Lithostrotians
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Lithostrotians
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node-based ...
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Titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as '' Patagotitan'', estimated at long with a mass of , and the comparably-sized '' Argentinosaurus'' and '' Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a ''nomen dubium)'' '' Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Fossil record Due to the near-global distr ...
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Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 Megaannum#SI prefix multipliers, Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian–Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining Anoxic event#Cretaceous, early Aptian Oceanic Anoxic Event 1a (OAE1a) was dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java Plateau, Ontong Java-Manihiki Plateau, Manihiki-Hikurangi Plateau, Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Onto ...
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Uriash
''Uriash'' is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous (Maastrichtian) of Romania. The holotype of this genus was originally referred to ''Magyarosaurus hungaricus'', which is now classified in a separate genus, '' Petrustitan''. Discovery and naming In 1914, the Hungarian geologist Ottokár Kadić in the Pârâul Budurone ravine near Vălioara, discovered a skeleton of a sauropod which was markedly larger than previously found sauropod remains in the area. In 1916, the discovery, consisting of limb bones and eight vertebrae, was reported in the scientific literature. By 12 January 1927 two vertebrae had been sent to Friedrich von Huene at the University of Tübingen. Von Huene described these in 1932, preliminary referring them to a ?''Magyarosaurus hungaricus'', presently ''Petrustitan''. In 2021 the site was rediscoveredBotfalvai, G., Csiki-Sava, Z., Kocsis, L., Albert, G., Magyar, J., Bodor, E.R., Ţabara, D., Ulyanov, A., & Makadi, L. 2021. "‘X ...
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Sauropod Caudal Pneumaticity
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their body), and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include ''Apatosaurus'', ''Argentinosaurus'', ''Alamosaurus'', ''Brachiosaurus'', ''Camarasaurus'', ''Diplodocus,'' and '' Mamenchisaurus''. The oldest known unequivocal sauropod dinosaurs are known from the Early Jurassic. ''Isanosaurus'' and '' Antetonitrus'' were originally described as Triassic sauropods, but their age, and in the case of ''Antetonitrus'' also its sauropod status, were subsequently questioned. Sauropod-like sauropodomorph tracks from the Fleming Fjord Formation (Greenland) might, however, indicate the occurrence of the group in ...
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Titanosaurus
''Titanosaurus'' (; ) is a dubious genus of sauropod dinosaurs, first described by Richard Lydekker in 1877.R. Lydekker. (1877). Notices of new and other Vertebrata from Indian Tertiary and Secondary rocks. ''Records of the Geological Survey of India'' 10(1):30-43 It is known from the Maastrichtian (Upper Cretaceous) Lameta and Kallakurichi Formations of India. Discovery and naming ''Titanosaurus'', literally meaning 'titanic lizard', was named after the mythological Titans. ''Titanosaurus'' was the first Indian dinosaur to be named and properly described, having been recorded for the first time in 1877. The type species, ''T. indicus'', was named in 1877, and the second species, ''T. blanfordi'', was named in 1879.R. Lydekker. (1879). Fossil Reptilia and Batrachia. ''Memoirs of the Geological Survey of India. Palaeontologia Indica, Series IV. Indian Pretertiary Vertebrata'' 1(3):1-36 Both species were named by Richard Lydekker. ''T. indicus'' and ''T. blanfordi'' are 70 mil ...
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Node-based Taxon
Phylogenetic nomenclature is a method of nomenclature for taxon, taxa in biology that uses phylogenetics, phylogenetic definitions for taxon names as explained below. This contrasts with Biological classification, the traditional method, by which taxon names are defined by a ''Type (biology), type'', which can be a specimen or a taxon of lower Taxonomic rank, rank, and a description in words. Phylogenetic nomenclature is regulated currently by the ''PhyloCode, International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature associates names with clades, groups consisting of an ancestor and all its descendants. Such groups are said to be Monophyly, monophyletic. There are slightly different methods of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the taxon named. Listing all these organisms (i.e. prov ...
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Caudal Vertebrae
Caudal vertebrae are the vertebrae of the tail in many vertebrates. In birds, the last few caudal vertebrae fuse into the pygostyle, and in apes, including humans, the caudal vertebrae are fused into the coccyx. In many reptiles, some of the caudal vertebrae bear ribs, the caudal ribs, though these are often fused with the vertebrae. The caudal vertebrae often articulate with haemal arches ventrally. The number of caudal vertebrae in animals can vary greatly. Anguid lizards have been reported to have as many as 111 caudal vertebrae, whereas as few as seven are present in the tail of the early therapsid ''Tapinocaninus''. In lepidosaurs and captorhinids, the caudal vertebrae possess fracture planes at mid-length that allow caudal autotomy. In frogs, the few caudal vertebrae are fused together to form part of the urostyle. References

Skeletal system Vertebrate anatomy {{vertebrate-anatomy-stub ...
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Osteoderms
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of Extant taxon, extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, Temnospondyli, temnospondyls (extinct amphibians), various groups of dinosaurs (most notably ankylosaurs and stegosaurians), phytosaurs, aetosaurs, placodonts, and hupehsuchians (marine reptiles with possible ichthyosaur affinities). Osteoderms are uncommon in mammals, although they have occurred in many xenarthrans (armadillos and the extinct glyptodonts and Mylodontidae, mylodontid ground sloths). The heavy, bony osteoderms have evolved independently in many different lineages. The armadillo osteoderm is believed to develop in subcutaneous dermal tissues. These varied structures should be thought of as anatomical analogues, not homology (biology), homologues, and do not necessarily indicate monophyly. The structures are however derived from scute, sc ...
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Ancient Greek
Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek Dark Ages, Dark Ages (), the Archaic Greece, Archaic or Homeric Greek, Homeric period (), and the Classical Greece, Classical period (). Ancient Greek was the language of Homer and of fifth-century Athens, fifth-century Athenian historians, playwrights, and Ancient Greek philosophy, philosophers. It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article primarily contains information about the Homeric Greek, Epic and Classical periods of the language, which are the best-attested periods and considered most typical of Ancient Greek. From the Hellenistic period (), Ancient Greek was followed by Koine Greek, which is regar ...
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Saltasaurus
''Saltasaurus'' (which means "lizard from Salta") is a genus of saltasaurid dinosaur of the Late Cretaceous Period (geology), period of Argentina. Small among sauropods, though still heavy by the standards of modern creatures, ''Saltasaurus'' was characterized by a short neck and stubby limbs. It was the first genus of sauropod known to possess Armour (zoology), armour of bony plates embedded in its skin. Such small bony plates, called osteoderms, have since been found on other titanosaurians. Discovery The fossils of ''Saltasaurus'' were excavated by José Bonaparte, Martín Vince and Juan C. Leal between 1975 and 1977 at the Estancia "El Brete". The find was in 1977 reported in the scientific literature. ''Saltasaurus'' was named and described by Bonaparte and Jaime E. Powell in 1980. The type species is ''Saltasaurus loricatus''. Its generic name is derived from Salta Province, the region of north-west Argentina where the first fossils were recovered. The specific name (zoolo ...
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Sauropods
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their body), and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include '' Apatosaurus'', '' Argentinosaurus'', '' Alamosaurus'', ''Brachiosaurus'', '' Camarasaurus'', '' Diplodocus,'' and '' Mamenchisaurus''. The oldest known unequivocal sauropod dinosaurs are known from the Early Jurassic. '' Isanosaurus'' and '' Antetonitrus'' were originally described as Triassic sauropods, but their age, and in the case of ''Antetonitrus'' also its sauropod status, were subsequently questioned. Sauropod-like sauropodomorph tracks from the Fleming Fjord Formation (Greenland) might, however, indicate the occurrence of the g ...
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Clade
In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach to taxonomy adopted by most biological fields. The common ancestor may be an individual, a population, or a species (extinct or Extant taxon, extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed ''monophyletic'' (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming Taxon, taxa that are not clades; that is, taxa that are not Monophyly, monophyletic. Some of the relationships between organisms that the molecul ...
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