The yellow grouper (''Epinephelus awoara''), also known as the banded grouper, is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...

, a

grouper Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...

from the subfamily

Epinephelinae Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...

which is part of the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some ca ...

, which also includes the

anthias Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name Anthi ...

and sea basses. It is found in eastern Asian waters of the

Western Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

. Its natural

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s are shallow

sea The sea, connected as the world ocean or simply the ocean, is the body of salty water that covers approximately 71% of the Earth's surface. The word sea is also used to denote second-order sections of the sea, such as the Mediterranean Sea, ...

s and rocky reefs.

Description

The yellow grouper has a body with a standard length which is 2.7 to 3.3 times greater than its depth. The dorsal profile of the head is convex between its eyes, and overall it is notably convex. The preopercle has an angle of less than 90° and there are 2-5 robust spines at the angle. The

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...

contains 11 spines and 15-16 soft rays while the anal fin has 3 spines and 8 soft rays. The membranes between the dorsal fin spines are deeply incised. The

pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

s are longer than the pelvic fins and the

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

is convex. The head and body are pale greyish brown on the upperparts and are normally golden yellow on the underparts, There are 4 wide dark bars on the upper portion of the body with one on the caudal peduncle and sometimes there is another showing on the nape. The head and body are marked with many small yellow spots. The fins are marked small greyish white spots while the soft rayed part of the dorsal fin and the caudal fin, as well as occasionally the anal fin, have a prominent yellow margin. The pectoral and pelvic fins are dusky yellow and there is a yellow moustache along the

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...

ry groove. The maximum published

total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...

recorded for this species is but a length of around is more common.

Distribution

The yellow grouper is found in the western Pacific Ocean from southern Japan and Korea to Vietnam, including the

South China Sea The South China Sea is a marginal sea of the Western Pacific Ocean. It is bounded in the north by the shores of South China (hence the name), in the west by the Indochinese Peninsula, in the east by the islands of Taiwan and northwestern Phi ...

, East China Sea,

Paracel Islands The Paracel Islands, also known as the Xisha Islands () and the Hoang Sa Archipelago ( vi, Quần đảo Hoàng Sa, lit=Yellow Sand Archipelago), are a disputed archipelago in the South China Sea. The archipelago includes about 130 small coral ...

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...

and the

Sea of Japan The Sea of Japan is the marginal sea between the Japanese archipelago, Sakhalin, the Korean Peninsula, and the mainland of the Russian Far East. The Japanese archipelago separates the sea from the Pacific Ocean. Like the Mediterranean Sea, i ...

Habitat and biology

The yellow grouper occurs over rock areas as well as over areas of sandy substrate, it can also ben found on coral reefs. Juveniles are found in tidal pools. The adults are found at depths of . It is an aggressive species which frequently chases and bites other fishes, especially those of its own species. It is a predatory species which preys on

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can ...

s such as shrimp and crabs, as well as fish and cephalopods. The yellow grouper is a

protogynous hermaphrodite Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods, and plants. Sequential hermaphroditism occurs when the individual changes its sex at some point in its life. In particular, ...

and, in Hong Kong,

spawning Spawn is the eggs and sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of releasing the eggs and sperm, and the act of both sexes is called spawning. Most aquatic animals, except for aquat ...

takes place from February and March through to May, in Taiwan these fish spawn in June and July, while in

Zhejiang Zhejiang ( or , ; , Chinese postal romanization, also romanized as Chekiang) is an East China, eastern, coastal Provinces of China, province of the People's Republic of China. Its capital and largest city is Hangzhou, and other notable citie ...

, China from May to July.

Taxonomy

The yellow grouper was first formally described as ''Serranus awoara'' in 1842 by the

Dutch Dutch commonly refers to: * Something of, from, or related to the Netherlands * Dutch people () * Dutch language () Dutch may also refer to: Places * Dutch, West Virginia, a community in the United States * Pennsylvania Dutch Country People E ...

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and d ...

Coenraad Jacob Temminck (1778-1858) and his student, the

German German(s) may refer to: * Germany (of or related to) ** Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...

ichthyologist

Hermann Schlegel Hermann Schlegel (10 June 1804 – 17 January 1884) was a German ornithologist, herpetologist and ichthyologist. Early life and education Schlegel was born at Altenburg, the son of a brassfounder. His father collected butterflies, which stimulate ...

(1804-1884), with the type locality given as

Nagasaki is the capital and the largest Cities of Japan, city of Nagasaki Prefecture on the island of Kyushu in Japan. It became the sole Nanban trade, port used for trade with the Portuguese and Dutch during the 16th through 19th centuries. The Hi ...

Utilisation

The yellow grouper is a species of high economic value as a food fish and is caught with trawls and hook and line. It is grown in aquaculture but this appears to be reliant on wild caught fry. In some countries it is used as an ornamental fish due to the attractive colours it shows. Female yellow groupers have been crossed with male '' Epinephelus tukula'' using artificial insemination to produce hybrids which have characteristics more desirable for aquaculture.

References

{{Taxonbar, from=Q672389 Yellow grouper Fish described in 1842 Taxonomy articles created by Polbot