Gleicheniales
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Gleicheniales
Gleicheniales is an order of ferns in the subclass Polypodiidae (the leptosporangiate ferns). The Gleicheniales has spore records potentially as early as the Early Carboniferous, but the oldest unambiguous macrofossil records date to the Early Permian. Description These ferns are characterized by root steles having 3–5 protoxylem poles and antheridia with 6–12 narrow, twisted or curved cells in their walls, Gleichenia-type spore wall ultrastructure. Otherwise, their habitus is highly diverse, including plants with the typical fern fronds, others whose leaves resemble those of palm trees, and yet others again which have undivided leaves. They are tropical ferns, most diverse in Asia and the Pacific region. Classification In the molecular phylogenetic classification of Smith et al. in 2006, the Gleicheniales were placed in class Polypodiopsida (the leptosporangiate ferns). Three families, Dipteridaceae, Gleicheniaceae, and Matoniaceae were recognized. The linear seque ...
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Matoniaceae
Matoniaceae is one of the three families of ferns in the Gleicheniales order of the Polypodiopsida class. Fossil records reveal that Matoniaceae ferns were abundant during the Mesozoic era (about 250-million to 66-million years ago), during which they lived on every continent, including Antarctica, with eight genera and 26 species, with the oldest known specimens being from the Middle Triassic of Antarctica. Today the family is much less abundant, and also less diverse, with only two extant genera and four species, which are limited to portions of southeastern Asia. The following diagram shows a likely phylogenic relationship with the other two families of the Gleicheniales. Extant taxa * genus ** species ** species * genus ** species ** species Extinct genera Some common Mesozoic Matoniaceae genera and a sampling of their species include: * genus ** species ** species * genus ** species ** species ** species ** species ** species * genus * genus
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Gleicheniaceae
The forked ferns are the family Gleicheniaceae, which includes six genera and about 160 known species. The formerly independent families Dicranopteridaceae and Stromatopteridaceae are generally included in the Gleicheniaceae, whereas the Dipteridaceae and Matoniaceae, although closely related, are considered separate families by most authors.Smith ''et al.'' (2006) Description These tropical ferns are the most widespread living lineage of Gleicheniales. Their rhizomes have a "vitalized" protostele or in some taxa a solenostele. The leaves are indeterminate, with pseudodichotomously forked leaves except in '' Stromatopteris'', and free veins. ''Stromatopteris'' also differs from the other species in having an achlorophyllous and subterranean gametophyte. The sori are abaxial but not marginal and carry 5–15 exindusiate round sporangia each. These have a transverse-oblique annulus and contain 128 to 800 bilateral or globose-tetrahedral spores. The sori and sporangia mature ...
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Dipteridaceae
The Dipteridaceae is a family of ferns in the order Gleicheniales of the class Polypodiopsida. They are commonly known as umbrella ferns and contain two genera, '' Cheiropleuria'' and '' Dipteris'', with a total of nine species confined to Asia, New Guinea and northern Australia While currently a small family, they were much more abundant in the Mesozoic era, with the oldest fossils being known from the Middle Triassic of Italy, Australia and Argentina. Seven fossil genera are recognised, including '' Hausmannia'', '' Clathropteris'', '' Dictyophyllum'', '' Thaumatopteris'', '' Camptopteris'', and '' Polyphacelus''. The following diagram shows the placement of Dipteridaceae in a likely phylogenic relationship with the other two families of Gleicheniales Gleicheniales is an order of ferns in the subclass Polypodiidae (the leptosporangiate ferns). The Gleicheniales has spore records potentially as early as the Early Carboniferous, but the oldest unambiguous macrofossil record ...
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Polypodiidae (plant)
The Polypodiidae, commonly called leptosporangiate ferns, formerly Leptosporangiatae, are one of four subclasses of ferns, the largest of these being the largest group of living ferns, including some 11,000 species worldwide. The group has also been treated as the class Pteridopsida or Polypodiopsida, although other classifications assign them a different rank. Older names for the group include Filicidae and Filicales, although at least the "water ferns" (now the Salviniales) were then treated separately. The leptosporangiate ferns are one of the four major groups of ferns, with the other three being the eusporangiate ferns comprising the marattioid ferns (Marattiidae, Marattiaceae), the horsetails (Equisetiidae, Equisetaceae), and whisk ferns and moonworts. There are approximately 8465 species of living leptosporangiate ferns, compared with about 2070 for all other ferns, totalling 10535 species of ferns. Almost a third of leptosporangiate fern species are epiphytes. These ...
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Fern
The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissues that conduct water and nutrients, and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaf, leaves called megaphylls that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled Fiddlehead fern, fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae (plant), Polypodiidae) and eusporangiate ferns, the latter group including horsetails, Psilotaceae, whisk ferns, marattioid ferns, and ophioglossoid ferns. The fern crown group, consisting of the leptosporangiates and ...
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Dipteris Conjugata
''Dipteris'' is a genus of about seven species of ferns, native to tropical regions across the world, particularly Asia, with a species in northeastern Queensland in Australia. It is one of two genera in the family Dipteridaceae. Description Species of ''Dipteris'' grow from creeping rhizomes, and have large stalks to the sporangium and annulus. The rhizomes have bristles (or hairs) and the fronds have uniseriate hairs (having one line or series). All species of ''Dipteris'' have spore-capsules that are carried on the lower surface of the broad lobed frond.A. C. Seward The fronds can reach up to 50 cm long. Taxonomy Caspar Georg Carl Reinwardt first published the genus in 1825, by describing ''Dipteris conjugata'' Reinw. which is the best known species. In 1839, R. Brown reduced the genus to a subgenus of ''Polypodium''. In 1901, Konrad Christ published ''Die Farnkrauter der Erde't'', within which he included the genus ''Dipteris'' in the family Polypodiaceae, (a subdiv ...
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Mesozoic
The Mesozoic Era is the Era (geology), era of Earth's Geologic time scale, geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Period (geology), Periods. It is characterized by the dominance of archosaurian reptiles such as the dinosaurs, and of Gymnosperm, gymnosperms such as cycads, ginkgoaceae and Araucariaceae, araucarian conifers; a hot Greenhouse and icehouse earth, greenhouse climate; and the tectonic break-up of Pangaea. The Mesozoic is the middle of the three eras since Cambrian explosion, complex life evolved: the Paleozoic, the Mesozoic, and the Cenozoic. The era began in the wake of the Permian–Triassic extinction event, the largest mass extinction in Earth's history, and ended with the Cretaceous–Paleogene extinction event, another mass extinction whose victims included the non-avian dinosaurs, Pterosaur, pterosaurs, Mosasaur, mosasaurs, and Plesiosaur, plesiosaurs. The Mesozoic was a time of significant tectonic, climatic, an ...
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Fossil Taxon
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geologic time, and assess the interactions between prehistoric organisms and their natural environment. While paleontological observations are known from at least the 6th century BC, the foundation of paleontology as a science dates back to the work of Georges Cuvier in 1796. Cuvier demonstrated evidence for the concept of extinction and how life of the past was not necessarily the same as that of the present. The field developed rapidly over the course of the following decades, and the French word ''paléontologie'' was introduced for the study in 1822, which was derived from the Ancient Greek word for "ancient" and words describing relatedness and a field of study. Further advances in the field accompanied the work of Charles Darwin who popula ...
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