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The WILD BOAR (_Sus scrofa_), also known as the WILD SWINE or EURASIAN WILD PIG, is a suid native to much of Eurasia
Eurasia
, North Africa , and the Greater Sunda Islands . Human
Human
intervention has spread its distribution further, making the species one of the widest-ranging mammals in the world, as well as the most widely spread suiform . Its wide range, high numbers, and adaptability mean that it is classed as least concern by the IUCN and it has become an invasive species in part of its introduced range. The animal probably originated in Southeast Asia during the Early Pleistocene
Pleistocene
, and outcompeted other suid species as it spread throughout the Old World .

As of 1990, up to 16 subspecies are recognised, which are divided into four regional groupings based on skull height and lacrimal bone length. The species lives in matriarchal societies consisting of interrelated females and their young (both male and female). Fully grown males are usually solitary outside the breeding season. The grey wolf is the wild boar's main predator throughout most of its range except in the Far East and the Lesser Sunda Islands , where it is replaced by the tiger and Komodo dragon
Komodo dragon
respectively. It has a long history of association with humans , having been the ancestor of most domestic pig breeds and a big-game animal for millennia.

CONTENTS

* 1 Terminology

* 1.1 Hunting

* 2 Taxonomy and evolution

* 2.1 Subspecies
Subspecies
* 2.2 Domestication

* 3 Physical description * 4 Social behaviour and life cycle

* 5 Ecology

* 5.1 Habitat
Habitat
and sheltering behaviour * 5.2 Diet * 5.3 Predators

* 6 Range

* 6.1 Reconstructed range * 6.2 Present range * 6.3 Status in Britain

* 6.4 Introduction to North America

* 6.4.1 History

* 7 Diseases and parasites

* 8 Relationships with humans

* 8.1 In culture * 8.2 As a game animal and food source * 8.3 Crop and garbage raiding * 8.4 Attacks on humans

* 9 See also * 10 Notes * 11 References * 12 Bibliography * 13 Further reading * 14 External links

TERMINOLOGY

As true wild boars became extinct in Britain before the development of modern English, the same terms are often used for both true wild boar and pigs, especially large or semiwild ones. The English 'boar' stems from the Old English
Old English
_bar_, which is thought to be derived from the West Germanic _*bairaz_, of unknown origin. Boar is sometimes used specifically to refer to males, and may also be used to refer to male domesticated pigs, especially breeding males that have not been castrated.

'Sow', the traditional name for a female, again comes from Old English and Germanic; it stems from Proto-Indo-European
Proto-Indo-European
, and is related to the Latin
Latin
_sus_ and Greek _hus_ and more closely to the modern German _Sau_. The young may be called 'piglets'.

The animals' specific name _scrofa_ is Latin
Latin
for 'sow'.

HUNTING

In hunting terminology, boars are given different designations according to their age:

DESIGNATION AGE IMAGE

SQUEAKER 0–10 months

JUVENILE 10–12 months

PIG OF THE SOUNDER Two years

BOAR OF THE 4TH/5TH/6TH YEAR 3–5 years

OLD BOAR Six years

GRAND OLD BOAR Over seven years

"SOLITARY BOAR"

TAXONOMY AND EVOLUTION

_ Skull of Sus strozzii _ ( Museo di Storia Naturale di Firenze ), a Pleistocene
Pleistocene
suid that was outcompeted by _S. scrofa_

MtDNA
MtDNA
studies indicate that the wild boar originated from islands in Southeast Asia such as Indonesia
Indonesia
and the Philippines
Philippines
, and subsequently spread onto mainland Eurasia
Eurasia
and North Africa. The earliest fossil finds of the species come from both Europe
Europe
and Asia, and date back to the Early Pleistocene
Pleistocene
. By the late Villafranchian , _S. scrofa_ largely displaced the related _S. strozzii _, a large, possibly swamp-adapted suid ancestral to the modern _S. verrucosus _ throughout the Eurasian mainland, restricting it to insular Asia. Its closest wild relative is the bearded pig of Malacca
Malacca
and surrounding islands.

SUBSPECIES

As of 2005 , 16 subspecies are recognised, which are divided into four regional groupings:

* WESTERN: Includes _S. s. scrofa_, _S. s. meridionalis_, _S. s. algira_, _S. s. attila_, _S. s. lybicus_, and _S. s. nigripes_. These subspecies are typically high-skulled (though _lybicus_ and some _scrofa_ are low-skulled), with thick underwool and (excepting _scrofa_ and _attila_) poorly developed manes. * INDIAN: Includes _S. s. davidi_ and _S. s. cristatus_. These subspecies have sparse or absent underwool, with long manes and prominent bands on the snout and mouth. While _S. s. cristatus_ is high-skulled, _S. s. davidi_ is low-skulled. * EASTERN: Includes _S. s. sibiricus_, _S. s. ussuricus_, _S. s. leucomystax_, _S. s. riukiuanus_, _S. s. taivanus_, and _S. s. moupinensis_. These subspecies are characterised by a whitish streak extending from the corners of the mouth to the lower jaw. With the exception of _S. s. ussuricus_, most are high-skulled. The underwool is thick, except in _S. s. moupinensis_, and the mane is largely absent. * INDONESIAN: Represented solely by _S. s. vittatus_, it is characterised by its sparse body hair, lack of underwool, fairly long mane, a broad reddish band extending from the muzzle to the sides of the neck. It is the most basal of the four groups, having the smallest relative brain size, more primitive dentition and unspecialised cranial structure.

SUBSPECIES IMAGE TRINOMIAL AUTHORITY DESCRIPTION RANGE SYNONYMS

CENTRAL EUROPEAN BOAR _S. s. scrofa _ Nominate subspecies _ Linnaeus, 1758 A medium-sized, dark to rusty-brown haired subspecies with long and relatively narrow lacrimal bones Northern Spain
Spain
, northern Italy
Italy
, France
France
, Germany
Germany
, Benelux
Benelux
, Croatia
Croatia
, Belarus
Belarus
, Denmark
Denmark
, Sweden
Sweden
, Lithuania
Lithuania
, Poland
Poland
, Czech Republic , Slovakia
Slovakia
and possibly Albania
Albania
anglicus_ (Reichenbach, 1846), _aper_ (Erxleben, 1777), _asiaticus_ (Sanson, 1878), _bavaricus_ (Reichenbach, 1846), _campanogallicus_ (Reichenbach, 1846), _capensis_ (Reichenbach, 1846), _castilianus_ (Thomas, 1911), _celticus_ (Sanson, 1878), _chinensis_ (Linnaeus, 1758), _crispus_ (Fitzinger, 1858), _deliciosus_ (Reichenbach, 1846), _domesticus _ (Erxleben, 1777), _europaeus_ (Pallas, 1811), _fasciatus_ (von Schreber, 1790), _ferox_ (Moore, 1870), _ferus_ (Gmelin, 1788), _gambianus_ (Gray, 1847), _hispidus_ (von Schreber, 1790), _hungaricus_ (Reichenbach, 1846), _ibericus_ (Sanson, 1878), _italicus_ (Reichenbach, 1846), _juticus_ (Fitzinger, 1858), _lusitanicus_ (Reichenbach, 1846), _macrotis_ (Fitzinger, 1858), _monungulus_ (G. Fischer , 1814), _moravicus_ (Reichenbach, 1846), _nanus_ (Nehring, 1884), _palustris_ (Rütimeyer, 1862), _pliciceps_ (Gray, 1862), _polonicus_ (Reichenbach, 1846), _sardous_ (Reichenbach, 1846), _scropha_ (Gray, 1827), _sennaarensis_ (Fitzinger, 1858), _sennaarensis_ (Gray, 1868), _sennaariensis_ (Fitzinger, 1860), _setosus_ (Boddaert, 1785), _siamensis_ (von Schreber, 1790), _sinensis_ (Erxleben, 1777), _suevicus_ (Reichenbach, 1846), _syrmiensis_ (Reichenbach, 1846), _turcicus_ (Reichenbach, 1846), _variegatus_ (Reichenbach, 1846), _vulgaris_ (S. D. W., 1836), _wittei_ (Reichenbach, 1846)

NORTH AFRICAN BOAR _S. s. algira_ _ Loche, 1867 Sometimes considered a junior synonym of S. s. scrofa_, but smaller and with proportionally longer tusks Tunisia
Tunisia
, Algeria
Algeria
and Morocco
Morocco
_barbarus_ (Sclater, 1860)

_sahariensis_ (Heim de Balzac, 1937)

CARPATHIAN BOAR _S. s. attila _ _ Thomas, 1912 A large-sized subspecies with long lacrimal bones and dark hair, though lighter-coloured than S. s. scrofa_ Romania
Romania
, Hungary
Hungary
, Ukraine
Ukraine
, Balkans
Balkans
, Caucasus
Caucasus
, Transcaucasia , Caspian coast, Asia Minor
Asia Minor
and northern Iran
Iran
_falzfeini_ (Matschie, 1918)

INDIAN BOAR _S. s. cristatus _ _ Wagner, 1839 A long-maned subspecies with a coat that is brindled black unlike S. s. davidi_, it is more lightly built than _S. s. scrofa_. Its head is larger and more pointed than that of _S. s. scrofa_, and its ears smaller and more pointed. The plane of the forehead is straight, while it is concave in _S. s. scrofa_. India
India
, Nepal
Nepal
, Burma
Burma
, western Thailand
Thailand
and Sri Lanka
Sri Lanka
_affinis_ (Gray, 1847), _aipomus_ (Gray, 1868), _aipomus_ (Hodgson, 1842), _bengalensis_ (Blyth, 1860), _indicus_ (Gray, 1843), _isonotus_ (Gray, 1868), _isonotus_ (Hodgson, 1842), _jubatus_ (Miller, 1906), _typicus_ (Lydekker, 1900), _zeylonensis_ (Blyth, 1851)

CENTRAL ASIAN BOAR _S. s. davidi _

Groves, 1981 A small, long-maned and light brown subspecies Pakistan
Pakistan
and northwest India
India
to southeastern Iran

JAPANESE BOAR _S. s. leucomystax _ _ Temminck, 1842 A small, almost maneless, yellowish-brown subspecies All of Japan
Japan
, save for Hokkaido
Hokkaido
and the Ryukyu Islands japonica_ (Nehring, 1885)

_nipponicus_ (Heude, 1899)

ANATOLIAN BOAR _S. s. libycus _ _ Gray, 1868 A small, pale and almost maneless subspecies Transcaucasia , Turkey
Turkey
, Levant
Levant
, Israel
Israel
and former Yugoslavia lybicus_ (Groves, 1981)

_mediterraneus_ (Ulmansky, 1911) _reiseri_ (Bolkay, 1925)

MAREMMAN BOAR _S. s. majori_ _ De Beaux and Festa, 1927 Smaller than S. s. scrofa_, with a higher and wider skull, since the 1950s, it has crossed extensively with _S. s. scrofa_, largely due to the two being kept together in meat farms and artificial introductions by hunters of _S. s. scrofa_ specimens into _S. s. majori_ habitats. Its separation from _S. s. scrofa_ is doubtful. Maremma (central Italy)

MEDITERRANEAN BOAR _S. s. meriodionalis_ _ Forsyth Major, 1882

Andalusia
Andalusia
, Corsica and Sardinia
Sardinia
baeticus_ (Thomas, 1912)

_sardous_ (Ströbel, 1882)

NORTHERN CHINESE BOAR _S. s. moupinensis _ _ Milne-Edwards, 1871 There are significant variations within this subspecies, and it is possible there are actually several subspecies involved. Coastal China
China
south to Vietnam
Vietnam
and west to Sichuan
Sichuan
acrocranius_ (Heude, 1892), _chirodontus_ (Heude, 1888), _chirodonticus_ (Heude, 1899), _collinus_ (Heude, 1892), _curtidens_ (Heude, 1892), _dicrurus_ (Heude, 1888), _flavescens_ (Heude, 1899), _frontosus_ (Heude, 1892), _laticeps_ (Heude, 1892), _leucorhinus_ (Heude, 1888), _melas_ (Heude, 1892), _microdontus_ (Heude, 1892), _oxyodontus_ (Heude, 1888), _paludosus_ (Heude, 1892), _palustris_ (Heude, 1888), _planiceps_ (Heude, 1892), _scrofoides_ (Heude, 1892), _spatharius_ (Heude, 1892), _taininensis_ (Heude, 1888)

MIDDLE ASIAN BOAR _S. s. nigripes_

Blanford, 1875 A light coloured subspecies with black legs which, though varied in size, it is generally quite large, the lacrimal bones and facial region of the skull are shorter than those of _S. s. scrofa_ and _S. s. attila_. Middle Asia , Kazakhstan
Kazakhstan
, eastern Tien Shan , western Mongolia
Mongolia
, Kashgar and possibly Afghanistan
Afghanistan
and southern Iran

RYUKYU BOAR _S. s. riukiuanus_

Kuroda, 1924 A small subspecies Ryukyu Islands

TRANS-BAIKAL BOAR _S. s. sibiricus_

Staffe, 1922 The smallest subspecies of the former Soviet
Soviet
region, it has dark brown, almost black hair and a light grey patch extending from the cheeks to the ears. The skull is squarish and the lacrimal bones short. Baikal
Baikal
, Transbaikalia , northern and northeastern Mongolia _raddeanus_ (Adlerberg, 1930)

FORMOSAN BOAR _S. s. taivanus_

Swinhoe, 1863 A small blackish subspecies Taiwan
Taiwan

USSURI BOAR _S. s. ussuricus_

Heude, 1888 The largest subspecies, it has usually dark hair and a white band extending from the corners of the mouth to the ears. The lacrimal bones are shortened, but longer than those of _S. s. sibiricus_. Eastern China
China
, Ussuri and Amur bay _canescens_ (Heude, 1888), _continentalis_ (Nehring, 1889), _coreanus_ (Heude, 1897), _gigas_ (Heude, 1892), _mandchuricus_ (Heude, 1897), _songaricus_ (Heude, 1897)

BANDED PIG _S. s. vittatus _ _ Boie, 1828 A small, short-faced and sparsely furred subspecies with a white band on the muzzle; it might be a separate species, and shows some similarities with some other suid species in Southeast Asia. From Peninsular Malaysia , and in Indonesia
Indonesia
from Sumatra
Sumatra
and Java east to Komodo andersoni_ (Thomas and Wroughton, 1909), _jubatulus_ (Miller, 1906), _milleri_ (Jentink, 1905), _pallidiloris_ (Mees, 1957), _peninsularis_ (Miller, 1906), _rhionis_ (Miller, 1906), _typicus_ (Heude, 1899)

Wild boar
Wild boar
(left) and domestic pig (right) skulls: Note the greatly shortened facial region of the latter.

DOMESTICATION

Male, domestic pig-wild boar cross

With the exception of domestic pigs in Timor
Timor
and Papua New Guinea (which appear to be of Sulawesi warty pig stock), the wild boar is the ancestor of most pig breeds. Archaeological evidence suggests that pigs were domesticated from wild boar as early as 13,000–12,700 BC in the Near East
Near East
in the Tigris
Tigris
Basin being managed in the wild in a way similar to the way they are managed by some modern New Guineans. Remains of pigs have been dated to earlier than 11,400 BC in Cyprus
Cyprus
. Those animals must have been introduced from the mainland, which suggests domestication in the adjacent mainland by then. There was also a separate domestication in China
China
which took place about 8000 years ago. DNA
DNA
evidence from sub-fossil remains of teeth and jawbones of Neolithic
Neolithic
pigs shows that the first domestic pigs in Europe
Europe
had been brought from the Near East. This stimulated the domestication of local European wild boar resulting in a third domestication event with the Near Eastern genes dying out in European pig stock. Modern domesticated pigs have involved complex exchanges, with European domesticated lines being exported in turn to the ancient Near East. Historical records indicate that Asian pigs were introduced into Europe
Europe
during the 18th and early 19th centuries. Domestic pigs tend to have much more developed hindquarters than their wild boar ancestors, to the point where 70% of their body weight is concentrated in the posterior, which is the opposite of wild boar, where most of the muscles are concentrated on the head and shoulders.

PHYSICAL DESCRIPTION

Dentition, as illustrated by Charles Knight

The wild boar is a bulky, massively built suid with short and relatively thin legs. The trunk is short and massive, while the hindquarters are comparatively underdeveloped. The region behind the shoulder blades rises into a hump, and the neck is short and thick, to the point of being nearly immobile. The animal's head is very large, taking up to one third of the body's entire length. The structure of the head is well suited for digging. The head acts as a plow, while the powerful neck muscles allow the animal to upturn considerable amounts of soil: it is capable of digging 8–10 cm (3.1–3.9 in) into frozen ground and can upturn rocks weighing 40–50 kg (88–110 lb). The eyes are small and deep-set, and the ears long and broad. The species has well developed canine teeth , which protrude from the mouths of adult males. The middle hooves are larger and more elongated than the lateral ones, and are capable of quick movements. The animal can run at a maximum speed of 40 km/h and jump at a height of 140–150 cm (55–59 in). Sexual dimorphism is very pronounced in the species, with males being typically 5–10% larger and 20–30% heavier than females. Males also sport a mane running down the back, which is particularly apparent during autumn and winter. The canine teeth are also much more prominent in males, and grow throughout life. The upper canines are relatively short and grow sideways early in life, though gradually curve upwards. The lower canines are much sharper and longer, with the exposed parts measuring 10–12 cm (3.9–4.7 in) in length. In the breeding period, males develop a coating of subcutaneous tissue , which may be 2–3 cm (0.79–1.18 in) thick, extending from the shoulder blades to the rump, thus protecting vital organs during fights. Males sport a roughly egg-sized sack near the opening of the penis, which collects urine and emits a sharp odour. The purpose of this is not fully understood. Skeleton, as illustrated by Richard Lydekker . A European wild boar piglet, painted by Hans Hoffman in 1578. Note the stripes, a characteristic feature of piglets.

Adult size and weight is largely determined by environmental factors; boars living in arid areas with little productivity tend to attain smaller sizes than their counterparts inhabiting areas with abundant food and water. In most of Europe
Europe
, males average 75–100 kg (165–220 lb) in weight, 75–80 cm (30–31 in) in shoulder height and 150 cm (59 in) in body length, whereas females average 60–80 kg (130–180 lb) in weight, 70 cm (28 in) in shoulder height and 140 cm (55 in) in body length. In Europe's Mediterranean regions , males may reach average weights as low as 50 kg (110 lb) and females 45 kg (99 lb), with shoulder heights of 63–65 cm (25–26 in). In the more productive areas of Eastern Europe
Europe
, males average 110–130 kg (240–290 lb) in weight, 95 cm (37 in) in shoulder height and 160 cm (63 in) in body length, while females weigh 95 kg (209 lb), reach 85–90 cm (33–35 in) in shoulder height and 145 cm (57 in) in body length. In Western and Central Europe
Europe
, the largest males weigh 200 kg (440 lb) and females 120 kg (260 lb). In Eastern Europe, large males can reach brown bear -like sizes, weighing 270 kg (600 lb) and measuring 110–118 cm (43–46 in) in shoulder height. Some adult males in Ussuriland and Manchuria
Manchuria
have been recorded to weigh 300–350 kg (660–770 lb) and measure 125 cm (49 in) in shoulder height. Adults of this size are generally immune from wolf predation. Such giants are rare in modern times, due to past overhunting preventing animals from attaining their full growth.

The winter coat consists of long, coarse bristles underlaid with short brown downy fur. The length of these bristles varies along the body, with the shortest being around the face and limbs and the longest running along the back. These back bristles form the aforementioned mane prominent in males, and stand erect when the animal is agitated. Colour is highly variable; specimens around Lake Balkhash are very lightly coloured, and can even be white, while some boars from Belarus
Belarus
and Ussuriland can be black. Some subspecies sport a light coloured patch running backwards from the corners of the mouth. Coat colour also varies with age, with piglets having light brown or rusty-brown fur with pale bands extending from the flanks and back.

The wild boar produces a number of different sounds which are divided into three categories:

* CONTACT CALLS: Grunting noises which differ in intensity according to the situation. Adult males are usually silent, while females frequently grunt and piglets whine. When feeding, boars express their contentment through purring. Studies have shown that piglets imitate the sounds of their mother, thus different litters may have unique vocalisations. * ALARM CALLS: Warning cries emitted in response to threats. When frightened, boars make loud huffing _ukh! ukh!_ sounds or emit screeches transcribed as _gu-gu-gu_. * COMBAT CALLS: High-pitched, piercing cries.

Its sense of smell is very well developed, to the point that the animal is used for drug detection in Germany. Its hearing is also acute, though its eyesight is comparatively weak, lacking colour vision and being unable to recognise a standing human 10–15 metres away.

Pigs are one of four known mammalian species which possess mutations in the nicotinic acetylcholine receptor that protect against snake venom. Mongooses , honey badgers , hedgehogs , and pigs all have modifications to the receptor pocket which prevents the snake venom α-neurotoxin from binding. These represent four separate, independent mutations.

SOCIAL BEHAVIOUR AND LIFE CYCLE

Boars are typically social animals, living in female-dominated sounders consisting of barren sows and mothers with young led by an old matriarch. Male boars leave their sounder at the age of 8–15 months, while females either remain with their mothers or establish new territories nearby. Subadult males may live in loosely knit groups, while adult and elderly males tend to be solitary outside the breeding season. _ Central European wild boar (S. s. scrofa_) piglets suckling

The breeding period in most areas lasts from November to January, though most mating only lasts a month and a half. Prior to mating, the males develop their subcutaneous armour, in preparation for confronting rivals. The testicles double in size and the glands secrete a foamy yellowish liquid. Once ready to reproduce, males travel long distances in search of a sounder of sows, eating little on the way. Once a sounder has been located, the male drives off all young animals and persistently chases the sows. At this point, the male fiercely fights potential rivals, A single male can mate with 5–10 sows. By the end of the rut, males are often badly mauled and have lost 20% of their body weight, with bite-induced injuries to the penis being common. The gestation period varies according to the age of the expecting mother. For first time breeders, it lasts 114–130 days, while it lasts 133–140 days in older sows. Farrowing occurs between March and May, with litter sizes depending on the age and nutrition of the mother. The average litter consists of 4–6 piglets, with the maximum being 10–12. The piglets are whelped in a nest constructed from twigs, grasses and leaves. Should the mother die prematurely, the piglets are adopted by the other sows in the sounder.

Newborn piglets weigh around 600–1,000 grams, lacking underfur and bearing a single milk incisor and canine on each half of the jaw. There is intense competition between the piglets over the most milk-rich nipples, as the best fed young grow faster and have stronger constitutions. The piglets do not leave the lair for their first week of life. Should the mother be absent, the piglets lie closely pressed to each other. By two weeks of age, the piglets begin accompanying their mother on her journeys. Should danger be detected, the piglets take cover or stand immobile, relying on their camouflage to keep them hidden. The neonatal coat fades after three months, with adult colouration being attained at eight months. Although the lactation period lasts 2.5–3.5 months, the piglets begin displaying adult feeding behaviours at the age of 2–3 weeks. The permanent dentition is fully formed by 1–2 years. With the exception of the canines in males, the teeth stop growing during the middle of the fourth year. The canines in old males continue to grow throughout their lives, curving strongly as they age. Sows attain sexual maturity at the age of one year, with males attaining it a year later. However, estrus usually first occurs after two years in sows, while males begin participating in the rut after 4–5 years, as they are not permitted to mate by the older males. The maximum lifespan in the wild is 10–14 years, though few specimens survive past 4–5 years. Boars in captivity have lived for 20 years.

ECOLOGY

HABITAT AND SHELTERING BEHAVIOUR

An individual from higher ridges of Himalayas
Himalayas
at 9,600 ft in Pangolakha Wildlife Sanctuary
Pangolakha Wildlife Sanctuary
, Sikkim
Sikkim
, India
India
. Wild boar frequently wallow in mud, possibly to regulate temperature or remove parasites

The wild boar inhabits a diverse array of habitats from boreal taigas to deserts . In mountainous regions, it can even occupy alpine zones, occurring up to 1,900 metres in the Carpathians
Carpathians
, 2,600 metres in the Caucasus
Caucasus
and up to 3,600–4,000 metres in the mountains in Central Asia and Kazakhstan
Kazakhstan
. In order to survive in a given area, wild boars require a habitat fulfilling three conditions: heavily brushed areas providing shelter from predators, water for drinking and bathing purposes and an absence of regular snowfall. The main habitats favoured by boars in Europe
Europe
are deciduous and mixed forests , with the most favourable areas consisting of forest composed of oak and beech enclosing marshes and meadows . In the Białowieża Forest , the animal's primary habitat consists of well developed, broad-leaved and mixed forests, along with marshy mixed forests, with coniferous forests and undergrowths being of secondary importance. Forests made up entirely of oak groves and beech are used only during the fruit-bearing season. This is in contrast to the Caucasian and Transcaucasian mountain areas, where boars will occupy such fruit-bearing forests year-round. In the mountainous areas of the Russian Far East , the species inhabits nutpine groves, hilly mixed forests where Mongolian oak and Korean pine are present, swampy mixed taiga and coastal oak forests. In Transbaikalia , boars are restricted to river valleys with nutpine and shrubs. Boars are regularly encountered in pistachio groves in winter in some areas of Tajikistan and Turkmenia , while in spring they migrate to open deserts; boar have also colonised deserts in several areas they have been introduced to. On the islands of Komodo and Rinca
Rinca
, the boar mostly inhabits savanna or open monsoon forests, avoiding heavily forested areas unless pursued by humans. Wild boar
Wild boar
are known to be competent swimmers, capable of covering long distances. In 2013, one boar was reported to have completed the seven mile swim from France
France
to Alderney in the Channel Islands
Channel Islands
. Due to concerns about disease it was shot and incinerated.

Wild boar
Wild boar
rest in shelters, which contain insulating material like spruce branches and dry hay . These resting places are occupied by whole families (though males lie separately), and are often located in the vicinity of streams, in swamp forests, in tall grass or shrub thickets. Boars never defecate in their shelters, and will cover themselves with soil and pine needles when irritated by insects.

DIET

_ Male Indian boar (S. s. cristatus_) feeding on a chital carcass

The wild boar is a highly versatile omnivore , whose diversity in choice of food rivals that of humans . Their foods can be divided into four categories:

* Rhizomes , roots , tubers and bulbs , all of which are dug up throughout the year in the animal's whole range. * Nuts , berries , and seeds , which are consumed when ripened and are dug up from the snow when abundant. * Leaves
Leaves
, bark , twigs, and shoots , along with garbage . * Earthworms , insects , mollusks , fish , rodents , insectivores , bird eggs , lizards , snakes , frogs , and carrion . Most of these prey items are taken in warm periods.

A 50 kg (110 lb) boar needs around 4,000–4,500 calories of food per day, though this required amount increases during winter and pregnancy, with the majority of its diet consisting of food items dug from the ground like underground plant material and burrowing animals. Acorns and beechnuts are invariably its most important food items in temperate zones , as they are rich in the carbohydrates necessary for the buildup of fat reserves needed to survive lean periods. In Western Europe, underground plant material favoured by boars includes bracken , willow herb , bulbs, meadow herb roots and bulbs, and the bulbs of cultivated crops. Such food is favoured in early spring and summer, but may also be eaten in autumn and winter during beechnut and acorn crop failures. Should regular wild foods become scarce, boars will eat tree bark and fungi , as well as visit cultivated potato and artichoke fields. Boar soil disturbance and foraging have been shown to facilitate invasive plants. Boars of the _vittatus_ subspecies in Ujung Kulon National Park in Java
Java
differ from most other populations by their primarily frugivorous diet, which consists of 50 different fruit species, especially figs , thus making them important seed dispersers. The wild boar can consume numerous genera of poisonous plants without ill effect, including _ Aconitum
Aconitum
_, _ Anemone
Anemone
_, _Calla _, _ Caltha _, _ Ferula _, and _ Pteridium _.

Boars may occasionally prey on small vertebrates like newborn deer fawns, leporids and galliform chicks. Boars inhabiting the Volga Delta and near some lakes and rivers of Kazakhstan
Kazakhstan
have been recorded to feed extensively on fish like carp and Caspian roach . Boars in the former area will also feed on cormorant and heron chicks, bivalved molluscs , trapped muskrats and mice. There is at least one record of a boar killing and eating a bonnet macaque in southern India's Bandipur National Park , though this may have been a case of intraguild predation , brought on by interspecific competition for human handouts.

PREDATORS

Tigers killing a wild boar in Kanha Tiger
Tiger
Reserve

Piglets are vulnerable to attack from medium-sized felids like lynx , jungle cats and snow leopards and other carnivorans like brown bears and yellow-throated martens .

The grey wolf is the main predator of wild boar throughout most of its range. A single wolf can kill around 50–80 boars of differing ages in one year. In Italy
Italy
and Belarus' Belovezhskaya Pushcha National Park , boars are the wolf's primary prey, despite an abundance of alternative, less powerful ungulates. Wolves are particularly threatening during the winter, when deep snow impedes the boars' movements. In the Baltic regions, heavy snowfall can allow wolves to eliminate boars from an area almost completely. Wolves primarily target piglets and subadults, and only rarely attack adult sows. Adult males are usually avoided entirely. Dholes may also prey on boars, to the point of keeping their numbers down in northwestern Bhutan, despite there being many more cattle in the area. _ Banded pig (S. s. vittatus_) eaten by Komodo dragons

Leopards are predators of wild boar in the Caucasus, Transcaucasia, the Russian Far East, India, China, and Iran. In most areas, boars constitute only a small part of the leopard's diet. However, in Iran's Sarigol National Park, boars are the second most frequently targeted prey species after mouflon , though adult individuals are generally avoided, as they are above the leopard's preferred weight range of 10–40 kg (22–88 lb). This dependence on wild boar is largely due in part to the local leopard subspecies' large size.

Boars of all ages were once the primary prey of tigers in Transcaucasia, Kazakhstan, Middle Asia and the Far East up until the late 19th century. In modern times, tiger numbers are too low to have a limiting effect on boar populations. A single tiger can systematically destroy an entire sounder by preying on its members one by one, before moving on to another herd. Tigers have been noted to chase boars for longer distances than with other prey. In two rare cases, boars were reported to gore a small tiger and a tigress to death in self-defense. In the Amur region , wild boars are one of the two most important prey species for tigers alongside the Manchurian wapiti , with the two species collectively comprising roughly 80% of the felid's prey. In Sikhote Alin , a tiger can kill 30–34 boars a year. Studies of tigers in India
India
indicate that boars are usually secondary in preference to various cervids and bovids , though when boars are targeted, healthy adults are caught more frequently than young and sick specimens.

On the islands of Komodo , Rinca
Rinca
, and Flores
Flores
, the boar's main predator is the Komodo dragon
Komodo dragon
.

RANGE

RECONSTRUCTED RANGE

The species originally occurred in North Africa
North Africa
and much of Eurasia
Eurasia
; from the British Isles
British Isles
to Korea
Korea
and the Sunda Islands . The northern limit of its range extended from southern Scandinavia
Scandinavia
to southern Siberia
Siberia
and Japan
Japan
. Within this range, it was only absent in extremely dry deserts and alpine zones. It was once found in North Africa
North Africa
along the Nile
Nile
valley up to Khartum and north of the Sahara
Sahara
. The species occurs on a few Ionian and Aegean Islands
Aegean Islands
, sometimes swimming between islands. The reconstructed northern boundary of the animal's Asian range ran from Lake Ladoga (at 60°N) through the area of Novgorod
Novgorod
and Moscow into the southern Urals , where it reached 52°N. From there, the boundary passed Ishim and farther east the Irtysh at 56°N. In the eastern Baraba steppe (near Novosibirsk
Novosibirsk
) the boundary turned steep south, encircled the Altai Mountains
Altai Mountains
, and went again eastward including the Tannu-Ola Mountains and Lake Baikal
Baikal
. From here the boundary went slightly north of the Amur River
Amur River
eastward to its lower reaches at the Sea of Okhotsk
Sea of Okhotsk
. On Sakhalin
Sakhalin
, there are only fossil reports of wild boar. The southern boundaries in Europe
Europe
and Asia were almost invariably identical to the sea shores of these continents. It is absent in the dry regions of Mongolia
Mongolia
from 44–46°N southward, in China
China
westward of Sichuan
Sichuan
and in India
India
north of the Himalayas
Himalayas
. It is absent in the higher elevations of Pamir and Tien Shan , though they do occur in the Tarim basin and on the lower slopes of the Tien Shan.

PRESENT RANGE

In recent centuries, the range of wild boar has changed dramatically, largely due to hunting by humans and more recently because of captive wild boar escaping into the wild. Prior to the 20th century, boar populations had declined in numerous areas, with British populations probably becoming extinct during the 13th century. In Denmark, the last boar was shot at the beginning of the 19th century, and in 1900 they were absent in Tunisia
Tunisia
and Sudan and large areas of Germany, Austria, and Italy. In Russia
Russia
they were extirpated in wide areas in the 1930s. The last boar in Egypt
Egypt
reportedly died on 20 December 1912 in the Giza Zoo , with wild populations having disappeared around 1894–1902. Prince Kamal el Dine Hussein attempted to repopulate Wadi El Natrun with boars of Hungarian stock, but they were quickly exterminated by poachers.

A revival of boar populations began in the middle of the 20th century. By 1950 wild boar had once again reached their original northern boundary in many parts of their Asiatic range. By 1960, they reached Leningrad
Leningrad
and Moscow, and by 1975 they were to be found in Archangelsk and Astrakhan
Astrakhan
. In the 1970s they again occurred in Denmark
Denmark
and Sweden, where captive animals escaped and now survive in the wild. In England, wild boar populations re-established themselves in the 1990s, after escaping from specialist farms that had imported European stock.

STATUS IN BRITAIN

Mixed sounder of wild boar and domestic pigs at Culzie, Scotland

Wild boar
Wild boar
were apparently already becoming rare by the 11th century, since a 1087 forestry law enacted by William the Conqueror
William the Conqueror
punishes through blinding the unlawful killing of boar. Charles I attempted to reintroduce the species into the New Forest
New Forest
, though this population was exterminated during the Civil War .

Between their medieval extinction and the 1980s, when wild boar farming began, only a handful of captive wild boar, imported from the continent, were present in Britain. Occasional escapes of wild boar from wildlife parks have occurred as early as the 1970s, but since the early 1990s significant populations have re-established themselves after escapes from farms, the number of which has increased as the demand for meat from the species has grown. A 1998 MAFF (now DEFRA ) study on wild boar living wild in Britain confirmed the presence of two populations of wild boar living in Britain; one in Kent
Kent
/East Sussex and another in Dorset
Dorset
. Another DEFRA report, in February 2008, confirmed the existence of these two sites as 'established breeding areas' and identified a third in Gloucestershire / Herefordshire
Herefordshire
; in the Forest of Dean
Forest of Dean
/ Ross on Wye area. A 'new breeding population' was also identified in Devon
Devon
. There is another significant population in Dumfries and Galloway. Populations estimates were as follows:

* The largest population, in Kent/East Sussex, was then estimated at approximately 200 animals in the core distribution area. * The second largest, in Gloucestershire/Herefordshire, was first estimated to be in excess of 100 animals. Legally classified as dangerous wild animals, the group is known to be feral descendants of domestic (Tamworth) pigs abandoned nearby. Their numbers grew by 2016 to at least 1500 and the Forestry Commission planned to reduce the total to a manageable 400. "Adult males can reach twenty stone (125 kg), run at thirty miles an hour, and can jump or barge through all but the strongest of fences. Also they are not afraid of humans, so (unlike deer) you can't just shoo them out of your garden." * The smallest, in west Dorset, was estimated to be fewer than 50 animals. * Since winter 2005/6 significant escapes/releases have also resulted in animals colonising areas around the fringes of Dartmoor , in Devon
Devon
. These are considered as an additional single 'new breeding population' and currently estimated to be up to 100 animals.

Population estimates for the Forest of Dean
Forest of Dean
are disputed as at the time that the DEFRA population estimate was 100, a photo of a boar sounder in the forest near Staunton with over 33 animals visible was published, and at about the same time over 30 boar were seen in a field near the original escape location of Weston under Penyard many miles away. In early 2010 the Forestry Commission embarked on a cull, with the aim of reducing the boar population from an estimated 150 animals to 100. By August it was stated that efforts were being made to reduce the population from 200 to 90, but that only 25 had been killed. The failure to meet cull targets was confirmed in February 2011.

Wild boar
Wild boar
have crossed the River Wye
River Wye
into Monmouthshire
Monmouthshire
, Wales. Iolo Williams, the BBC Wales wildlife expert, attempted to film Welsh boar in late 2012. Many other sightings, across the UK, have also been reported. The effects of wild boar on the UK's woodlands were discussed with Ralph Harmer of the Forestry Commission on the BBC Radio's _ Farming Today _ radio programme in 2011. The programme prompted activist writer George Monbiot to propose a thorough population study, followed by the introduction of permit-controlled culling .

INTRODUCTION TO NORTH AMERICA

"Razorbacks " confronting an alligator in Florida
Florida

Wild boars are an invasive species in the Americas, and cause problems including outcompeting native species for food, destroying the nests of ground-nesting species, killing fawns and young domestic livestock, destroying agricultural crops, eating tree seeds and seedlings, destroying native vegetation and wetlands through wallowing, damaging water quality, coming into violent conflict with humans and pets, and carrying pig and human diseases including brucellosis , trichinosis , and pseudorabies . In some jurisdictions, it is illegal to import, breed, release, possess, sell, distribute, trade, transport, hunt, or trap Eurasian boars. Hunting and trapping is done systematically, to increase the chance of eradication and to remove the incentive to illegally release boars, which have mostly been spread deliberately by sport hunters.

History

While domestic pigs, both captive and feral (popularly termed "razorbacks "), have been in North America since the earliest days of European colonization , pure wild boar were not introduced into the New World until the 19th century. The suids were released into the wild by wealthy landowners as big game animals. The initial introductions took place in fenced enclosures, though several escapes occurred, with the escapees sometimes intermixing with already established feral pig populations.

The first of these introductions occurred in New Hampshire
New Hampshire
in 1890. Thirteen wild boar from Germany
Germany
were purchased by Austin Corbin from Carl Hagenbeck , and released into a 9,500 hectare game preserve in Sullivan County . Several of these boars escaped, though they were quickly hunted down by locals. Two further introductions were made since the original stocking, with several escapes taking place due to breaches in the game preserve's fencing. These escapees have ranged widely, with some specimens having been observed crossing into Vermont .

In 1902, 15–20 wild boar from Germany
Germany
were released into a 3,200 hectare estate in Hamilton County, New York . Several specimens escaped six years later, dispersing into the William C. Whitney Wilderness Area , with their descendants surviving for at least 20 years.

The most extensive boar introduction in the US took place in western North Carolina
North Carolina
in 1912, when 13 boars of undetermined European origin were released into two fenced enclosures in a game preserve in Hooper Bald , Graham County . Most of the specimens remained in the preserve for the next decade, until a large-scale hunt caused the remaining animals to break through their confines and escape. Some of the boars migrated to Tennessee
Tennessee
, where they intermixed with both free ranging and feral pigs in the area. In 1924, a dozen Hooper Bald wild pigs were shipped to California
California
and released in a property between Carmel Valley and the Los Padres National Forest . These hybrid boar were later used as breeding stock on various private and public lands throughout the state, as well as in other states like Florida
Florida
, Georgia , South Carolina
South Carolina
, West Virginia
West Virginia
and Mississippi
Mississippi
.

Several wild boars from Leon Springs and the San Antonio , Saint Louis and San Diego Zoos were released in the Powder Horn Ranch in Calhoun County, Texas , in 1939. These specimens escaped and established themselves in surrounding ranchlands and coastal areas, with some crossing the Espiritu Santo Bay and colonising Matagorda Island . Descendants of the Powder Horn Ranch boars were later released onto San José Island and the coast of Chalmette, Louisiana .

Wild boar
Wild boar
of unknown origin were stocked in a ranch in the Edwards Plateau in the 1940s, only to escape during a storm and hybridise with local feral pig populations, later spreading into neighbouring counties.

Starting in the mid-80s, several boars purchased from the San Diego Zoo and Tierpark Berlin were released into the United States. A decade later, more specimens from farms in Canada
Canada
and Białowieża Forest were let loose. In recent years, wild pig populations have been reported in 44 states within the US, most of which are likely wild boar-feral hog hybrids. Pure wild boar populations may still be present, but are extremely localised.

DISEASES AND PARASITES

Lesions consistent with bovine tuberculosis on the lower jaw and lung of a wild boar

Wild boars are known to host at least 20 different parasitic worm species, with maximum infections occurring in summer. Young animals are vulnerable to helminths like _ Metastrongylus _, which are consumed by boars through earthworms, and cause death by parasitising the lungs. Wild boar
Wild boar
also carry parasites known to infect humans, including _ Gastrodiscoides _, _ Trichinella spiralis _, _Taenia solium _, and _ Balantidium coli _. Wild boar
Wild boar
in southern regions are frequently infested with ticks (_ Dermacentor _, _ Rhipicephalus _, and _ Hyalomma _) and hog lice . The species also suffers from blood-sucking flies , which it escapes by bathing frequently or hiding in dense shrubs.

Swine plague spreads very quickly in wild boar, with epizootics being recorded in Germany, Poland, Hungary, Belarus, the Caucasus, the Far East, Kazakhstan, and other regions. Foot-and-mouth disease can also take on epidemic proportions in boar populations. The species occasionally, but rarely contracts Pasteurellosis , hemorrhagic septicemia , tularemia and anthrax . Wild boar
Wild boar
may on occasion contract swine erysipelas through rodents or hog lice and ticks.

RELATIONSHIPS WITH HUMANS

IN CULTURE

Upper Paleolithic cave painting , Altamira , Spain
Spain
. This is one of the earliest known depictions of the species. Depiction of wild boars at Lake Balaton on silver dish (part of the 4th century Sevso Treasure

)

The wild boar features prominently in the cultures of Indo-European people, many of which saw the animal as embodying warrior virtues. Cultures throughout Europe
Europe
and Asia Minor
Asia Minor
saw the killing of a boar as proof of one's valor and strength. Neolithic
Neolithic
hunter gatherers depicted reliefs of ferocious wild boars on their temple pillars at Göbekli Tepe some 11,600 years ago. Virtually all heroes in Greek mythology fight or kill a boar at one point. The demigod Herakles
Herakles
' third labour involves the capture of the Erymanthian Boar
Erymanthian Boar
, Theseus slays the wild sow Phaea , and a disguised Odysseus
Odysseus
is recognised by his handmaiden Eurycleia
Eurycleia
by the scars inflicted on him by a boar during a hunt in his youth. To the mythical Hyperboreans , the boar represented spiritual authority. Several Greek myths use the boar as a symbol of darkness, death and winter. One example is the story of the youthful Adonis
Adonis
, who is killed by a boar and is permitted by Zeus
Zeus
to depart from Hades only during the spring and summer period. This theme also occurs in Irish and Egyptian mythology , where the animal is explicitly linked to the month of October, therefore autumn. This association likely arose from aspects of the boar's actual nature. Its dark colour was linked to the night, while its solitary habits, proclivity to consume crops and nocturnal nature were associated with evil. The foundation myth of Ephesus
Ephesus
has the city being built over the site where prince Androklos of Athens
Athens
killed a boar. Boars were frequently depicted on Greek funerary monuments alongside lions , representing gallant losers who have finally met their match, as opposed to victorious hunters as lions are. The theme of the doomed, yet valorous boar warrior also occurred in Hittite culture, where it was traditional to sacrifice a boar alongside a dog and a prisoner of war after a military defeat.

The boar as a warrior also appears in Scandinavian , Germanic and Anglo-Saxon culture , with its image having been frequently engraved on helmets, shields and swords. According to Tacitus
Tacitus
, the Baltic Aesti featured boars on their helmets, and may have also worn boar masks. The boar and pig were held in particularly high esteem by the Celts
Celts
, who considered them to be their most important sacred animal. Some Celtic deities
Celtic deities
linked to boars include Moccus and Veteris
Veteris
. It has been suggested that some early myths surrounding the Welsh hero Culhwch
Culhwch
involved the character being the son of a boar god. Nevertheless, the importance of the boar as a culinary item among Celtic tribes may have been exaggerated in popular culture by the _ Asterix
Asterix
_ series, as wild boar bones are rare among Celtic archaeological sites, and the few that occur show no signs of butchery, having probably been used in sacrificial rituals. The boar also appears in Vedic mythology
Vedic mythology
. A story present in the Brāhmaṇas has Indra
Indra
slaying an avaricious boar, who has stolen the treasure of the asuras , then giving its carcass to Vishnu
Vishnu
, who offered it as a sacrifice to the gods. In the story's retelling in the Charaka Samhita , the boar is described as a form of Prajāpti , and is credited with having raised the earth from the primeval waters. In the Rāmāyaṇa and the Purāṇas , the same boar is portrayed as an avatar of Vishnu. Herakles
Herakles
brings Eurystheus
Eurystheus
the Erymanthian boar
Erymanthian boar
, as depicted on a black-figure amphora (c. 550 BC) from Vulci .

In Japanese culture , the boar is widely seen as a fearsome and reckless animal, to the point that several words and expressions in Japanese referring to recklessness include references to boars. The boar is the last animal of the oriental zodiac , with people born during the year of the Pig
Pig
being said to embody the boar-like traits of determination and impetuosity. Among Japanese hunters, the boar's courage and defiance is a source of admiration, and it is not uncommon for hunters and mountain people to name their sons after the animal _inoshishi_ (猪). Boars are also seen as symbols of fertility and prosperity; in some regions, it is thought that boars are drawn to fields owned by families including pregnant women, and hunters with pregnant wives are thought to have greater chances of success when boarhunting. The animal's link to prosperity was illustrated by its inclusion on the ¥ 10 note during the Meiji period , and it was once believed that a man could become wealthy by keeping a clump of boar hair in his wallet.

In the folklore of the Mongol Altai Uriankhai
Altai Uriankhai
tribe, the wild boar was associated with the watery underworld, as it was thought that the spirits of the dead entered the animal's head, to be ultimately transported to the water. Prior to the conversion to Islam, the Kyrgyz people
Kyrgyz people
believed that they were descended from boars, and thus did not eat pork. In Buryat mythology, the forefathers of the Buryats descended from heaven and were nourished by a boar. In China
China
, the boar is the emblem of the Miao people .

The boar (sanglier ) is frequently displayed in English , Scottish and Welsh heraldry . As with the lion , the boar is often shown as armed and langued. As with the bear , Scottish and Welsh heraldry displays the boar's head with the neck cropped, unlike the English version, which retains the neck. The white boar served as the badge of King Richard III of England , who distributed it among his northern retainers during his tenure as Duke of Gloucester
Duke of Gloucester
.

AS A GAME ANIMAL AND FOOD SOURCE

Main article: Boar hunting Wild boar
Wild boar
haunches and trophy , Umbria
Umbria
, Italy
Italy
. A wild boar dish served in Helsinki
Helsinki
, Finland .

Humans have been hunting boar for millennia, with the earliest artistic depictions of such activities dating back to the Upper Paleolithic . The animal was seen as a source of food among the Ancient Greeks , as well as a sporting challenge and source of epic narratives. The Romans inherited this tradition, with one of its first practitioners being Scipio Aemilianus . Boar hunting became particularly popular among the young nobility during the 3rd century BC as preparation for manhood and battle. A typical Roman boarhunting tactic involved surrounding a given area with large nets, then flushing the boar with dogs and immobilising it with smaller nets. The animal would then be dispatched with a _venabulum_, a short spear with a crossguard at the base of the blade. More than their Greek predecessors, the Romans extensively took inspiration from boarhunting in their art and sculpture. With the ascension of Constantine the Great , boarhunting took on Christian allegorical themes, with the animal being portrayed as a "black beast" analogous to the dragon of Saint George . Boarhunting continued after the fall of the Western Roman Empire , though the Germanic tribes considered the red deer to be a more noble and worthy quarry. The post-Roman nobility hunted boar as their predecessors did, but primarily as training for battle rather than sport. It was not uncommon for medieval hunters to deliberately hunt boars during the breeding season, when the animals were more aggressive. During the Renaissance
Renaissance
, when deforestation and the introduction of firearms reduced boar numbers, boarhunting became the sole prerogative of the nobility, one of many charges brought up against the rich during the German Peasants\' War and the French Revolution . During the mid-20th century, 7,000–8,000 boars were caught in the Caucasus, 6,000–7,000 in Kazakhstan, and about 5,000 in Central Asia
Central Asia
during the Soviet
Soviet
period, primarily through use of dogs and beats. In Nepal, farmers and poachers eliminate boars by baiting balls of wheat flour containing explosives with kerosene oil, with the animals' chewing motions triggering the devices.

Wild boar
Wild boar
can thrive in captivity, though piglets grow slowly and poorly without their mothers. Products derived from wild boar include meat, hide and bristles. _ Apicius _ devotes a whole chapter to the cooking of boar meat, providing ten recipes involving roasting, boiling and what sauces to use. The Romans usually served boar meat with garum . Boar\'s head was the centrepiece of most medieval Christmas
Christmas
celebrations among the nobility. Although growing in popularity as a captive-bred source of food, the wild boar takes longer to mature than most domestic pigs, and is usually smaller and produces less meat. Nevertheless, wild boar meat is leaner and healthier than pork , being of higher nutritional value and having a much higher concentration of essential amino acids . Most meat-dressing organisations agree that a boar carcass should yield 50 kg (110 lb) of meat on average. Large specimens can yield 15–20 kg (33–44 lb) of fat, with some giants yielding 30 kg (66 lb) or more. A boar hide can measure 300 dm2 , and can yield 350–1000 grams of bristle and 400 grams of underwool.

*

Roman relief of a dog confronting a boar, Cologne
Cologne
*

Southern Indian depiction of boar hunt, c. 1540 *

Pigsticking in British India
India
*

Boar shot in Volgograd Oblast , Russia
Russia
*

The Boar Hunt – Hans Wertinger, c. 1530, Danube Valley

CROP AND GARBAGE RAIDING

An adult sow and young that have broken open a litter bag in Berlin seeking food.

Boars can be damaging to agriculture . Populations living on the outskirts of towns or farms can dig up potatoes and damage melons , watermelons and maize . They generally only encroach upon farms when natural food is scarce. In the Belovezh forest for example, 34–47% of the local boar population will enter fields in years of moderate availability of natural foods. While the role of boars in damaging crops is often exaggerated, cases are known of boar depredations causing famines , as was the case in Hachinohe , Japan
Japan
in 1749, where 3,000 people died of what became known as the 'wild boar famine'. Still within Japanese culture, the boar's status as vermin is expressed through its title as "king of pests" and the popular saying (addressed to young men in rural areas) "When you get married, choose a place with no wild boar." In Central Europe, farmers typically repel boars through distraction or fright, while in Kazakhstan
Kazakhstan
it is usual to employ guard dogs in plantations. Although large boar populations can play an important role in limiting forest growth, they are also useful in keeping pest populations such as June bugs under control. The growth of urban areas and corresponding decline in natural boar habitats has led to some sounders entering human habitations in search of food. As in natural conditions, sounders in peri-urban areas are matriarchal, though males tend to be much less represented, and adults of both sexes can be up to 35% heavier than their forest-dwelling counterparts. As of 2010, at least 44 cities in 15 countries have experienced problems of some kind relating to the presence of habituated wild boar.

ATTACKS ON HUMANS

Depiction of a stylised boar attacking a man, Bhimbetaka , India

Actual attacks on humans are rare, but can be serious, resulting in multiple penetrating injuries to the lower part of the body. They generally occur during the boars' rutting season from November–January, in agricultural areas bordering forests or on paths leading through forests. The animal typically attacks by charging and pointing its tusks towards the intended victim, with most injuries occurring on the thigh region. Once the initial attack is over, the boar steps back, takes position and attacks again if the victim is still moving, only ending once the victim is completely incapacitated.

Boar attacks on humans have been documented since the Stone Age
Stone Age
, with one of the oldest depictions being a cave painting in Bhimbetaka , India
India
. The Romans and Ancient Greeks wrote of these attacks ( Odysseus
Odysseus
was wounded by a boar, and Adonis
Adonis
killed by one), and several attacks are shown on the headstones of England's 12th century Severn Temple graveyard. A 2012 study compiling recorded attacks from 1825–2012 found accounts of 665 human victims of both wild boars and feral pigs, with the majority (19%) of attacks in the animal's native range occurring in India. Most of the attacks occurred in rural areas during the winter months in non-hunting contexts, and were committed by solitary males.

SEE ALSO

* Mammals portal * Animals portal

* Domestic pig * Feral pig * Peccary * Indian pig

NOTES

* ^ It is from the male boar's solitary habits that the species gets its name in numerous Romance languages . Although the Latin
Latin
word for "boar" was _aper_, the French _sanglier_ and Italian _cinghiale_ derive from _singularis porcus_, which is Latin
Latin
for "solitary pig". * ^ Thirteen has been observed in a captive specimen.

REFERENCES

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Mammal
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Grassland". _Biological Invasions_. 8 (5): 1073–1089. doi :10.1007/s10530-005-6829-7 . * ^ Oldfield, Callie A.; Evans, Jonathan P. (2016-03-01). "Twelve years of repeated wild hog activity promotes population maintenance of an invasive clonal plant in a coastal dune ecosystem". _Ecology and Evolution_. 6 (8). * ^ Gupta, S. et al. A wild boar hunting: predation on a bonnet macaque by a wild boar in the Bandipur National Park, southern India. _Current Science_. Vol. 106, No. 9. (10 May 2014) * ^ _A_ _B_ Marsan Kamler, JF; Wang, SW; Lham, K; Stenkewitz, U; et al. (2011). "Seasonal diet of dholes (_Cuon alpinus_) in northwestern Bhutan". _Mamm Biol_. 76 (4): 518–520. doi :10.1016/j.mambio.2011.02.003 . CS1 maint: Explicit use of et al. (link ) * ^ Heptner, V. G.; Sludskii, A. A. (1992). _Mammals of the Soviet Union: Carnivora (hyaenas and cats), Volume 2_. Smithsonian Institution Libraries and National Science Foundation. p. 248–252. * ^ Taghdisi, M.; et al. (2013). "Diet and habitat use of the endangered Persian leopard (_Panthera pardus saxicolor_) in northeastern Iran" (PDF). _Turkish Journal of Zoologist_. 37: 554–561. doi :10.3906/zoo-1301-20 . CS1 maint: Explicit use of et al. (link ) * ^ Elmira Sharbafi, Mohammad S. Farhadinia, Hamid R. Rezaie, Alex Richard Braczkowski. (2016) Prey of the Persian Leopard
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* ^ " Forest of Dean
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BIBLIOGRAPHY

* Cabanau, Laurent (2001). _The Hunter's Library: Wild Boar in Europe_. Könemann. ISBN 3-8290-5528-5 . * Marsan, Andrea; Mattioli, Stefano (2013). _Il Cinghiale_ (in Italian). Il Piviere (collana Fauna selvatica. Biologia e gestione). ISBN 978-88-96348-178 . * Scheggi, Massimo (1999). _La bestia nera: Caccia al cinghiale fra mito, storia e attualità_ (in Italian). Editoriale Olimpia (collana Caccia). ISBN 88-253-7904-8 .

FURTHER READING

* Apollonio, M. et al. (1988), "The systematics of the wild boar (_Sus scrofa_ L.) in Italy", _Bolletino di Zoologia, 3:213–221_ * Carden, R.F. (2012) "Review of the Natural History of Wild Boar (Sus scrofa) on the island of Ireland", Report prepared by Ruth Carden for the Northern Ireland Environment Agency, Northern Ireland, UK, National Parks & Wildlife Service, Department of Arts, Heritage and the Gaeltacht, Dublin, Ireland and the National Museum of Ireland – Education & Outreach Department. * (in French) Durantel, P. (2007), _Le sanglier et ses chasses_, Editions Artemis, ISBN 2844166032 * Greene, J. (2011), _The Golden-Bristled Boar: Last Ferocious Beast of the Forest_, University of Virginia Press, ISBN 0-8139-3103-7 * (in French) Marillier, B. (2003), _Le sanglier héraldique_, Editions Cheminements, ISBN 2844781845 * Mayer, J. J. & Shedrow, C. B. (2007), _Annotated Bibliography of the Wild Pig
Pig
(Sus scrofa): Environmental Information Document, Washington Savannah River Company_ * (in Italian) Padiglione, V. (1989), _Il cinghiale cacciatore: Antropologia simbolica della caccia in Sardegna_, Armando Editore (collana Antropologia culturale) * Ronald M. Nowak (1999), _Walker’s Mammals of the World_ (6 ed.), Baltimore: Johns Hopkins University Press, ISBN 0-8018-5789-9 , LCCN 98023686

EXTERNAL LINKS

_ Wikisource has the text of the 1911 Encyclopædia Britannica_ article _BOAR _.

_ Wikispecies has information related to: SUS SCROFA_

_ Wikiquote has quotations related to: WILD BOAR _

* _ News related to Saskatchewan

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