Wild yak
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The wild yak (''Bos mutus'') is a large, wild bovine native to the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
. It is the ancestor of the domestic yak (''Bos grunniens'').


Taxonomy

The ancestor of the wild and domestic yak is thought to have diverged from '' Bos primigenius'' at a point between one and five million years ago. The wild yak is now normally treated as a separate species from the domestic yak (''Bos grunniens'').


Description

The wild yak is among the largest extant bovid species. Adults stand about tall at the shoulder, and weigh . The head and body length is , not counting the tail of .Han Jianlin, M. Melletti, J. Burton, 2014, Wild yak (Bos mutus Przewalski, 1883), Ecology, Evolution and Behavior of Wild Cattle: Implications for Conservation, Chapter 1, p.203,
Cambridge University Press Cambridge University Press is the university press of the University of Cambridge. Granted letters patent by King Henry VIII in 1534, it is the oldest university press in the world. It is also the King's Printer. Cambridge University Pr ...
The females are about one-third the weight and are about 30% smaller in their linear dimensions when compared to bull wild yaks. Domesticated yaks are somewhat smaller. They are heavily built animals with a bulky frame, sturdy legs, and rounded cloven hooves. To protect against the cold, the udder in females and the
scrotum The scrotum or scrotal sac is an anatomical male reproductive structure located at the base of the penis that consists of a suspended dual-chambered sac of skin and smooth muscle. It is present in most terrestrial male mammals. The scrotum co ...
in males are small, and covered in a layer of hair. Females have four teats. Both sexes have long shaggy hair, with a dense woolly undercoat over the chest, flanks, and thighs for insulation against the cold. In males especially, this undercoat may form a long "skirt" that can reach the ground. The tail is long and horse-like, rather than tufted like the tails of cattle or
bison Bison are large bovines in the genus ''Bison'' (Greek: "wild ox" (bison)) within the tribe Bovini. Two extant and numerous extinct species are recognised. Of the two surviving species, the American bison, ''B. bison'', found only in North A ...
. The coat is typically black or dark brown, covering most of the body, with a grey muzzle (although some wild golden-brown individuals have been reported). Wild yaks with gold coloured hair are known as the wild golden yak (). They are considered an endangered subspecies in China, with an estimated population of 170 left in the wild. Two morphological types have been identified, so-called ''Qilian'' and ''Kunlun''.


Distribution and habitat

Wild yaks once ranged up to southern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
to the east of
Lake Baikal Lake Baikal (, russian: Oзеро Байкал, Ozero Baykal ); mn, Байгал нуур, Baigal nuur) is a rift lake in Russia. It is situated in southern Siberia, between the federal subjects of Irkutsk Oblast to the northwest and the ...
, but became extinct in Russia around the 17th century. Today, wild yaks are found primarily in northern
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
and western
Qinghai Qinghai (; alternately romanized as Tsinghai, Ch'inghai), also known as Kokonor, is a landlocked province in the northwest of the People's Republic of China. It is the fourth largest province of China by area and has the third smallest po ...
, with some populations extending into the southernmost parts of
Xinjiang Xinjiang, SASM/GNC: ''Xinjang''; zh, c=, p=Xīnjiāng; formerly romanized as Sinkiang (, ), officially the Xinjiang Uygur Autonomous Region (XUAR), is an autonomous region of the People's Republic of China (PRC), located in the northwes ...
, and into
Ladakh Ladakh () is a region administered by India as a union territory which constitutes a part of the larger Kashmir region and has been the subject of dispute between India, Pakistan, and China since 1947. (subscription required) Quote: "Jammu a ...
in India. Small, isolated populations of wild yak are also found farther afield, primarily in western Tibet and eastern Qinghai. In historic times, wild yaks were also found in
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainou ...
, but they are now considered extinct there. The primary habitat of wild yaks consists of treeless uplands between , dominated by mountains and
plateau In geology and physical geography, a plateau (; ; ), also called a high plain or a tableland, is an area of a highland consisting of flat terrain that is raised sharply above the surrounding area on at least one side. Often one or more sides ...
s. They are most commonly found in
alpine tundra Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alpine tundra gets ...
with a relatively thick carpet of grasses and sedges rather than the more barren
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate gras ...
country. The wild yak was thought to be regionally extinct in
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
in the 1970s, but was rediscovered in Humla in 2014. This discovery later made the species to be painted on Nepal's currency.


Behaviour and ecology

The diet of wild yaks consists largely of grasses and sedges, such as '' Carex'', '' Stipa'', and '' Kobresia''. They also eat a smaller amount of herbs, winterfat shrubs, and mosses, and have even been reported to eat
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Himalayan wolf, but Himalayan black bears,
Himalayan brown bear The Himalayan brown bear (''Ursus arctos isabellinus''), also known as the Himalayan red bear, isabelline bear or Dzu-Teh, is a subspecies of the brown bear and is known from northern Afghanistan, northern Pakistan, northern India, west China an ...
s and snow leopards have also been reported as predators in some areas, likely of young or infirm wild yaks. Thubten Jigme Norbu, the elder brother of the
14th Dalai Lama The 14th Dalai Lama (spiritual name Jetsun Jamphel Ngawang Lobsang Yeshe Tenzin Gyatso, known as Tenzin Gyatso (Tibetan: བསྟན་འཛིན་རྒྱ་མཚོ་, Wylie: ''bsTan-'dzin rgya-mtsho''); né Lhamo Thondup), known as ...
, reported on his journey from Kumbum in
Amdo Amdo ( am˥˥.to˥˥ ) is one of the three traditional Tibetan regions, the others being U-Tsang in the west and Kham in the east. Ngari (including former Guge kingdom) in the north-west was incorporated into Ü-Tsang. Amdo is also the bi ...
to
Lhasa Lhasa (; Lhasa dialect: ; bo, text=ལྷ་ས, translation=Place of Gods) is the urban center of the prefecture-level Lhasa City and the administrative capital of Tibet Autonomous Region in Southwest China. The inner urban area of Lhasa ...
in 1950: Wild yaks are
herd A herd is a social group of certain animals of the same species, either wild or domestic. The form of collective animal behavior associated with this is called '' herding''. These animals are known as gregarious animals. The term ''herd'' i ...
animals. Herds can contain several hundred individuals, although many are much smaller. Herds consist primarily of females and their young, with a smaller number of adult males. On average female yaks graze 100m higher than males. Females with young tend to choose grazing ground on high, steep slopes. The remaining males are either solitary, or found in much smaller groups, averaging around six individuals. Groups move into lower altitude ranges during the winter. Although wild yaks can become aggressive when defending young, or during the rut, they generally avoid humans, and may flee for great distances if approached.


Reproduction

Wild yaks mate in summer and give birth to a single calf the following spring.Wiener, G.; Jianlin, H.; Ruijun, L. (2003)
"4 The Yak in Relation to Its Environment"
''The Yak, Second Edition''. Bangkok: Regional Office for Asia and the Pacific Food and Agriculture Organization of the United Nations, . Accessed 8 August 2008.
Females typically only give birth every other year.


Conservation

The wild yak is currently listed as Vulnerable on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
. It was previously classified as
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
, but was downlisted in 1996 based on the estimated rate of population decline and current population sizes. The latest assessment in 2008 suggested a total population of no more than 10,000 mature individuals. The wild yak is experiencing threats applied by several sources. Poaching, including commercial poaching, has remained the most serious threat; males are particularly impacted because of their more solitary habits. Disturbance by and interbreeding with livestock herds is also common. This may include the transmission of cattle-borne diseases, although no direct evidence of this has yet been found. Conflicts with herders themselves, as in preventive and retaliatory killings for abduction of domestic yaks by wild herds, also occur but appear to be relatively rare. Recent protection from poaching particularly appears to have stabilized or even increased population sizes in several areas, leading to the IUCN downlisting in 2008. In both China and India, the species is officially protected; in China it is present in a number of large nature reserves.


Impact on humans

The wild yak is a reservoir for
zoonotic diseases A zoonosis (; plural zoonoses) or zoonotic disease is an infectious disease of humans caused by a pathogen (an infectious agent, such as a bacterium, virus, parasite or prion) that has jumped from a non-human (usually a vertebrate) to a ...
of both bacterial and viral origins. Such bacterial diseases include
anthrax Anthrax is an infection caused by the bacterium '' Bacillus anthracis''. It can occur in four forms: skin, lungs, intestinal, and injection. Symptom onset occurs between one day and more than two months after the infection is contracted. The s ...
, botulism,
tetanus Tetanus, also known as lockjaw, is a bacterial infection caused by ''Clostridium tetani'', and is characterized by muscle spasms. In the most common type, the spasms begin in the jaw and then progress to the rest of the body. Each spasm usually ...
, and
tuberculosis Tuberculosis (TB) is an infectious disease usually caused by '' Mycobacterium tuberculosis'' (MTB) bacteria. Tuberculosis generally affects the lungs, but it can also affect other parts of the body. Most infections show no symptoms, ...
.


References


External links

*ARKive
images and movies of the wild yak ''(Bos grunniens)''
{{Taxonbar, from=Q26547 Mammals described in 1883 Mammals of Nepal Mammals of India Mammals of China Bovines Extant Zanclean first appearances Yaks Taxa named by Nikolay Przhevalsky