Guadalupian First Appearances
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Guadalupian First Appearances
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian is also known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. ...
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International Commission On Stratigraphy
The International Commission on Stratigraphy (ICS), sometimes unofficially referred to as the International Stratigraphic Commission, is a daughter or major subcommittee grade scientific organization that concerns itself with stratigraphy, stratigraphical, geology, geological, and chronology, geochronological matters, worldwide. It is the largest subordinate body of the International Union of Geological Sciences (IUGS). The ICS is essentially a permanent working committee, working subcommittee, which meets far more regularly than the quadrennial meetings scheduled by the IUGS, when it meets as a congress or committee, membership of the whole. Aims One of its main aims, a project begun in 1974, is to establish a multidisciplinary standard and global geologic time scale that will ease paleontology, paleontological and geobiology, geobiological comparisons region to region by benchmarks with stringent and rigorous strata criteria called Global Boundary Stratotype Section and Points ...
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Lopingian
The Lopingian is the uppermost series/last epoch of the Permian. It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic. The Lopingian is often synonymous with the informal terms late Permian or upper Permian. The name was introduced by Amadeus William Grabau in 1931 and derives from Leping, Jiangxi in China. It consists of two stages/ ages. The earlier is the Wuchiapingian and the later is the Changhsingian. The International Chronostratigraphic Chart (v2018/07) provides a numerical age of 259.1 ±0.5 Ma. If a Global Boundary Stratotype Section and Point (GSSP) has been approved, the lower boundary of the earliest stage determines numerical age of an epoch. The GSSP for the Wuchiapingian has a numerical age of 259.8 ± 0.4 Ma. Evidence from Milankovitch cycles suggests that the length of an Earth day during this epoch was approximately 22 hours. Geography During the Lopingian, most of the earth was ...
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Biarmosuchia
Biarmosuchia is an extinct clade of non-mammalian synapsids from the Permian. Biarmosuchians are the most basal group of the therapsids. They were moderately-sized, lightly built carnivores, intermediate in form between basal sphenacodont " pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation. Characteristics The biarmosuchian skull is very similar to the sphenacodontid skull, differing only in the larger temporal fenestra (although these are still small relative to later therapsids), slightly backward-sloping occiput (the reverse of the pelycosaur condition), reduced number of teeth, and single large canine teeth in both upper and lower jaws, and other features (Carroll 1988 pp. 370, Benton 2000 p. 114). In later specialised Biarmosuchia, these resemble the enlarged canines of the Gorgonopsia. T ...
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Anomodont
Anomodontia is an extinct group of non-mammalian therapsids from the Permian and Triassic periods. By far the most speciose group are the dicynodonts, a clade of beaked, tusked herbivores. Anomodonts were very diverse during the Middle Permian, including primitive forms like '' Anomocephalus'' and '' Patranomodon'' and groups like Venyukovioidea and Dromasauria. Dicynodonts became the most successful and abundant of all herbivores in the Late Permian, filling ecological niches ranging from large browsers down to small burrowers. Few dicynodont families survived the Permian–Triassic extinction event, but one lineage ( Kannemeyeriiformes) evolved into large, stocky forms that became dominant terrestrial herbivores right until the Late Triassic, when changing conditions caused them to decline, finally going extinct during the Triassic–Jurassic extinction event. Classification Taxonomy * Order Therapsida * Suborder Anomodontia ** '' Biseridens'' ** '' Patranomodon'' ** ...
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Dinocephalia
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They ...
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Clades
In biology, a clade (), also known as a monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach to taxonomy adopted by most biological fields. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed ''monophyletic'' (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms that the molecular biology arm of cladistics has revealed ...
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Sphenacodontia
Sphenacodontia is a stem-based taxon, stem-based clade of derived synapsids. It was defined by Amson and Laurin (2011) as "the largest clade that includes ''Haptodus baylei'', ''Haptodus garnettensis'' and ''Sphenacodon ferox'', but not ''Edaphosaurus pogonias''". They first appear during the Late Pennsylvanian (Upper Carboniferous) epoch. From the end of the Carboniferous to the end of the Permian, most of them remained large, with only some secondarily becoming small in size. Basal (phylogenetics), Basal Sphenacodontia constitute a Transitional fossils, transitional evolutionary series from early pelycosaurs to ancestral therapsids (which in turn were the ancestors of cynodont, more advanced forms and finally the mammals). One might say that the sphenacodontians are proto-therapsids (even though there is almost a 30-million-years gap between the separation of the ancestors of therapsids from other sphenacodontians and the first appearance of therapsids in the fossil record). Ch ...
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Pelycosaur
Pelycosaur ( ) is an older term for basal or primitive Late Paleozoic synapsids, excluding the therapsids and their descendants. Previously, the term mammal-like reptile was used, and Pelycosauria was considered an order, but this is now thought to be incorrect and outdated. Because it excludes the advanced synapsid group Therapsida, the term is paraphyletic and contrary to modern formal naming practice. Thus the name ''pelycosaurs'', similar to the term ''mammal-like reptiles'', fell out of favor among scientists by the 21st century, and is only used informally, if at all, in the modern scientific literature. The terms stem mammals, protomammals, and basal or primitive synapsids are instead used where needed. Etymology The modern word was created from Greek meaning 'basin' and meaning 'lizard'. The term ''pelycosaur'' has been fairly well abandoned by paleontologists because it no longer matches the features that distinguish a clade. Pelycosauria is a paraphyletic ...
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End-Capitanian Extinction Event
The Capitanian mass extinction event (also known as the end-Guadalupian extinction event, the Guadalupian-Lopingian boundary mass extinction, the pre-Lopingian crisis, or the Middle Permian extinction) was a major mass extinction event that occurred towards the end of the Capitanian age and Guadalupian (Middle Permian) epoch of the Permian period. The mass extinction occurred during a period of decreased species richness and increased extinction rates. It is often called the end-Guadalupian extinction event because of its initial recognition between the Guadalupian and Lopingian series; however, more refined stratigraphic study suggests that extinction peaks in many taxonomic groups occurred within the Guadalupian, in the latter half of the Capitanian age.Bond, D. P. G., Wignall, P. B., Wang, W., Izon, G., Jiang, H. S., Lai, X. L., Sund, Y.-D., Newtona, R.J., Shaoe, L.-Y., Védrinea, S. & Cope, H. (2010). "The mid-Capitanian (Middle Permian) mass extinction and carbon isotope reco ...
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Extinction Event
An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp fall in the diversity and abundance of multicellular organisms. It occurs when the rate of extinction increases with respect to the background extinction rate and the rate of speciation. Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from disagreement as to what constitutes a "major" extinction event, and the data chosen to measure past diversity. The "Big Five" mass extinctions In a landmark paper published in 1982, Jack Sepkoski and David M. Raup identified five particular geological intervals with excessive diversity loss. They were originally identified as outliers on a general trend of decreasing extinction rates during the Phanerozoic, but as more stringent statistical tests have been applied t ...
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Olson's Extinction
Olson's Extinction was a mass extinction that occurred in the late Cisuralian or early Guadalupian epoch of the Permian period, predating the much larger Permian–Triassic extinction event. The event is named after American paleontologist Everett C. Olson, who first identified the gap in fossil record indicating a sudden change between the early Permian and middle/ late Permian faunas. Some authors also place a hiatus in the continental fossil record around that time, but others disagree. This event has been argued by some authors to have affected many taxa, including embryophytes, marine metazoans, and tetrapods. Identification The first evidence of extinction came when Everett C. Olson noted a hiatus between early Permian faunas dominated by pelycosaurs and therapsid-dominated faunas of the middle and late Permian. First considered to be a preservational gap in the fossil record, the event was originally dubbed 'Olson's Gap'. To compound the difficulty in identifyin ...
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