue of blood for most arthropods. An arthropod has an open circulatory system, with a body cavity called a haemocoel through which haemolymph circulates to the interior Organ (anatomy), organs. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. They have ladder-like nervous systems, with paired Anatomical terms of location#Dorsal and ventral, ventral Ventral nerve cord, nerve cord ...
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Arachnomorpha
Arachnomorpha is a proposed subdivision or clade of Arthropoda, comprising the group formed by the trilobites and their close relatives (Artiopoda), Megacheira (which may be paraphyletic) and chelicerates. Under this proposed classification scheme, Arachnomorpha is considered the sister group to Mandibulata (including insects, crustaceans and myriapods). The arachnomorph concept has been challenged by suggestions that the artiopods are more closely related to mandibulates, forming the clade Antennulata instead. There is no consensus as to assigning Arachnomorpha a formal Linnean rank. Classification Arachnomorpha Lameere, 1890 Arachnata Paulus, 1979, = Palaeopoda Packard, 1903*†'' Nettapezoura'' *†'' Dicranocaris'' *† Trilobita Walch, 1771 *† Megacheira Hou & Bergstrom, 1997 *† Strabopida Hou & Bergström, 1997 *† Aglaspida Walcott, 1911 *† Cheloniellida Broili, 1932 *Chelicerata Heymons 1901 **Family † Sanctacarididae Legg & Pates, 2017 **Class Pycnogoni ...
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Skania
''Skania'' is a Cambrian fossil arthropod. The type species, ''S. fragillis'', is known from the Burgess Shale of British Columbia, Canada. A second possible species ''"S." sundbergi'' is known from the Kaili Formation of China, but its placement within the genus has been questioned. Description Specimens of ''S. fragillis'' range in length from The cordiform dorsal shield/carapace covers the entire body, with its midline having a raised keel. The shield curls around the front to form a doublure, with the posterior edge of the doublure having a pair of lateral spines. A pair of spines is also present on the posterior of the dorsal shield. A hypostome was present on the underside of the head. There are two sets of appendages, the first 5 pairs of appendages are elongate, the first one of which is described as an antenna and has small subquadrate segments/podomeres, while the others have elongate subrectangular segments. These appendages are somewhat curved and extend beyo ...
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Burgessia
''Burgessia'' is a genus of arthropod known from the mid-Cambrian aged Burgess Shale of British Columbia, Canada. It is relatively abundant, with over 1,700 specimens having been collected. Description The body had an unsegmented approximately circular carapace that was somewhat convex. It was likely thin and had only weak sclerotization. A pair of tapering flexible segmented antennae projected forwards from the head, which were about equal in length to the carapace, these were likely tactile in function. There are three pairs of cephalic appendages excluding the antennae that functioned as walking limbs, as well as seven pairs of biramous (two-branched) walking limbs with gills, otherwise similar to the cephalic limbs running along the trunk, which decreased in size posteriorly. Although not visible on any specimens, the mouth was almost certainly located on the underside of the body. The circular carapace was largely occupied by the guts, which were divided into two sect ...
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Mandibulata
The clade Mandibulata constitutes one of the major subdivisions of the phylum Arthropoda, alongside Chelicerata. Mandibulates include the crustaceans, myriapods (centipedes and millipedes, among others), and all true insects. The name "Mandibulata" refers to the mandibles, a modified pair of limbs used in food processing, the presence of which are characteristic of most members of the group. The mandibulates are divided between the extant groups Myriapoda (millipedes and centipedes, among others) and Pancrustacea (including crustaceans and hexapods, the latter group containing insects). Molecular phylogenetic studies suggest that the living arthropods are related as shown in the cladogram below. Crustaceans do not form a monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent ...
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Xylokorys
''Xylokorys'' is a genus of marrellomorph known from two specimens from the Silurian The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of t ... Herefordshire lagerstatte; it filter-fed on mud particles on the sea floor. It is the only marrellomorph known from the Silurian period. References Silurian arthropods of Europe Marrellomorpha {{paleo-arthropod-stub ...
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Uniramous
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, : ''coxae''), ''trochanter'', ''femur'' (: ''femora''), ''tibia'' (: ''tibiae''), ''tarsus'' (: ''tarsi''), ''ischium'' (: ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (: ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments articulates with the next segmen ...
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Karl Beurlen
Karl Beurlen (17 April 1901 – 27 December 1985) was a German paleontologist. Beurlen was born in Aalen. He attended University of Tübingen. He completed a PhD in 1923. p. 115-119/ref> Beurlen was a proponent of orthogenesis and saltational evolution. He used the term ''metakinesis'' (coined by Otto Jaekel) to describe sudden changes of development in organisms. He also invented the term ''palingenesis'' as a mechanism for his orthogenetic theory of evolution. He was an assistant of Edwin Hennig. He was a proponent of National Socialist ideology and wrote about the Aryan race The Aryan race is a pseudoscientific historical race concepts, historical race concept that emerged in the late-19th century to describe people who descend from the Proto-Indo-Europeans as a Race (human categorization), racial grouping. The ter .... He was director of the Zoologische Staatssammlung München. See also * Otto Schindewolf References 1901 births 1985 deaths German paleonto ...
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Cambrian Stage 3
Cambrian Stage 3 is the still unnamed third stage of the Cambrian. It succeeds Cambrian Stage 2 and precedes Cambrian Stage 4, although neither its base nor top have been formally defined. The plan is for its lower boundary to correspond approximately to the first appearance of trilobites, about million years ago, though the globally asynchronous appearance of trilobites warrants the use of a separate, globally synchronous marker to define the base. The upper boundary and beginning of Cambrian Stage 4 is informally defined as the first appearance of the trilobite genera ''Olenellus'' or ''Redlichia'' around million years ago. Naming The International Commission on Stratigraphy has not officially named the 3rd stage of the Cambrian. The stage approximately corresponds to the "Atdabanian", which is used by geologists working in Siberia. Biostratigraphy The oldest trilobite known is ''Lemdadella'' which appears at the beginning of the ''Fallotaspis'' zone. The Cambrian radiation of ...
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Maotianshan Shales
The Maotianshan Shales () are a series of Early Cambrian sedimentary deposits in the Chiungchussu Formation or Heilinpu Formation, famous for their '' Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale of British Columbia, Canada. They take their name from Maotianshan Hill () in Chengjiang County, Yunnan Province, China. The most famous assemblage of organisms are referred to as the Chengjiang biota for the multiple scattered fossil sites in Chengjiang. The age of the Chengjiang Lagerstätte is locally termed Qiongzhusian, a stage correlated to the late Atdabanian Stage in Siberian sequences of the middle of the Early Cambrian. The shales date to ≤. Along with the Burgess Shale, the Maotianshan Shales ...
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