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The wild boar (Sus scrofa), also known as the wild swine[3] Eurasian wild pig,[4] or simply wild pig,[5] is a suid native to much of Eurasia, North Africa, and the Greater Sunda Islands. Human intervention has spread its distribution further, making the species one of the widest-ranging mammals in the world, as well as the most widely spread suiform.[4] Its wide range, high numbers, and adaptability mean that it is classed as least concern by the IUCN[1] and it has become an invasive species in part of its introduced range. The animal probably originated in Southeast Asia during the Early Pleistocene,[6] and outcompeted other suid species as it spread throughout the Old World.[7] As of 1990, up to 16 subspecies are recognized, which are divided into four regional groupings based on skull height and lacrimal bone length.[2] The species lives in matriarchal societies consisting of interrelated females and their young (both male and female). Fully grown males are usually solitary outside the breeding season.[8] The grey wolf is the wild boar's main predator throughout most of its range except in the Far East and the Lesser Sunda Islands, where it is replaced by the tiger and Komodo dragon, respectively.[9][10] It has a long history of association with humans, having been the ancestor of most domestic pig breeds and a big-game animal for millennia.

Contents

1 Terminology

1.1 Hunting

2 Taxonomy and evolution

2.1 Subspecies 2.2 Domestication

3 Physical description 4 Social behavior and life cycle 5 Ecology

5.1 Habitat
Habitat
and sheltering behaviour 5.2 Diet 5.3 Predators

6 Range

6.1 Reconstructed range 6.2 Present range 6.3 Status in Britain 6.4 Introduction to North America

6.4.1 History

7 Diseases and parasites 8 Relationships with humans

8.1 In culture 8.2 As a game animal and food source 8.3 Crop and garbage raiding 8.4 Attacks on humans

9 See also 10 Notes 11 References 12 Bibliography 13 Further reading 14 External links

Terminology[edit] As true wild boars became extinct in Britain before the development of modern English, the same terms are often used for both true wild boar and pigs, especially large or semi-wild ones. The English 'boar' stems from the Old English
Old English
bar, which is thought to be derived from the West Germanic *bairaz, of unknown origin.[11] Boar is sometimes used specifically to refer to males, and may also be used to refer to male domesticated pigs, especially breeding males that have not been castrated. 'Sow', the traditional name for a female, again comes from Old English and Germanic; it stems from Proto-Indo-European, and is related to the Latin
Latin
sus and Greek hus and more closely to the modern German Sau. The young may be called 'piglets'. The animals' specific name scrofa is Latin
Latin
for 'sow'.[12] Hunting[edit] In hunting terminology, boars are given different designations according to their age:[13]

Designation Age Image

Squeaker 0–10 months

Juvenile 10–12 months

Pig
Pig
of the sounder Two years

Boar of the 4th/5th/6th year 3–5 years

Old boar Six years

Grand old boar Over seven years

"Solitary boar"

Taxonomy and evolution[edit]

Skull of Sus strozzii (Museo di Storia Naturale di Firenze), a Pleistocene
Pleistocene
suid that was outcompeted by S. scrofa

MtDNA
MtDNA
studies indicate that the wild boar originated from islands in Southeast Asia such as Indonesia
Indonesia
and the Philippines, and subsequently spread onto mainland Eurasia
Eurasia
and North Africa.[6] The earliest fossil finds of the species come from both Europe
Europe
and Asia, and date back to the Early Pleistocene.[14] By the late Villafranchian, S. scrofa largely displaced the related S. strozzii, a large, possibly swamp-adapted suid ancestral to the modern S. verrucosus throughout the Eurasian mainland, restricting it to insular Asia.[7] Its closest wild relative is the bearded pig of Malacca
Malacca
and surrounding islands.[3] Subspecies[edit] As of 2005[update],[2] 16 subspecies are recognised, which are divided into four regional groupings:

Western: Includes S. s. scrofa, S. s. meridionalis, S. s. algira, S. s. attila, S. s. lybicus, and S. s. nigripes. These subspecies are typically high-skulled (though lybicus and some scrofa are low-skulled), with thick underwool and (excepting scrofa and attila) poorly developed manes.[15] Indian: Includes S. s. davidi and S. s. cristatus. These subspecies have sparse or absent underwool, with long manes and prominent bands on the snout and mouth. While S. s. cristatus is high-skulled, S. s. davidi is low-skulled.[15] Eastern: Includes S. s. sibiricus, S. s. ussuricus, S. s. leucomystax, S. s. riukiuanus, S. s. taivanus, and S. s. moupinensis. These subspecies are characterised by a whitish streak extending from the corners of the mouth to the lower jaw. With the exception of S. s. ussuricus, most are high-skulled. The underwool is thick, except in S. s. moupinensis, and the mane is largely absent.[15] Indonesian: Represented solely by S. s. vittatus, it is characterised by its sparse body hair, lack of underwool, fairly long mane, a broad reddish band extending from the muzzle to the sides of the neck.[15] It is the most basal of the four groups, having the smallest relative brain size, more primitive dentition and unspecialised cranial structure.[16]

Subspecies Image Trinomial authority Description Range Synonyms

Central European boar S. s. scrofa Nominate subspecies

Linnaeus, 1758 A medium-sized, dark to rusty-brown haired subspecies with long and relatively narrow lacrimal bones[3] Much of continental Europe
Europe
and into Eurasia anglicus (Reichenbach, 1846), aper (Erxleben, 1777), asiaticus (Sanson, 1878), bavaricus (Reichenbach, 1846), campanogallicus (Reichenbach, 1846), capensis (Reichenbach, 1846), castilianus (Thomas, 1911), celticus (Sanson, 1878), chinensis (Linnaeus, 1758), crispus (Fitzinger, 1858), deliciosus (Reichenbach, 1846), domesticus (Erxleben, 1777), europaeus (Pallas, 1811), fasciatus (von Schreber, 1790), ferox (Moore, 1870), ferus (Gmelin, 1788), gambianus (Gray, 1847), hispidus (von Schreber, 1790), hungaricus (Reichenbach, 1846), ibericus (Sanson, 1878), italicus (Reichenbach, 1846), juticus (Fitzinger, 1858), lusitanicus (Reichenbach, 1846), macrotis (Fitzinger, 1858), monungulus (G. Fischer [von Waldheim], 1814), moravicus (Reichenbach, 1846), nanus (Nehring, 1884), palustris (Rütimeyer, 1862), pliciceps (Gray, 1862), polonicus (Reichenbach, 1846), sardous (Reichenbach, 1846), scropha (Gray, 1827), sennaarensis (Fitzinger, 1858), sennaarensis (Gray, 1868), sennaariensis (Fitzinger, 1860), setosus (Boddaert, 1785), siamensis (von Schreber, 1790), sinensis (Erxleben, 1777), suevicus (Reichenbach, 1846), syrmiensis (Reichenbach, 1846), turcicus (Reichenbach, 1846), variegatus (Reichenbach, 1846), vulgaris (S. D. W., 1836), wittei (Reichenbach, 1846)

North African boar S. s. algira

Loche, 1867 Sometimes considered a junior synonym of S. s. scrofa, but smaller and with proportionally longer tusks[17] Tunisia, Algeria
Algeria
and Morocco barbarus (Sclater, 1860) sahariensis (Heim de Balzac, 1937)

Carpathian boar S. s. attila

Thomas, 1912 A large-sized subspecies with long lacrimal bones and dark hair, though lighter-coloured than S. s. scrofa[3] Romania, Hungary, Ukraine, Balkans, Caucasus, Transcaucasia, Caspian coast, Asia Minor
Asia Minor
and northern Iran falzfeini (Matschie, 1918)

Indian boar S. s. cristatus

Wagner, 1839 A long-maned subspecies with a coat that is brindled black unlike S. s. davidi,[18] it is more lightly built than S. s. scrofa. Its head is larger and more pointed than that of S. s. scrofa, and its ears smaller and more pointed. The plane of the forehead is straight, while it is concave in S. s. scrofa.[19] India, Nepal, Burma, western Thailand
Thailand
and Sri Lanka affinis (Gray, 1847), aipomus (Gray, 1868), aipomus (Hodgson, 1842), bengalensis (Blyth, 1860), indicus (Gray, 1843), isonotus (Gray, 1868), isonotus (Hodgson, 1842), jubatus (Miller, 1906), typicus (Lydekker, 1900), zeylonensis (Blyth, 1851)

Central Asian boar S. s. davidi

Groves, 1981 A small, long-maned and light brown subspecies[18] Pakistan
Pakistan
and northwest India
India
to southeastern Iran

Japanese boar S. s. leucomystax

Temminck, 1842 A small, almost maneless, yellowish-brown subspecies[18] All of Japan, save for Hokkaido
Hokkaido
and the Ryukyu Islands japonica (Nehring, 1885) nipponicus (Heude, 1899)

Anatolian boar S. s. libycus

Gray, 1868 A small, pale and almost maneless subspecies[18] Transcaucasia, Turkey, Levant, and former Yugoslavia lybicus (Groves, 1981) mediterraneus (Ulmansky, 1911) reiseri (Bolkay, 1925)

Maremman boar S. s. majori

De Beaux and Festa, 1927 Smaller than S. s. scrofa, with a higher and wider skull, since the 1950s, it has crossed extensively with S. s. scrofa, largely due to the two being kept together in meat farms and artificial introductions by hunters of S. s. scrofa specimens into S. s. majori habitats.[20] Its separation from S. s. scrofa is doubtful.[21] Maremma
Maremma
(central Italy)

Mediterranean boar S. s. meridionalis

Forsyth Major, 1882

Andalusia, Corsica
Corsica
and Sardinia baeticus (Thomas, 1912) sardous (Ströbel, 1882)

Northern Chinese boar S. s. moupinensis

Milne-Edwards, 1871 There are significant variations within this subspecies, and it is possible there are actually several subspecies involved.[18] Coastal China
China
south to Vietnam
Vietnam
and west to Sichuan acrocranius (Heude, 1892), chirodontus (Heude, 1888), chirodonticus (Heude, 1899), collinus (Heude, 1892), curtidens (Heude, 1892), dicrurus (Heude, 1888), flavescens (Heude, 1899), frontosus (Heude, 1892), laticeps (Heude, 1892), leucorhinus (Heude, 1888), melas (Heude, 1892), microdontus (Heude, 1892), oxyodontus (Heude, 1888), paludosus (Heude, 1892), palustris (Heude, 1888), planiceps (Heude, 1892), scrofoides (Heude, 1892), spatharius (Heude, 1892), taininensis (Heude, 1888)

Middle Asian boar S. s. nigripes

Blanford, 1875 A light coloured subspecies with black legs which, though varied in size, it is generally quite large, the lacrimal bones and facial region of the skull are shorter than those of S. s. scrofa and S. s. attila.[3] Middle Asia, Kazakhstan, eastern Tien Shan, western Mongolia, Kashgar and possibly Afghanistan
Afghanistan
and southern Iran

Ryukyu boar S. s. riukiuanus

Kuroda, 1924 A small subspecies[18] Ryukyu Islands

Trans-baikal boar S. s. sibiricus

Staffe, 1922 The smallest subspecies of the former Soviet
Soviet
region, it has dark brown, almost black hair and a light grey patch extending from the cheeks to the ears. The skull is squarish and the lacrimal bones short.[3] Baikal, Transbaikalia, northern and northeastern Mongolia raddeanus (Adlerberg, 1930)

Formosan boar S. s. taivanus

Swinhoe, 1863 A small blackish subspecies[18] Taiwan

Ussuri boar S. s. ussuricus

Heude, 1888 The largest subspecies, it has usually dark hair and a white band extending from the corners of the mouth to the ears. The lacrimal bones are shortened, but longer than those of S. s. sibiricus.[3] Eastern China, Ussuri and Amur bay canescens (Heude, 1888), continentalis (Nehring, 1889), coreanus (Heude, 1897), gigas (Heude, 1892), mandchuricus (Heude, 1897), songaricus (Heude, 1897)

Banded pig S. s. vittatus

Boie, 1828 A small, short-faced and sparsely furred subspecies with a white band on the muzzle; it might be a separate species, and shows some similarities with some other suid species in Southeast Asia.[18] From Peninsular Malaysia, and in Indonesia
Indonesia
from Sumatra
Sumatra
and Java
Java
east to Komodo andersoni (Thomas and Wroughton, 1909), jubatulus (Miller, 1906), milleri (Jentink, 1905), pallidiloris (Mees, 1957), peninsularis (Miller, 1906), rhionis (Miller, 1906), typicus (Heude, 1899)

Wild boar
Wild boar
(left) and domestic pig (right) skulls: Note the greatly shortened facial region of the latter.[22]

Domestication[edit]

Male, domestic pig–wild boar cross

With the exception of domestic pigs in Timor
Timor
and Papua New Guinea (which appear to be of Sulawesi warty pig stock), the wild boar is the ancestor of most pig breeds.[16][23] Archaeological evidence suggests that pigs were domesticated from wild boar as early as 13,000–12,700 BC in the Near East
Near East
in the Tigris
Tigris
Basin[24] being managed in the wild in a way similar to the way they are managed by some modern New Guineans.[25] Remains of pigs have been dated to earlier than 11,400 BC in Cyprus. Those animals must have been introduced from the mainland, which suggests domestication in the adjacent mainland by then.[26] There was also a separate domestication in China
China
which took place about 8000 years ago.[27][28] DNA
DNA
evidence from sub-fossil remains of teeth and jawbones of Neolithic
Neolithic
pigs shows that the first domestic pigs in Europe
Europe
had been brought from the Near East. This stimulated the domestication of local European wild boar resulting in a third domestication event with the Near Eastern genes dying out in European pig stock. Modern domesticated pigs have involved complex exchanges, with European domesticated lines being exported in turn to the ancient Near East.[29][30] Historical records indicate that Asian pigs were introduced into Europe
Europe
during the 18th and early 19th centuries.[27] Domestic pigs tend to have much more developed hindquarters than their wild boar ancestors, to the point where 70% of their body weight is concentrated in the posterior, which is the opposite of wild boar, where most of the muscles are concentrated on the head and shoulders.[31]

Physical description[edit]

Dentition, as illustrated by Charles Knight

The wild boar is a bulky, massively built suid with short and relatively thin legs. The trunk is short and massive, while the hindquarters are comparatively underdeveloped. The region behind the shoulder blades rises into a hump, and the neck is short and thick, to the point of being nearly immobile. The animal's head is very large, taking up to one third of the body's entire length.[3] The structure of the head is well suited for digging. The head acts as a plow, while the powerful neck muscles allow the animal to upturn considerable amounts of soil:[32] it is capable of digging 8–10 cm (3.1–3.9 in) into frozen ground and can upturn rocks weighing 40–50 kg (88–110 lb).[9] The eyes are small and deep-set, and the ears long and broad. The species has well developed canine teeth, which protrude from the mouths of adult males. The middle hooves are larger and more elongated than the lateral ones, and are capable of quick movements.[3] The animal can run at a maximum speed of 40 km/h and jump at a height of 140–150 cm (55–59 in).[9] Sexual dimorphism
Sexual dimorphism
is very pronounced in the species, with males being typically 5–10% larger and 20–30% heavier than females. Males also sport a mane running down the back, which is particularly apparent during autumn and winter.[33] The canine teeth are also much more prominent in males, and grow throughout life. The upper canines are relatively short and grow sideways early in life, though gradually curve upwards. The lower canines are much sharper and longer, with the exposed parts measuring 10–12 cm (3.9–4.7 in) in length. In the breeding period, males develop a coating of subcutaneous tissue, which may be 2–3 cm (0.79–1.18 in) thick, extending from the shoulder blades to the rump, thus protecting vital organs during fights. Males sport a roughly egg-sized sack near the opening of the penis, which collects urine and emits a sharp odour. The purpose of this is not fully understood.[3]

Skeleton, as illustrated by Richard Lydekker.

A European wild boar piglet, painted by Hans Hoffman in 1578. Note the stripes, a characteristic feature of piglets.

Adult size and weight is largely determined by environmental factors; boars living in arid areas with little productivity tend to attain smaller sizes than their counterparts inhabiting areas with abundant food and water. In most of Europe, males average 75–100 kg (165–220 lb) in weight, 75–80 cm (30–31 in) in shoulder height and 150 cm (59 in) in body length, whereas females average 60–80 kg (130–180 lb) in weight, 70 cm (28 in) in shoulder height and 140 cm (55 in) in body length. In Europe's Mediterranean regions, males may reach average weights as low as 50 kg (110 lb) and females 45 kg (99 lb), with shoulder heights of 63–65 cm (25–26 in). In the more productive areas of Eastern Europe, males average 110–130 kg (240–290 lb) in weight, 95 cm (37 in) in shoulder height and 160 cm (63 in) in body length, while females weigh 95 kg (209 lb), reach 85–90 cm (33–35 in) in shoulder height and 145 cm (57 in) in body length. In Western and Central Europe, the largest males weigh 200 kg (440 lb) and females 120 kg (260 lb). In North-Eastern Asia, large males can reach brown bear-like sizes, weighing 270 kg (600 lb) and measuring 110–118 cm (43–46 in) in shoulder height. Some adult males in Ussuriland and Manchuria
Manchuria
have been recorded to weigh 300–350 kg (660–770 lb) and measure 125 cm (49 in) in shoulder height. Adults of this size are generally immune from wolf predation.[34] Such giants are rare in modern times, due to past overhunting preventing animals from attaining their full growth.[3] The winter coat consists of long, coarse bristles underlaid with short brown downy fur. The length of these bristles varies along the body, with the shortest being around the face and limbs and the longest running along the back. These back bristles form the aforementioned mane prominent in males and stand erect when the animal is agitated. Colour is highly variable; specimens around Lake Balkhash
Lake Balkhash
are very lightly colored, and can even be white, while some boars from Belarus and Ussuriland can be black. Some subspecies sport a light colored patch running backward from the corners of the mouth. Coat color also varies with age, with piglets having light brown or rusty-brown fur with pale bands extending from the flanks and back.[3] The wild boar produces a number of different sounds which are divided into three categories:

Contact calls: Grunting noises which differ in intensity according to the situation.[35] Adult males are usually silent, while females frequently grunt and piglets whine.[3] When feeding, boars express their contentment through purring. Studies have shown that piglets imitate the sounds of their mother, thus different litters may have unique vocalisations.[35] Alarm calls: Warning cries emitted in response to threats.[35] When frightened, boars make loud huffing ukh! ukh! sounds or emit screeches transcribed as gu-gu-gu.[3] Combat calls: High-pitched, piercing cries.[35]

Its sense of smell is very well developed, to the point that the animal is used for drug detection in Germany.[36] Its hearing is also acute, though its eyesight is comparatively weak,[3] lacking colour vision[36] and being unable to recognise a standing human 10–15 metres away.[9]

Pigs are one of four known mammalian species which possess mutations in the nicotinic acetylcholine receptor that protect against snake venom. Mongooses, honey badgers, hedgehogs, and pigs all have modifications to the receptor pocket which prevents the snake venom α-neurotoxin from binding. These represent four separate, independent mutations.[37] Social behavior and life cycle[edit] Boars are typically social animals, living in female-dominated sounders consisting of barren sows and mothers with young led by an old matriarch. Male boars leave their sounder at the age of 8–15 months, while females either remain with their mothers or establish new territories nearby. Subadult males may live in loosely knit groups, while adult and elderly males tend to be solitary outside the breeding season.[8][a]

Central European wild boar (S. s. scrofa) piglets suckling

The breeding period in most areas lasts from November to January, though most mating only lasts a month and a half. Prior to mating, the males develop their subcutaneous armor, in preparation for confronting rivals. The testicles double in size and the glands secrete a foamy yellowish liquid. Once ready to reproduce, males travel long distances in search of a sounder of sows, eating little on the way. Once a sounder has been located, the male drives off all young animals and persistently chases the sows. At this point, the male fiercely fights potential rivals,[3] A single male can mate with 5–10 sows.[9] By the end of the rut, males are often badly mauled and have lost 20% of their body weight,[3] with bite-induced injuries to the penis being common.[39] The gestation period varies according to the age of the expecting mother. For first time breeders, it lasts 114–130 days, while it lasts 133–140 days in older sows. Farrowing occurs between March and May, with litter sizes depending on the age and nutrition of the mother. The average litter consists of 4–6 piglets, with the maximum being 10–12.[3][b] The piglets are whelped in a nest constructed from twigs, grasses and leaves. Should the mother die prematurely, the piglets are adopted by the other sows in the sounder.[41] Newborn piglets weigh around 600–1,000 grams, lacking underfur and bearing a single milk incisor and canine on each half of the jaw.[3] There is intense competition between the piglets over the most milk-rich nipples, as the best fed young grow faster and have stronger constitutions.[41] The piglets do not leave the lair for their first week of life. Should the mother be absent, the piglets lie closely pressed to each other. By two weeks of age, the piglets begin accompanying their mother on her journeys. Should danger be detected, the piglets take cover or stand immobile, relying on their camouflage to keep them hidden. The neonatal coat fades after three months, with adult coloration being attained at eight months. Although the lactation period lasts 2.5–3.5 months, the piglets begin displaying adult feeding behaviors at the age of 2–3 weeks. The permanent dentition is fully formed by 1–2 years. With the exception of the canines in males, the teeth stop growing during the middle of the fourth year. The canines in old males continue to grow throughout their lives, curving strongly as they age. Sows attain sexual maturity at the age of one year, with males attaining it a year later. However, estrus usually first occurs after two years in sows, while males begin participating in the rut after 4–5 years, as they are not permitted to mate by the older males.[3] The maximum lifespan in the wild is 10–14 years, though few specimens survive past 4–5 years.[42] Boars in captivity have lived for 20 years.[9]

Ecology[edit] Habitat
Habitat
and sheltering behaviour[edit]

An individual from higher ridges of Himalayas
Himalayas
at 9,600 ft in Pangolakha Wildlife Sanctuary, Sikkim, India.

Wild boar
Wild boar
frequently wallow in mud, possibly to regulate temperature or remove parasites

The wild boar inhabits a diverse array of habitats from boreal taigas to deserts.[3] In mountainous regions, it can even occupy alpine zones, occurring up to 1,900 meters in the Carpathians, 2,600 meters in the Caucasus
Caucasus
and up to 3,600–4,000 meters in the mountains in Central Asia
Central Asia
and Kazakhstan.[3] In order to survive in a given area, wild boars require a habitat fulfilling three conditions: heavily brushed areas providing shelter from predators, water for drinking and bathing purposes and an absence of regular snowfall.[43] The main habitats favored by boars in Europe
Europe
are deciduous and mixed forests, with the most favorable areas consisting of forest composed of oak and beech enclosing marshes and meadows. In the Białowieża Forest, the animal's primary habitat consists of well developed, broad-leaved and mixed forests, along with marshy mixed forests, with coniferous forests and undergrowths being of secondary importance. Forests made up entirely of oak groves and beeches are used only during the fruit-bearing season. This is in contrast to the Caucasian and Transcaucasian mountain areas, where boars will occupy such fruit-bearing forests year-round. In the mountainous areas of the Russian Far East, the species inhabits nutpine groves, hilly mixed forests where Mongolian oak
Mongolian oak
and Korean pine
Korean pine
are present, swampy mixed taiga and coastal oak forests. In Transbaikalia, boars are restricted to river valleys with nut pine and shrubs. Boars are regularly encountered in pistachio groves in winter in some areas of Tajikistan and Turkmenia, while in spring they migrate to open deserts; boar have also colonized deserts in several areas they have been introduced to.[3][43][44] On the islands of Komodo and Rinca, the boar mostly inhabits savanna or open monsoon forests, avoiding heavily forested areas unless pursued by humans.[10] Wild boar
Wild boar
are known to be competent swimmers, capable of covering long distances. In 2013, one boar was reported to have completed the seven mile swim from France
France
to Alderney
Alderney
in the Channel Islands. Due to concerns about disease it was shot and incinerated.[45] Wild boar
Wild boar
rest in shelters, which contain insulating material like spruce branches and dry hay. These resting places are occupied by whole families (though males lie separately), and are often located in the vicinity of streams, in swamp forests, in tall grass or shrub thickets. Boars never defecate in their shelters, and will cover themselves with soil and pine needles when irritated by insects.[9] Diet[edit]

Male Indian boar
Indian boar
(S. s. cristatus) feeding on a chital carcass

The wild boar is a highly versatile omnivore, whose diversity in choice of food rivals that of humans.[32] Their foods can be divided into four categories:

Rhizomes, roots, tubers and bulbs, all of which are dug up throughout the year in the animal's whole range.[3] Nuts, berries, and seeds, which are consumed when ripened and are dug up from the snow when abundant.[3] Leaves, bark, twigs, and shoots, along with garbage.[3] Earthworms, insects, mollusks, fish, rodents, insectivores, bird eggs, lizards, snakes, frogs, and carrion. Most of these prey items are taken in warm periods.[3]

A 50 kg (110 lb) boar needs around 4,000–4,500 calories of food per day, though this required amount increases during winter and pregnancy,[32] with the majority of its diet consisting of food items dug from the ground like underground plant material and burrowing animals.[3] Acorns and beechnuts are invariably its most important food items in temperate zones,[citation needed] as they are rich in the carbohydrates necessary for the buildup of fat reserves needed to survive lean periods.[32] In Western Europe, underground plant material favoured by boars includes bracken, willow herb, bulbs, meadow herb roots and bulbs, and the bulbs of cultivated crops. Such food is favoured in early spring and summer, but may also be eaten in autumn and winter during beechnut and acorn crop failures. Should regular wild foods become scarce, boars will eat tree bark and fungi, as well as visit cultivated potato and artichoke fields.[3] Boar soil disturbance and foraging have been shown to facilitate invasive plants.[46][47] Boars of the vittatus subspecies in Ujung Kulon National Park in Java
Java
differ from most other populations by their primarily frugivorous diet, which consists of 50 different fruit species, especially figs, thus making them important seed dispersers.[4] The wild boar can consume numerous genera of poisonous plants without ill effect, including Aconitum, Anemone, Calla, Caltha, Ferula, and Pteridium.[9] Boars may occasionally prey on small vertebrates like newborn deer fawns, leporids and galliform chicks.[32] Boars inhabiting the Volga Delta and near some lakes and rivers of Kazakhstan
Kazakhstan
have been recorded to feed extensively on fish like carp and Caspian roach. Boars in the former area will also feed on cormorant and heron chicks, bivalved molluscs, trapped muskrats and mice.[3] There is at least one record of a boar killing and eating a bonnet macaque in southern India's Bandipur National Park, though this may have been a case of intraguild predation, brought on by interspecific competition for human handouts.[48]

Predators[edit]

Tigers killing a wild boar in Kanha Tiger
Tiger
Reserve

Piglets are vulnerable to attack from medium-sized felids like lynx, jungle cats and snow leopards and other carnivorans like brown bears and yellow-throated martens.[3] The grey wolf is the main predator of wild boar throughout most of its range. A single wolf can kill around 50 to 80 boars of differing ages in one year.[3] In Italy[49] and Belarus' Belovezhskaya Pushcha National Park, boars are the wolf's primary prey, despite an abundance of alternative, less powerful ungulates.[49] Wolves are particularly threatening during the winter, when deep snow impedes the boars' movements. In the Baltic regions, heavy snowfall can allow wolves to eliminate boars from an area almost completely. Wolves primarily target piglets and subadults, and only rarely attack adult sows. Adult males are usually avoided entirely.[3] Dholes may also prey on boars, to the point of keeping their numbers down in northwestern Bhutan, despite there being many more cattle in the area.[50]

Banded pig
Banded pig
(S. s. vittatus) eaten by Komodo dragons

Leopards are predators of wild boar in the Caucasus, Transcaucasia, the Russian Far East, India, China,[51] and Iran. In most areas, boars constitute only a small part of the leopard's diet. However, in Iran's Sarigol National Park, boars are the second most frequently targeted prey species after mouflon, though adult individuals are generally avoided, as they are above the leopard's preferred weight range of 10–40 kg (22–88 lb).[52] This dependence on wild boar is largely due in part to the local leopard subspecies' large size.[53] Boars of all ages were once the primary prey of tigers in Transcaucasia, Kazakhstan, Middle Asia
Middle Asia
and the Far East up until the late 19th century. In modern times, tiger numbers are too low to have a limiting effect on boar populations. A single tiger can systematically destroy an entire sounder by preying on its members one by one, before moving on to another herd. Tigers have been noted to chase boars for longer distances than with other prey. In two rare cases, boars were reported to gore a small tiger and a tigress to death in self-defense.[54] In the Amur region, wild boars are one of the two most important prey species for tigers alongside the Manchurian wapiti, with the two species collectively comprising roughly 80% of the felid's prey.[55] In Sikhote Alin, a tiger can kill 30–34 boars a year.[9] Studies of tigers in India
India
indicate that boars are usually secondary in preference to various cervids and bovids,[56] though when boars are targeted, healthy adults are caught more frequently than young and sick specimens.[57] On the islands of Komodo, Rinca, and Flores, the boar's main predator is the Komodo dragon.[10]

Range[edit] Reconstructed range[edit] The species originally occurred in North Africa
North Africa
and much of Eurasia; from the British Isles
British Isles
to Korea
Korea
and the Sunda Islands. The northern limit of its range extended from southern Scandinavia
Scandinavia
to southern Siberia
Siberia
and Japan. Within this range, it was only absent in extremely dry deserts and alpine zones. It was once found in North Africa
North Africa
along the Nile
Nile
valley up to Khartum
Khartum
and north of the Sahara. The species occurs on a few Ionian and Aegean Islands, sometimes swimming between islands.[58] The reconstructed northern boundary of the animal's Asian range ran from Lake Ladoga
Lake Ladoga
(at 60°N) through the area of Novgorod
Novgorod
and Moscow into the southern Urals, where it reached 52°N. From there, the boundary passed Ishim and farther east the Irtysh
Irtysh
at 56°N. In the eastern Baraba steppe
Baraba steppe
(near Novosibirsk) the boundary turned steep south, encircled the Altai Mountains, and went again eastward including the Tannu-Ola Mountains
Tannu-Ola Mountains
and Lake Baikal. From here the boundary went slightly north of the Amur River
Amur River
eastward to its lower reaches at the Sea of Okhotsk. On Sakhalin, there are only fossil reports of wild boar. The southern boundaries in Europe
Europe
and Asia were almost invariably identical to the seashores of these continents. It is absent in the dry regions of Mongolia
Mongolia
from 44–46°N southward, in China
China
westward of Sichuan
Sichuan
and in India
India
north of the Himalayas. It is absent in the higher elevations of Pamir and Tien Shan, though they do occur in the Tarim basin
Tarim basin
and on the lower slopes of the Tien Shan.[3] Present range[edit] In recent centuries, the range of wild boar has changed dramatically, largely due to hunting by humans and more recently because of captive wild boar escaping into the wild. Prior to the 20th century, boar populations had declined in numerous areas, with British populations probably becoming extinct during the 13th century.[59] In Denmark, the last boar was shot at the beginning of the 19th century, and in 1900 they were absent in Tunisia
Tunisia
and Sudan and large areas of Germany, Austria, and Italy. In Russia, they were extirpated in wide areas in the 1930s.[3] The last boar in Egypt
Egypt
reportedly died on 20 December 1912 in the Giza Zoo, with wild populations having disappeared around 1894–1902. Prince Kamal el Dine Hussein
Prince Kamal el Dine Hussein
attempted to repopulate Wadi El Natrun with boars of Hungarian stock, but they were quickly exterminated by poachers.[60] A revival of boar populations began in the middle of the 20th century. By 1950 wild boar had once again reached their original northern boundary in many parts of their Asiatic range. By 1960, they reached Leningrad
Leningrad
and Moscow, and by 1975 they were to be found in Archangelsk and Astrakhan. In the 1970s they again occurred in Denmark and Sweden, where captive animals escaped and now survive in the wild. In England, wild boar populations re-established themselves in the 1990s, after escaping from specialist farms that had imported European stock.[59] Status in Britain[edit]

Mixed sounder of wild boar and domestic pigs at Culzie, Scotland

Wild boars were apparently already becoming rare by the 11th century since a 1087 forestry law enacted by William the Conqueror
William the Conqueror
punishes through blinding the unlawful killing of a boar. Charles I attempted to reintroduce the species into the New Forest, though this population was exterminated during the Civil War. Between their medieval extinction and the 1980s, when wild boar farming began, only a handful of captive wild boar, imported from the continent, were present in Britain. Occasional escapes of wild boar from wildlife parks have occurred as early as the 1970s, but since the early 1990s significant populations have re-established themselves after escapes from farms, the number of which has increased as the demand for meat from the species has grown. A 1998 MAFF (now DEFRA) study on wild boar living wild in Britain confirmed the presence of two populations of wild boar living in Britain; one in Kent/East Sussex and another in Dorset.[59] Another DEFRA report, in February 2008,[61] confirmed the existence of these two sites as 'established breeding areas' and identified a third in Gloucestershire/Herefordshire; in the Forest of Dean/ Ross on Wye
Ross on Wye
area. A 'new breeding population' was also identified in Devon. There is another significant population in Dumfries
Dumfries
and Galloway. Populations estimates were as follows:

The largest population, in Kent/East Sussex, was then estimated at approximately 200 animals in the core distribution area. The second largest, in Gloucestershire/Herefordshire, was first estimated to be in excess of 100 animals. Legally classified as dangerous wild animals, the group is known to be feral descendants of domestic (Tamworth) pigs abandoned nearby. Their numbers grew by 2016 to at least 1500 and the Forestry Commission
Forestry Commission
planned to reduce the total to a manageable 400. "Adult males can reach twenty stone (125 kg), run at thirty miles an hour, and can jump or barge through all but the strongest fences. Also, they are not afraid of humans, so (unlike deer) you can't just shoo them out of your garden."[62] The smallest, in west Dorset, was estimated to be fewer than 50 animals. Since winter 2005/6 significant escapes/releases have also resulted in animals colonizing areas around the fringes of Dartmoor, in Devon. These are considered as an additional single 'new breeding population' and currently estimated to be up to 100 animals.

Population estimates for the Forest of Dean
Forest of Dean
are disputed as at the time that the DEFRA population estimate was 100, a photo of a boar sounder in the forest near Staunton with over 33 animals visible was published, and at about the same time over 30 boar were seen in a field near the original escape location of Weston under Penyard many miles away. In early 2010 the Forestry Commission
Forestry Commission
embarked on a cull,[63] with the aim of reducing the boar population from an estimated 150 animals to 100. By August it was stated that efforts were being made to reduce the population from 200 to 90, but that only 25 had been killed.[64] The failure to meet cull targets was confirmed in February 2011.[65] Wild boars have crossed the River Wye
River Wye
into Monmouthshire, Wales. Iolo Williams, the BBC Wales wildlife expert, attempted to film Welsh boar in late 2012.[66] Many other sightings, across the UK, have also been reported.[67] The effects of wild boar on the UK's woodlands were discussed with Ralph Harmer of the Forestry Commission
Forestry Commission
on the BBC Radio's Farming Today radio programme in 2011. The programme prompted activist writer George Monbiot
George Monbiot
to propose a thorough population study, followed by the introduction of permit-controlled culling.[68] Introduction to North America[edit]

"Razorbacks" confronting an alligator in Florida

Wild boars are an invasive species in the Americas, and cause problems including outcompeting native species for food, destroying the nests of ground-nesting species, killing fawns and young domestic livestock, destroying agricultural crops, eating tree seeds and seedlings, destroying native vegetation and wetlands through wallowing, damaging water quality, coming into violent conflict with humans and pets, and carrying pig and human diseases including brucellosis, trichinosis, and pseudorabies. In some jurisdictions, it is illegal to import, breed, release, possess, sell, distribute, trade, transport, hunt, or trap Eurasian boars. Hunting and trapping is done systematically, to increase the chance of eradication and to remove the incentive to illegally release boars, which have mostly been spread deliberately by sport hunters.[69] History[edit] While domestic pigs, both captive and feral (popularly termed "razorbacks"), have been in North America since the earliest days of European colonization, pure wild boars were not introduced into the New World
New World
until the 19th century. The suids were released into the wild by wealthy landowners as big game animals. The initial introductions took place in fenced enclosures, though several escapes occurred, with the escapees sometimes intermixing with already established feral pig populations. The first of these introductions occurred in New Hampshire
New Hampshire
in 1890. Thirteen wild boars from Germany were purchased by Austin Corbin from Carl Hagenbeck, and released into a 9,500-hectare game preserve in Sullivan County. Several of these boars escaped, though they were quickly hunted down by locals. Two further introductions were made from the original stocking, with several escapes taking place due to breaches in the game preserve's fencing. These escapees have ranged widely, with some specimens having been observed crossing into Vermont.[70] In 1902, 15–20 wild boar from Germany were released into a 3,200-hectare estate in Hamilton County, New York. Several specimens escaped six years later, dispersing into the William C. Whitney Wilderness Area, with their descendants surviving for at least 20 years.[70] The most extensive boar introduction in the US took place in western North Carolina
North Carolina
in 1912, when 13 boars of undetermined European origin were released into two fenced enclosures in a game preserve in Hooper Bald, Graham County. Most of the specimens remained in the preserve for the next decade, until a large-scale hunt caused the remaining animals to break through their confines and escape. Some of the boars migrated to Tennessee, where they intermixed with both free-ranging and feral pigs in the area. In 1924, a dozen Hooper Bald
Hooper Bald
wild pigs were shipped to California
California
and released in a property between Carmel Valley and the Los Padres National Forest. These hybrid boar were later used as breeding stock on various private and public lands throughout the state, as well as in other states like Florida, Georgia, South Carolina, West Virginia
West Virginia
and Mississippi.[70] Several wild boars from Leon Springs and the San Antonio, Saint Louis and San Diego Zoos were released in the Powder Horn Ranch in Calhoun County, Texas, in 1939. These specimens escaped and established themselves in surrounding ranchlands and coastal areas, with some crossing the Espiritu Santo Bay and colonizing Matagorda Island. Descendants of the Powder Horn Ranch boars were later released onto San José Island and the coast of Chalmette, Louisiana.[70] Wild boar
Wild boar
of unknown origin were stocked in a ranch in the Edwards Plateau in the 1940s, only to escape during a storm and hybridize with local feral pig populations, later spreading into neighboring counties.[70] Starting in the mid-80s, several boars purchased from the San Diego Zoo and Tierpark Berlin
Tierpark Berlin
were released into the United States. A decade later, more specimens from farms in Canada
Canada
and Białowieża Forest
Białowieża Forest
was let loose. In recent years, wild pig populations have been reported in 44 states within the US, most of which are likely wild boar–feral hog hybrids. Pure wild boar populations may still be present, but are extremely localized.[70] Diseases and parasites[edit]

Lesions consistent with bovine tuberculosis on the lower jaw and lung of a wild boar

Wild boars are known to host at least 20 different parasitic worm species, with maximum infections occurring in summer. Young animals are vulnerable to helminths like Metastrongylus, which are consumed by boars through earthworms, and cause death by parasitising the lungs. Wild boar
Wild boar
also carry parasites known to infect humans, including Gastrodiscoides, Trichinella spiralis, Taenia solium, and Balantidium coli. Wild boar
Wild boar
in southern regions are frequently infested with ticks (Dermacentor, Rhipicephalus, and Hyalomma) and hog lice. The species also suffers from blood-sucking flies, which it escapes by bathing frequently or hiding in dense shrubs.[3] Swine plague spreads very quickly in wild boar, with epizootics being recorded in Germany, Poland, Hungary, Belarus, the Caucasus, the Far East, Kazakhstan, and other regions. Foot-and-mouth disease
Foot-and-mouth disease
can also take on epidemic proportions in boar populations. The species occasionally, but rarely contracts Pasteurellosis, hemorrhagic septicemia, tularemia and anthrax. Wild boar
Wild boar
may on occasion contract swine erysipelas through rodents or hog lice and ticks.[3]

Relationships with humans[edit] In culture[edit]

Upper Paleolithic
Upper Paleolithic
cave painting, Altamira, Spain. This is a modern interpretation of one of the earliest known depictions of the species.[71]

Depiction of wild boars at Lake Balaton
Lake Balaton
on silver dish (part of the 4th century Sevso Treasure

The wild boar features prominently in the cultures of Indo-European people, many of which saw the animal as embodying warrior virtues. Cultures throughout Europe
Europe
and Asia Minor
Asia Minor
saw the killing of a boar as proof of one's valor and strength. Neolithic
Neolithic
hunter gatherers depicted reliefs of ferocious wild boars on their temple pillars at Göbekli Tepe some 11,600 years ago.[72][73] Virtually all heroes in Greek mythology fight or kill a boar at one point. The demigod Herakles' third labour involves the capture of the Erymanthian Boar, Theseus slays the wild sow Phaea, and a disguised Odysseus
Odysseus
is recognised by his handmaiden Eurycleia
Eurycleia
by the scars inflicted on him by a boar during a hunt in his youth.[74] To the mythical Hyperboreans, the boar represented spiritual authority.[71] Several Greek myths use the boar as a symbol of darkness, death, and winter. One example is the story of the youthful Adonis, who is killed by a boar and is permitted by Zeus
Zeus
to depart from Hades only during the spring and summer period. This theme also occurs in Irish and Egyptian mythology, where the animal is explicitly linked to the month of October, therefore autumn. This association likely arose from aspects of the boar's actual nature. Its dark color was linked to the night, while its solitary habits, proclivity to consume crops and nocturnal nature were associated with evil.[75] The foundation myth of Ephesus
Ephesus
has the city being built over the site where prince Androklos of Athens
Athens
killed a boar.[76] Boars were frequently depicted on Greek funerary monuments alongside lions, representing gallant losers who have finally met their match, as opposed to victorious hunters as lions are. The theme of the doomed, yet valorous boar warrior also occurred in Hittite culture, where it was traditional to sacrifice a boar alongside a dog and a prisoner of war after a military defeat.[74]

A 3rd century CE sculpture from Uttar Pradesh
Uttar Pradesh
of Varaha, an avatar of Vishnu

The boar as a warrior also appears in Scandinavian, Germanic and Anglo-Saxon culture, with its image having been frequently engraved on helmets, shields, and swords. According to Tacitus, the Baltic Aesti featured boars on their helmets, and may have also worn boar masks (see for example the Guilden Morden boar). The boar and pig were held in particularly high esteem by the Celts, who considered them to be their most important sacred animal. Some Celtic deities
Celtic deities
linked to boars include Moccus and Veteris. It has been suggested that some early myths surrounding the Welsh hero Culhwch
Culhwch
involved the character being the son of a boar god.[74] Nevertheless, the importance of the boar as a culinary item among Celtic tribes may have been exaggerated in popular culture by the Asterix
Asterix
series, as wild boar bones are rare among Celtic archaeological sites, and the few that occur show no signs of butchery, having probably been used in sacrificial rituals.[77] The boar also appears in Vedic mythology. A story present in the Brāhmaṇas has Indra
Indra
slaying an avaricious boar, who has stolen the treasure of the asuras, then giving its carcass to Vishnu, who offered it as a sacrifice to the gods. In the story's retelling in the Charaka Samhita, the boar is described as a form of Prajāpti, and is credited with having raised the earth from the primeval waters. In the Rāmāyaṇa and the Purāṇas, the same boar is portrayed as an avatar of Vishnu.[78]

Herakles
Herakles
brings Eurystheus
Eurystheus
the Erymanthian boar, as depicted on a black-figure amphora (c. 550 BC) from Vulci.

In Japanese culture, the boar is widely seen as a fearsome and reckless animal, to the point that several words and expressions in Japanese referring to recklessness include references to boars. The boar is the last animal of the oriental zodiac, with people born during the year of the Pig
Pig
being said to embody the boar-like traits of determination and impetuosity. Among Japanese hunters, the boar's courage and defiance is a source of admiration, and it is not uncommon for hunters and mountain people to name their sons after the animal inoshishi (猪). Boars are also seen as symbols of fertility and prosperity; in some regions, it is thought that boars are drawn to fields owned by families including pregnant women, and hunters with pregnant wives are thought to have greater chances of success when boar hunting. The animal's link to prosperity was illustrated by its inclusion on the ¥10 note during the Meiji period, and it was once believed that a man could become wealthy by keeping a clump of boar hair in his wallet.[79] In the folklore of the Mongol Altai Uriankhai
Altai Uriankhai
tribe, the wild boar was associated with the watery underworld, as it was thought that the spirits of the dead entered the animal's head, to be ultimately transported to the water.[80] Prior to the conversion to Islam, the Kyrgyz people
Kyrgyz people
believed that they were descended from boars, and thus did not eat pork. In Buryat mythology, the forefathers of the Buryats descended from heaven and were nourished by a boar.[81] In China, the boar is the emblem of the Miao people.[71] The boar (sanglier) is frequently displayed in English, Scottish and Welsh heraldry. As with the lion, the boar is often shown as armed and langued. As with the bear, Scottish and Welsh heraldry
Welsh heraldry
displays the boar's head with the neck cropped, unlike the English version, which retains the neck.[82] The white boar served as the badge of King Richard III of England, who distributed it among his northern retainers during his tenure as Duke of Gloucester.[83]

As a game animal and food source[edit] Main article: Boar hunting

Wild boar
Wild boar
haunches and trophy, Umbria, Italy.

A wild boar dish served in Helsinki, Finland.

Humans have been hunting boar for millennia, with the earliest artistic depictions of such activities dating back to the Upper Paleolithic.[74] The animal was seen as a source of food among the Ancient Greeks, as well as a sporting challenge and source of epic narratives. The Romans inherited this tradition, with one of its first practitioners being Scipio Aemilianus. Boar hunting
Boar hunting
became particularly popular among the young nobility during the 3rd century BC as preparation for manhood and battle. A typical Roman boar hunting tactic involved surrounding a given area with large nets, then flushing the boar with dogs and immobilizing it with smaller nets. The animal would then be dispatched with a venabulum, a short spear with a crossguard at the base of the blade. More than their Greek predecessors, the Romans extensively took inspiration from boar hunting in their art and sculpture. With the ascension of Constantine the Great, boar hunting took on Christian allegorical themes, with the animal being portrayed as a "black beast" analogous to the dragon of Saint George. Boar hunting
Boar hunting
continued after the fall of the Western Roman Empire, though the Germanic tribes considered the red deer to be a more noble and worthy quarry. The post-Roman nobility hunted boar as their predecessors did, but primarily as training for battle rather than sport. It was not uncommon for medieval hunters to deliberately hunt boars during the breeding season when the animals were more aggressive. During the Renaissance, when deforestation and the introduction of firearms reduced boar numbers, boarhunting became the sole prerogative of the nobility, one of many charges brought up against the rich during the German Peasants' War
German Peasants' War
and the French Revolution.[84] During the mid-20th century, 7,000–8,000 boars were caught in the Caucasus, 6,000–7,000 in Kazakhstan, and about 5,000 in Central Asia
Central Asia
during the Soviet
Soviet
period, primarily through use of dogs and beats.[3] In Nepal, farmers and poachers eliminate boars by baiting balls of wheat flour containing explosives with kerosene oil, with the animals' chewing motions triggering the devices.[85] Wild boar
Wild boar
can thrive in captivity, though piglets grow slowly and poorly without their mothers. Products derived from wild boar include meat, hide and bristles.[3] Apicius
Apicius
devotes a whole chapter to the cooking of boar meat, providing ten recipes involving roasting, boiling and what sauces to use. The Romans usually served boar meat with garum.[86] Boar's head was the centrepiece of most medieval Christmas
Christmas
celebrations among the nobility.[87] Although growing in popularity as a captive-bred source of food, the wild boar takes longer to mature than most domestic pigs, and is usually smaller and produces less meat. Nevertheless, wild boar meat is leaner and healthier than pork,[88] being of higher nutritional value and having a much higher concentration of essential amino acids.[89] Most meat-dressing organizations agree that a boar carcass should yield 50 kg (110 lb) of meat on average. Large specimens can yield 15–20 kg (33–44 lb) of fat, with some giants yielding 30 kg (66 lb) or more. A boar hide can measure 300 dm2, and can yield 350–1000 grams of bristle and 400 grams of underwool.[3]

Roman relief of a dog confronting a boar, Cologne

Southern Indian depiction of boar hunt, c. 1540

Pigsticking in British India

Boar shot in Volgograd Oblast, Russia

The Boar Hunt – Hans Wertinger, c. 1530, Danube Valley

Crop and garbage raiding[edit]

An adult sow and young that have broken open a litter bag in Berlin seeking food.

Boars can be damaging to agriculture. Populations living on the outskirts of towns or farms can dig up potatoes and damage melons, watermelons and maize. They generally only encroach upon farms when natural food is scarce. In the Belovezh forest for example, 34–47% of the local boar population will enter fields in years of moderate availability of natural foods. While the role of boars in damaging crops is often exaggerated,[3] cases are known of boar depredations causing famines, as was the case in Hachinohe, Japan
Japan
in 1749, where 3,000 people died of what became known as the 'wild boar famine'. Still, within Japanese culture, the boar's status as vermin is expressed through its title as "king of pests" and the popular saying (addressed to young men in rural areas) "When you get married, choose a place with no wild boar."[79][90] In Central Europe, farmers typically repel boars through distraction or fright, while in Kazakhstan
Kazakhstan
it is usual to employ guard dogs in plantations. Although large boar populations can play an important role in limiting forest growth, they are also useful in keeping pest populations such as June bugs under control.[3] The growth of urban areas and the corresponding decline in natural boar habitats has led to some sounders entering human habitations in search of food. As in natural conditions, sounders in peri-urban areas are matriarchal, though males tend to be much less represented, and adults of both sexes can be up to 35% heavier than their forest-dwelling counterparts. As of 2010, at least 44 cities in 15 countries have experienced problems of some kind relating to the presence of habituated wild boar.[91] Attacks on humans[edit]

Depiction of a stylised boar attacking a man, Bhimbetaka, India

Actual attacks on humans are rare, but can be serious, resulting in multiple penetrating injuries to the lower part of the body. They generally occur during the boars' rutting season from November to January, in agricultural areas bordering forests or on paths leading through forests. The animal typically attacks by charging and pointing its tusks towards the intended victim, with most injuries occurring on the thigh region. Once the initial attack is over, the boar steps back, takes position and attacks again if the victim is still moving, only ending once the victim is completely incapacitated.[92][93] Boar attacks on humans have been documented since the Stone Age, with one of the oldest depictions being a cave painting in Bhimbetaka, India. The Romans and Ancient Greeks wrote of these attacks (Odysseus was wounded by a boar, and Adonis
Adonis
was killed by one). A 2012 study compiling recorded attacks from 1825–2012 found accounts of 665 human victims of both wild boars and feral pigs, with the majority (19%) of attacks in the animal's native range occurring in India. Most of the attacks occurred in rural areas during the winter months in non-hunting contexts and were committed by solitary males.[94] See also[edit]

Mammals portal Animals portal

Boar-baiting Domestic pig Feral pig Peccary Indian boar Guilden Morden boar

Notes[edit]

^ It is from the male boar's solitary habits that the species gets its name in numerous Romance languages. Although the Latin
Latin
word for "boar" was aper, the French sanglier and Italian cinghiale derive from singularis porcus, which is Latin
Latin
for "solitary pig".[38] ^ Thirteen has been observed in a captive specimen.[40]

References[edit]

^ a b c Oliver, W. & Leus, K. (2008). "Sus scrofa". The IUCN
IUCN
Red List of Threatened Species. IUCN. 2008: e.T41775A10559847. doi:10.2305/IUCN.UK.2008.RLTS.T41775A10559847.en. Retrieved 13 January 2018.  Database entry includes a brief justification of why this species is of least concern. ^ a b c Wozencraft, W.C. (2005). "Order Carnivora". In Wilson, D.E.; Reeder, D.M. Mammal
Mammal
Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 532–628. ISBN 978-0-8018-8221-0. OCLC 62265494.  ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap Heptner, V. G. ; Nasimovich, A. A. ; Bannikov, A. G. ; Hoffman, R. S. (1988) Mammals of the Soviet
Soviet
Union, Volume I, Washington, D.C. : Smithsonian Institution Libraries and National Science Foundation, pp. 19–82 ^ a b c Oliver, W. L. R. et al. 1993. The Eurasian Wild Pig
Pig
(Sus scrofa). In Oliver, W. L. R., ed., Pigs, Peccaries, and Hippos – 1993 Status Survey and Conservation Action Plan, 112–121. IUCN/SSC Pigs and Peccaries Specialist Group, ISBN 2-8317-0141-4 ^ https://www.britannica.com/animal/boar-mammal ^ a b Chen, K.; et al. (2007). "Genetic Resources, Genome Mapping and Evolutionary Genomics of the Pig
Pig
(Sus scrofa)". Int J Biol Sci. 3 (3): 153–165. doi:10.7150/ijbs.3.153. CS1 maint: Explicit use of et al. (link) ^ a b Kurtén, Björn (1968). Pleistocene
Pleistocene
mammals of Europe. Weidenfeld and Nicolson. pp. 153–155 ^ a b Marsan & Mattioli 2013, pp. 75–76 ^ a b c d e f g h i Baskin, L. & Danell, K. (2003), Ecology of Ungulates: A Handbook of Species in Eastern Europe
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and Northern and Central Asia, Springer Science & Business Media, pp. 15–38, ISBN 3-540-43804-1 ^ a b c Affenberg, W. (1981), The Behavioral Ecology of the Komodo Monitor, University Press of Florida, pp. 248, ISBN 081300621X ^ "Online Etymological Dictionary". Retrieved 2014-10-08.  ^ " Latin
Latin
Dictionary and Grammar Resources". Retrieved 2014-10-08.  ^ Cabanau 2001, pp. 24 ^ Ruvinsky, A. et al. (2011). "Systematics and evolution of the pig". In: Ruvinsky A, Rothschild MF (eds), The Genetics of the Pig. 2nd ed. CAB International, Oxon. pp. 1–13. ISBN 978-1-84593-756-0 ^ a b c d Groves, C. P. et al. 1993. The Eurasian Suids Sus and Babyrousa. In Oliver, W. L. R., ed., Pigs, Peccaries, and Hippos – 1993 Status Survey and Conservation Action Plan, 107–108. IUCN/SSC Pigs and Peccaries Specialist Group, ISBN 2-8317-0141-4 ^ a b Hemmer, H. (1990), Domestication: The Decline of Environmental Appreciation, Cambridge University Press, pp. 55–59, ISBN 0-521-34178-7 ^ Kingdon, J. (1997). The Kingdon Guide to African Mammals. p. 329. Academic Press Limited. ISBN 0-12-408355-2 ^ a b c d e f g h Groves, C. (2008). Current views on the taxonomy and zoogeography of the genus Sus. pp. 15–29 in Albarella, U., Dobney, K, Ervynck, A. & Rowley-Conwy, P. Eds. (2008). Pigs and Humans: 10,000 Years of Interaction. Oxford University Press. ISBN 978-0-19-920704-6 ^ Sterndale, R. A. (1884), Natural history of the Mammalia of India and Ceylon, Calcutta : Thacker, Spink, pp. 415–420 ^ Scheggi 1999, pp. 86–89 ^ Marsan & Mattioli 2013, pp. 14–15 ^ Clutton-Brock, J. (1999), A Natural History of Domesticated Mammals, Cambridge University Press, pp. 91–99, ISBN 0-521-63495-4 ^ The related peccary (Pecari tajacu) has been domesticated in the New World. "Commercial Farming of Collared Peccary: A Large-scale Commercial Farming of Collared Peccary
Peccary
(Tayassu tajacu) in North-eastern Brazil", 2007-4-30, http://pigtrop.cirad.fr/subjects/genetic_and_biodiversity/commercial_farming_of_collared_peccary . ^ *Sarah M. Nelson Ancestors for the Pigs. Pigs in prehistory. (1998) ^ Rosenberg M, Nesbitt R, Redding RW, Peasnall BL (1998). Hallan Çemi, pig husbandry, and post- Pleistocene
Pleistocene
adaptations along the Taurus-Zagros Arc (Turkey). Paleorient, 24(1):25–41. ^ Vigne, JD; Zazzo, A; Saliège, JF; Poplin, F; Guilaine, J; Simmons, A (2009). "Pre- Neolithic
Neolithic
wild boar management and introduction to Cyprus
Cyprus
more than 11,400 years ago". Proceedings of the National Academy of Sciences of the United States of America. 106 (38): 16135–8. doi:10.1073/pnas.0905015106. PMC 2752532 . PMID 19706455.  ^ a b Giuffra, E; Kijas, JM; Amarger, V; Carlborg, O; Jeon, JT; Andersson, L (2000). "The origin of the domestic pig: independent domestication and subsequent introgression". Genetics. 154 (4): 1785–91. PMC 1461048 . PMID 10747069.  ^ Jean-Denis Vigne; Anne Tresset & Jean-Pierre Digard (July 3, 2012). History of domestication (PDF) (Speech).  ^ BBC News, " Pig
Pig
DNA
DNA
reveals farming history" 4 September 2007. The report concerns an article in the journal PNAS ^ Larson, G; Albarella, U; Dobney, K; Rowley-Conwy, P; Schibler, J; Tresset, A; Vigne, JD; Edwards, CJ; et al. (2007). "Ancient DNA, pig domestication, and the spread of the Neolithic
Neolithic
into Europe". Proceedings of the National Academy of Sciences of the United States of America. 104 (39): 15276–81. doi:10.1073/pnas.0703411104. PMC 1976408 . PMID 17855556.  ^ Scheggi 1999, pp. 87 ^ a b c d e Marsan & Mattioli 2013, pp. 70–72 ^ Marsan & Mattioli 2013, pp. 26 ^ Marsan & Mattioli 2013, pp. 28 ^ a b c d Cabanau 2001, pp. 29 ^ a b Cabanau 2001, pp. 28 ^ Drabeck, D.H.; Dean, A.M.; Jansa, S.A. (June 1, 2015). "Why the honey badger don't care: Convergent evolution of venom-targeted nicotinic acetylcholine receptors in mammals that survive venomous snake bites". Toxicon. Elsevier. 99: 68–72. doi:10.1016/j.toxicon.2015.03.007. PMID 25796346.  ^ Scheggi 1999, pp. 20–22 ^ Weiler, Ulrike; et al. (2016). "Penile Injuries in Wild and Domestic Pigs". Animals. 6 (4): 25. doi:10.3390/ani6040025. CS1 maint: Explicit use of et al. (link) ^ "Eight little piggies went a-marching". Whipsnade Zoo. Retrieved 22 May 2015.  ^ a b Marsan & Mattioli 2013, pp. 83–86 ^ Marsan & Mattioli 2013, pp. 87–90 ^ a b Marsan & Mattioli 2013, pp. 55–58 ^ Long, J. L. (2003), Introduced Mammals of the World: Their History, Distribution and Influence, Cabi Publishing, ISBN 978-0-85199-748-3 ^ " Alderney
Alderney
wild boar that swam from France
France
shot over disease fear". BBC News. 14 November 2013.  ^ Tierney, Trisha A.; Cushman, J. Hall (2006-05-12). "Temporal Changes in Native and Exotic Vegetation and Soil Characteristics following Disturbances by Feral Pigs in a California
California
Grassland". Biological Invasions. 8 (5): 1073–1089. doi:10.1007/s10530-005-6829-7.  ^ Oldfield, Callie A.; Evans, Jonathan P. (2016-03-01). "Twelve years of repeated wild hog activity promotes population maintenance of an invasive clonal plant in a coastal dune ecosystem". Ecology and Evolution. 6 (8): 2569–2578. doi:10.1002/ece3.2045.  ^ Gupta, S. et al. A wild boar hunting: predation on a bonnet macaque by a wild boar in the Bandipur National Park, southern India. Current Science. Vol. 106, No. 9. (10 May 2014) ^ a b Marsan & Mattioli 2013, pp. 96–97 ^ Thinley, P; Kamler, JF; Wang, SW; Lham, K; Stenkewitz, U; et al. (2011). "Seasonal diet of dholes (Cuon alpinus) in northwestern Bhutan". Mamm Biol. 76 (4): 518–520. doi:10.1016/j.mambio.2011.02.003. CS1 maint: Explicit use of et al. (link) ^ Heptner, V. G.; Sludskii, A. A. (1992). Mammals of the Soviet
Soviet
Union: Carnivora (hyaenas and cats), Volume 2. Smithsonian Institution Libraries and National Science Foundation. p. 248–252. ^ Taghdisi, M.; et al. (2013). "Diet and habitat use of the endangered Persian leopard (Panthera pardus saxicolor) in northeastern Iran" (PDF). Turkish Journal of Zoologist. 37: 554–561. doi:10.3906/zoo-1301-20. CS1 maint: Explicit use of et al. (link) ^ Elmira Sharbafi, Mohammad S. Farhadinia, Hamid R. Rezaie, Alex Richard Braczkowski. (2016) Prey of the Persian Leopard
Leopard
( Panthera pardus saxicolor ) in a mixed forest-steppe landscape in northeastern Iran
Iran
(Mammalia: Felidae). Zoology in the Middle East, 1–8. ^ Heptner, V. G.; Sludskii, A. A. (1992). Mammals of the Soviet
Soviet
Union: Carnivora (hyaenas and cats), Volume 2. Smithsonian Institution Libraries and National Science Foundation. p. 174, 185. ^ Miquelle, Dale G.; et al. (1996). "Food habits of Amur tigers in the Sikhote-Alin Zapovednik and the Russian Far East, and implications for conservation" (PDF). Journal of Wildlife Research. 1 (2): 138. Archived from the original (PDF) on 1 November 2012.  ^ Ramesh, T.; Snehalatha, V.; Sankar, K. & Qureshi, Qamar (2009). "Food habits and prey selection of tiger and leopard in Mudumalai tiger Reserve, Tamil Nadu, India". J. Sci. Trans. Environ. Technov. 2 (3): 170–181.  ^ Schaller, G (1967). The deer and the tiger: a study of wildlife in India. p. 321.  ^ Masseti, M. (2012), Atlas of terrestrial mammals of the Ionian and Aegean islands, Walter de Gruyter, pp. 139–141, ISBN 3-11-025458-1 ^ a b c " Wild boar
Wild boar
in Britain". Britishwildboar.org.uk. 21 October 1998. Retrieved 30 July 2013.  ^ Osborn, Dale. J.; Helmy, Ibrahim (1980), "The contemporary land mammals of Egypt
Egypt
(including Sinai)", Field Museum of Natural History, pp. 475–477 ^ Government supports local communities to manage wild boar. Department for Environment, Food and Rural Affairs. 19 February 2008 ^ http://www.bbc.co.uk/programmes/b080wbwy ^ " Wild boar
Wild boar
cull is given go ahead". BBC News. 4 January 2010.  ^ " Forest of Dean
Forest of Dean
rangers battle to meet boar cull target". BBC News. 20 August 2010. Retrieved 13 November 2010.  ^ Cull failing to control wild boar. The Forester. 25 February 2011. ^ "BBC Wales – Nature – Wildlife – Wild boar". Bbc.co.uk. 1 January 1970. Retrieved 30 July 2013.  ^ "Wild Boar in Britain". Britishwildboar.org.uk. 31 December 2010. Retrieved 30 July 2013.  ^ Monbiot, George (16 September 2011). "How the UK's zoophobic legacy turned on wild boar". The Guardian. London. Retrieved 16 September 2011. I was prompted to write this article by an item I heard on the BBC's Farming Today programme at the beginning of the week. It was an interview with Ralph Harmer, who works for the Forestry Commission, about whether or not the returning boars are damaging our woodlands. I was struck by what the item did not say. Not once did the programme mention that this is a native species. The boar was discussed as if it were an exotic invasive animal, such as the mink or the grey squirrel. […] Then, once we've found out how many boars, […] should be culled to allow a gentle expansion but not an explosion, permits to shoot them should be sold, and the money used to compensate farmers whose crops the boar have damaged. Other hunting should be banned. This is how they do it in France.  ^ http://www.dec.ny.gov/animals/70843.html ^ a b c d e f Mayer, J. J. et al. (2009), Wild Pigs: Biology, Damage, Control Techniques and Management Archived 24 October 2014 at the Wayback Machine., Savannah River National Laboratory Aiken, South Carolina, SRNL-RP-2009-00869 ^ a b c Cabanau 2001, pp. 63 ^ Charles C. Mann, Göbekli Tepe: The Birth of Religion, National Geographic (June 2011) ^ Sandra Scham The World's First Temple, Archaeology, Volume 61 Number 6, November/December 2008 ^ a b c d Mallory, J. P. & Adams, D. Q. (1997), Encyclopedia of Indo-European Culture, Taylor & Francis, pp. 426–428, ISBN 1-884964-98-2 ^ Scheggi 1999, pp. 14–15 ^ Scheggi 1999, pp. 16 ^ Green, M. (2002), Animals in Celtic Life and Myth, Routledge, p. 46, ISBN 1-134-66531-8 ^ Macdonell, A. A. (1898), Vedic Mythology, Motilal Banarsidass Publ., p. 41 ^ a b Knight, J. (2003), Waiting for Wolves in Japan: An Anthropological Study of People-wildlife Relations, Oxford University Press, pp. 49–73, ISBN 0-19-925518-0 ^ Pegg, C. (2001), Mongolian Music, Dance, & Oral Narrative: Performing Diverse Identities, University of Washington Press, p. 140, ISBN 0-295-98112-1 ^ Holmberg, U. (1927), The Mythology of All Races volume 4: Finno-Ugric, Siberian, New York, Cooper Square Publishing Inc. pp. 502–503 ^ Fox-Davies, A. C. (1909), A complete guide to heraldry, London, Edinburgh, T.C. & E.C. Jack, pp. 198–199 ^ Wagner, J. A. (2001) Encyclopedia of the Wars of the Roses, ABC-CLIO, p. 15, ISBN 1-85109-358-3 ^ Scheggi 1999, pp. 9–58 ^ Shreshta, Tej Kumar (1997). Mammals of Nepal: (with reference to those of India, Bangladesh, Bhutan and Pakistan). Steven Simpson Books. p. 207. ISBN 0-9524390-6-9 ^ Scheggi 1999, pp. 30–35 ^ Adamson, M. W. (2004), Food in Medieval Times, Greenwood Publishing Group, p. 35, ISBN 0-313-32147-7 ^ Harris, C. (2009), A Guide to Traditional Pig
Pig
Keeping, Good Life Press, pp. 26–27, ISBN 1-904871-60-7 ^ Strazdina, V. et al. "Nutritional Characteristics of Wild Boar Meat Hunted in Latvia", Foodbalt (2014) ^ Walker, B. "Commercial Growth and Environmental Change in Early Modern Japan: Hachinohe’s Wild Boar Famine," The Journal of Asian Studies, Vol. 60, No. 2 (May, 2001), pp. 331. ^ Cahill, S.; Llimona, F.; Cabañeros, L.; Calomardo, F. (2012). "Characteristics of wild boar (Sus scrofa) habituation to urban areas in the Collserola Natural Park (Barcelona) and comparison with other locations" (PDF). Animal
Animal
Biodiversity and Conservation. 35 (2): 221–233.  ^ Manipady, S.; et al. (2006). "Death by attack from a wild boar" (PDF). Journal of clinical forensic medicine. 13 (2): 89–91. doi:10.1016/j.jcfm.2005.08.007. PMID 16263321. CS1 maint: Explicit use of et al. (link) ^ Gunduz, A.; et al. (2007). "Wild Boar Attacks" (PDF). Wilderness and Environmental Medicine. 18 (2): 117–119. doi:10.1580/06-weme-cr-033r1.1. PMID 17590063. CS1 maint: Explicit use of et al. (link) ^ Mayer, John J. (2013) "Wild Pig
Pig
Attacks on Humans". Wildlife Damage Management Conferences – Proceedings. Paper 151.

Bibliography[edit]

Cabanau, Laurent (2001). The Hunter's Library: Wild Boar in Europe. Könemann. ISBN 3-8290-5528-5.  Marsan, Andrea; Mattioli, Stefano (2013). Il Cinghiale (in Italian). Il Piviere (collana Fauna selvatica. Biologia e gestione). ISBN 978-88-96348-178.  Scheggi, Massimo (1999). La bestia nera: Caccia al cinghiale fra mito, storia e attualità (in Italian). Editoriale Olimpia (collana Caccia). ISBN 88-253-7904-8. 

Further reading[edit]

Apollonio, M. et al. (1988), "The systematics of the wild boar (Sus scrofa L.) in Italy", Bolletino di Zoologia, 3:213–221 Carden, R.F. (2012) "Review of the Natural History of Wild Boar (Sus scrofa) on the island of Ireland", Report prepared by Ruth Carden for the Northern Ireland Environment Agency, Northern Ireland, UK, National Parks & Wildlife Service, Department of Arts, Heritage and the Gaeltacht, Dublin, Ireland and the National Museum of Ireland – Education & Outreach Department. (in French) Durantel, P. (2007), Le sanglier et ses chasses, Editions Artemis, ISBN 2844166032 Greene, J. (2011), The Golden-Bristled Boar: Last Ferocious Beast of the Forest, University of Virginia Press, ISBN 0-8139-3103-7 (in French) Marillier, B. (2003), Le sanglier héraldique, Editions Cheminements, ISBN 2844781845 Mayer, J. J. & Shedrow, C. B. (2007), Annotated Bibliography of the Wild Pig
Pig
(Sus scrofa): Environmental Information Document, Washington Savannah River Company (in Italian) Padiglione, V. (1989), Il cinghiale cacciatore: Antropologia simbolica della caccia in Sardegna, Armando Editore (collana Antropologia culturale) Ronald M. Nowak (1999), Walker’s Mammals of the World (6th ed.), Baltimore: Johns Hopkins University Press, ISBN 0-8018-5789-9, LCCN 98023686 

External links[edit]

Wikisource
Wikisource
has the text of the 1911 Encyclopædia Britannica
Encyclopædia Britannica
article Boar.

Wikispecies
Wikispecies
has information related to Sus scrofa

Wikiquote has quotations related to: Wild boar

News related to Saskatchewan places moratorium on boar farming, says escaped boars should be killed at Wikinews Media related to Sus scrofa at Wikimedia Commons BBC profile  "Boar, Wild". Encyclopædia Britannica. 3 (9th ed.). 1878.  Jokelainen, P.; Näreaho, A.; Hälli, O.; Heinonen, M.; Sukura, A. (2012). "Farmed wild boars exposed to Toxoplasma gondii and Trichinella spp". Veterinary Parasitology. 187 (1–2): 323–327. doi:10.1016/j.vetpar.2011.12.026. PMID 22244535.  Species Profile- Wild Boar (Sus scrofa), National Invasive Species Information Center, United States National Agricultural Library. Lists general information and resources for wild boar. View the susScr3 genome assembly in the UCSC Genome Browser.

v t e

Pigs

Kingdom Animalia Phylum Chordata Class Mammalia Order Artiodactyla Suborder Suina Family Suidae Subfamily Suinae Genus Sus

Domestic

Breeding Shows

Farming Intensive farming Sty Pannage Toilet Hog oiler Gestation crate Cross-fostering Swineherd

As food Bacon Ham Lard Pork Scalder Slaughter Suckling Blood Religious restrictions Scottish pork taboo

Cuts Back bacon Boston butt Fatback Ham
Ham
hock Pig's trotters Pork
Pork
belly Pork
Pork
chop Pork
Pork
jowl Loin Tenderloin Ribs Spare ribs Pork
Pork
rind Pork
Pork
steak Ear Tail

Other uses Bladder Racing War Wrestling Truffling

Wild and Feral

Wild boar

Heraldry Hunting

In the Philippines Razorback

Pigs in culture

Iron Age pig Flitch of bacon custom Miss Piggy

Pigs in Space

Porky Pig Piglet "The Three Little Pigs" Animal
Animal
Farm

Old Major Napoleon Snowball Squealer

Babe Babe: Pig
Pig
in the City Bad Piggies My Brother the Pig Charlotte's Web The Sheep-Pig Peppa Pig Super Pig Fair, then Partly Piggy Jakers! The Adventures of Piggley Winks Olivia (character TV series) Pink Floyd pigs Pinky and Perky Preston Pig Rasher Porco Rosso Spider-Ham Spider pig The Tale of Little Pig
Pig
Robinson The Tale of Pigling Bland "This Little Piggy" Huxley Pig Wibbly Pig Zhu Bajie

Other

List of pigs List of fictional pigs Piganino Pigasus (politics) "When pigs fly"

v t e

Extant Artiodactyla species

Kingdom: Animalia Phylum: Chordata Class: Mammalia Infraclass: Eutheria Superorder: Laurasiatheria

Suborder Ruminantia

Antilocapridae

Antilocapra

Pronghorn
Pronghorn
(A. americana)

Giraffidae

Okapia

Okapi
Okapi
(O. johnstoni)

Giraffa

Northern giraffe
Northern giraffe
(G. camelopardalis) Southern giraffe
Southern giraffe
(G. giraffa) Reticulated giraffe
Reticulated giraffe
(G. reticulata) Masai giraffe
Masai giraffe
(G. tippelskirchi)

Moschidae

Moschus

Anhui musk deer
Anhui musk deer
(M. anhuiensis) Dwarf musk deer
Dwarf musk deer
(M. berezovskii) Alpine musk deer
Alpine musk deer
(M. chrysogaster) Kashmir musk deer
Kashmir musk deer
(M. cupreus) Black musk deer
Black musk deer
(M. fuscus) Himalayan musk deer (M. leucogaster) Siberian musk deer
Siberian musk deer
(M. moschiferus)

Tragulidae

Hyemoschus

Water chevrotain
Water chevrotain
(H. aquaticus)

Moschiola

Indian spotted chevrotain
Indian spotted chevrotain
(M. indica) Yellow-striped chevrotain
Yellow-striped chevrotain
(M. kathygre) Sri Lankan spotted chevrotain
Sri Lankan spotted chevrotain
(M. meminna)

Tragulus

Java
Java
mouse-deer (T. javanicus) Lesser mouse-deer
Lesser mouse-deer
(T. kanchil) Greater mouse-deer
Greater mouse-deer
(T. napu) Philippine mouse-deer
Philippine mouse-deer
(T. nigricans) Vietnam
Vietnam
mouse-deer (T. versicolor) Williamson's mouse-deer
Williamson's mouse-deer
(T. williamsoni)

Cervidae

Large family listed below

Bovidae

Large family listed below

Family Cervidae

Cervinae

Muntiacus

Indian muntjac
Indian muntjac
(M. muntjak) Reeves's muntjac
Reeves's muntjac
(M. reevesi) Hairy-fronted muntjac
Hairy-fronted muntjac
(M. crinifrons) Fea's muntjac
Fea's muntjac
(M. feae) Bornean yellow muntjac
Bornean yellow muntjac
(M. atherodes) Roosevelt's muntjac
Roosevelt's muntjac
(M. rooseveltorum) Gongshan muntjac
Gongshan muntjac
(M. gongshanensis) Giant muntjac
Giant muntjac
(M. vuquangensis) Truong Son muntjac
Truong Son muntjac
(M. truongsonensis) Leaf muntjac
Leaf muntjac
(M. putaoensis) Sumatran muntjac
Sumatran muntjac
(M. montanus) Pu Hoat muntjac
Pu Hoat muntjac
(M. puhoatensis)

Elaphodus

Tufted deer
Tufted deer
(E. cephalophus)

Dama

Fallow deer
Fallow deer
(D. dama) Persian fallow deer
Persian fallow deer
(D. mesopotamica)

Axis

Chital
Chital
(A. axis)

Rucervus

Barasingha
Barasingha
(R. duvaucelii)

Panolia

Eld's deer
Eld's deer
(P. eldii)

Elaphurus

Père David's deer
Père David's deer
(E. davidianus)

Hyelaphus

Hog deer (H. porcinus) Calamian deer
Calamian deer
(H. calamianensis) Bawean deer
Bawean deer
(H. kuhlii)

Rusa

Sambar deer
Sambar deer
(R. unicolor) Rusa deer (R. timorensis) Philippine sambar (R. mariannus) Philippine spotted deer (R. alfredi)

Cervus

Red deer
Red deer
(C. elaphus) Elk
Elk
(C. canadensis) Thorold's deer
Thorold's deer
(C. albirostris) Sika deer
Sika deer
(C. nippon)

Capreolinae

Alces

Moose
Moose
(A. alces)

Hydropotes

Water deer
Water deer
(H. inermis)

Capreolus

Roe deer
Roe deer
(C. capreolus) Siberian roe deer
Siberian roe deer
(C. pygargus)

Rangifer

Reindeer
Reindeer
(R. tarandus)

Hippocamelus

Taruca
Taruca
(H. antisensis) South Andean deer
South Andean deer
(H. bisulcus)

Mazama

Red brocket
Red brocket
(M. americana) Small red brocket
Small red brocket
(M. bororo) Merida brocket
Merida brocket
(M. bricenii) Dwarf brocket
Dwarf brocket
(M. chunyi) Gray brocket
Gray brocket
(M. gouazoubira) Pygmy brocket
Pygmy brocket
(M. nana) Amazonian brown brocket
Amazonian brown brocket
(M. nemorivaga) Yucatan brown brocket
Yucatan brown brocket
(M. pandora) Little red brocket
Little red brocket
(M. rufina) Central American red brocket
Central American red brocket
(M. temama)

Ozotoceros

Pampas deer
Pampas deer
(O. bezoarticus)

Blastocerus

Marsh
Marsh
deer (B. dichotomus)

Pudu

Northern pudú (P. mephistophiles) Southern pudú (P. pudu)

Odocoileus

White-tailed deer
White-tailed deer
(O. virginianus) Mule deer
Mule deer
(O. hemionus)

Family Bovidae

Cephalophinae

Cephalophus

Abbott's duiker
Abbott's duiker
(C. spadix) Aders's duiker
Aders's duiker
(C. adersi) Bay duiker
Bay duiker
(C. dorsalis) Black duiker
Black duiker
(C. niger) Black-fronted duiker
Black-fronted duiker
(C. nigrifrons) Brooke's duiker (C. brookei) Harvey's duiker
Harvey's duiker
(C. harveyi) Jentink's duiker
Jentink's duiker
(C. jentinki) Ogilby's duiker
Ogilby's duiker
(C. ogilbyi) Peters's duiker (C. callipygus) Red-flanked duiker
Red-flanked duiker
(C. rufilatus) Red forest duiker
Red forest duiker
(C. natalensis) Ruwenzori duiker
Ruwenzori duiker
(C. rubidis) Weyns's duiker
Weyns's duiker
(C. weynsi) White-bellied duiker
White-bellied duiker
(C. leucogaster) White-legged duiker
White-legged duiker
(C. crusalbum) Yellow-backed duiker
Yellow-backed duiker
(C. Sylvicultor) Zebra duiker
Zebra duiker
(C. zebra)

Philantomba

Blue duiker
Blue duiker
(P. monticola) Maxwell's duiker
Maxwell's duiker
(P. maxwellii) Walter's duiker
Walter's duiker
(P. walteri)

Sylvicapra

Common duiker
Common duiker
(S. grimmia)

Hippotraginae

Hippotragus

Roan antelope
Roan antelope
(H. equinus) Sable antelope
Sable antelope
(H. niger)

Oryx

East African oryx
East African oryx
(O. beisa) Scimitar oryx
Scimitar oryx
(O. dammah) Gemsbok
Gemsbok
(O. gazella) Arabian oryx
Arabian oryx
(O. leucoryx)

Addax

Addax
Addax
(A. nasomaculatus)

Reduncinae

Kobus

Upemba lechwe
Upemba lechwe
(K. anselli) Waterbuck
Waterbuck
(K. ellipsiprymnus) Kob
Kob
(K. kob) Lechwe
Lechwe
(K. leche) Nile
Nile
lechwe (K. megaceros) Puku
Puku
(K. vardonii)

Redunca

Southern reedbuck
Southern reedbuck
(R. arundinum) Mountain reedbuck
Mountain reedbuck
(R. fulvorufula) Bohor reedbuck
Bohor reedbuck
(R. redunca)

Aepycerotinae

Aepyceros

Impala
Impala
(A. melampus)

Peleinae

Pelea

Grey rhebok
Grey rhebok
(P. capreolus)

Alcelaphinae

Beatragus

Hirola
Hirola
(B. hunteri)

Damaliscus

Topi
Topi
(D. korrigum) Common tsessebe
Common tsessebe
(D. lunatus) Bontebok
Bontebok
(D. pygargus) Bangweulu tsessebe
Bangweulu tsessebe
(D. superstes)

Alcelaphus

Hartebeest
Hartebeest
(A. buselaphus) Red hartebeest
Red hartebeest
(A. caama) Lichtenstein's hartebeest
Lichtenstein's hartebeest
(A. lichtensteinii)

Connochaetes

Black wildebeest
Black wildebeest
(C. gnou) Blue wildebeest
Blue wildebeest
(C. taurinus)

Pantholopinae

Pantholops

Tibetan antelope
Tibetan antelope
(P. hodgsonii)

Caprinae

Large subfamily listed below

Bovinae

Large subfamily listed below

Antilopinae

Large subfamily listed below

Family Bovidae
Bovidae
(subfamily Caprinae)

Ammotragus

Barbary sheep
Barbary sheep
(A. lervia)

Budorcas

Takin
Takin
(B. taxicolor)

Capra

Wild goat
Wild goat
(C. aegagrus) Domestic goat (C. aegagrus hircus) West Caucasian tur
West Caucasian tur
(C. caucasia) East Caucasian tur
East Caucasian tur
(C. cylindricornis) Markhor
Markhor
(C. falconeri) Alpine ibex
Alpine ibex
(C. ibex) Nubian ibex
Nubian ibex
(C. nubiana) Spanish ibex
Spanish ibex
(C. pyrenaica) Siberian ibex
Siberian ibex
(C. sibirica) Walia ibex
Walia ibex
(C. walie)

Capricornis

Japanese serow
Japanese serow
(C. crispus) Taiwan
Taiwan
serow (C. swinhoei) Sumatran serow
Sumatran serow
(C. sumatraensis) Mainland serow
Mainland serow
(C. milneedwardsii) Red serow
Red serow
(C. rubidusi) Himalayan serow
Himalayan serow
(C. thar)

Hemitragus

Nilgiri tahr
Nilgiri tahr
(H. hylocrius) Arabian tahr
Arabian tahr
(H. jayakari) Himalayan tahr
Himalayan tahr
(H. jemlahicus)

Naemorhedus

Red goral
Red goral
(N. baileyi) Long-tailed goral
Long-tailed goral
(N. caudatus) Himalayan goral
Himalayan goral
(N. goral) Chinese goral
Chinese goral
(N. griseus)

Oreamnos

Mountain goat
Mountain goat
(O. americanus)

Ovibos

Muskox
Muskox
(O. moschatus)

Ovis

Argali
Argali
(O. ammon) Domestic sheep (O. aries) Bighorn sheep
Bighorn sheep
(O. canadensis) Dall sheep
Dall sheep
(O. dalli) Mouflon
Mouflon
(O. musimon) Snow sheep
Snow sheep
(O. nivicola) Urial
Urial
(O. orientalis)

Pseudois

Bharal
Bharal
(P. nayaur) Dwarf blue sheep
Dwarf blue sheep
(P. schaeferi)

Rupicapra

Pyrenean chamois
Pyrenean chamois
(R. pyrenaica) Chamois
Chamois
(R. rupicapra)

Family Bovidae
Bovidae
(subfamily Bovinae)

Boselaphini

Tetracerus

Four-horned antelope
Four-horned antelope
(T. quadricornis)

Boselaphus

Nilgai
Nilgai
(B. tragocamelus)

Bovini

Bubalus

Water buffalo
Water buffalo
(B. bubalis) Wild Water Buffalo (B. arnee) Lowland anoa (B. depressicornis) Mountain anoa (B. quarlesi) Tamaraw
Tamaraw
(B. mindorensis)

Bos

Banteng
Banteng
(B. javanicus) Gaur
Gaur
(B. gaurus) Gayal
Gayal
(B. frontalis) Domestic yak
Domestic yak
(B. grunniens) Wild yak
Wild yak
(B. mutus) Cattle
Cattle
(B. taurus) Kouprey
Kouprey
(B. sauveli)

Pseudonovibos

Kting voar (P. spiralis)

Pseudoryx

Saola
Saola
(P. nghetinhensis)

Syncerus

African buffalo
African buffalo
(S. caffer)

Bison

American bison
American bison
(B. bison) European bison
European bison
(B. bonasus)

Tragelaphini

Tragelaphus (including kudus)

Sitatunga
Sitatunga
(T. spekeii) Nyala
Nyala
(T. angasii) Kéwel
Kéwel
(T. scriptus) Cape bushbuck
Cape bushbuck
(T. sylvaticus) Mountain nyala
Mountain nyala
(T. buxtoni) Lesser kudu
Lesser kudu
(T. imberbis) Greater kudu
Greater kudu
(T. strepsiceros) Bongo (T. eurycerus)

Taurotragus

Common eland
Common eland
(T. oryx) Giant eland
Giant eland
(T. derbianus)

Family Bovidae
Bovidae
(subfamily Antilopinae)

Antilopini

Ammodorcas

Dibatag
Dibatag
(A. clarkei)

Antidorcas

Springbok
Springbok
(A. marsupialis)

Antilope

Blackbuck
Blackbuck
(A. cervicapra)

Eudorcas

Mongalla gazelle
Mongalla gazelle
(E. albonotata) Red-fronted gazelle
Red-fronted gazelle
(E. rufifrons) Thomson's gazelle
Thomson's gazelle
(E. thomsonii) Heuglin's gazelle
Heuglin's gazelle
(E. tilonura)

Gazella

Mountain gazelle
Mountain gazelle
(G. gazella) Neumann's gazelle (G. erlangeri) Speke's gazelle
Speke's gazelle
(G. spekei) Dorcas gazelle
Dorcas gazelle
(G. dorcas) Chinkara
Chinkara
(G. bennettii) Cuvier's gazelle
Cuvier's gazelle
(G. cuvieri) Rhim gazelle
Rhim gazelle
(G. leptoceros) Goitered gazelle
Goitered gazelle
(G. subgutturosa)

Litocranius

Gerenuk
Gerenuk
(L. walleri)

Nanger

Dama gazelle
Dama gazelle
(N. dama) Grant's gazelle
Grant's gazelle
(N. granti) Soemmerring's gazelle
Soemmerring's gazelle
(N. soemmerringii)

Procapra

Mongolian gazelle
Mongolian gazelle
(P. gutturosa) Goa (P. picticaudata) Przewalski's gazelle
Przewalski's gazelle
(P. przewalskii)

Saigini

Pantholops

Tibetan antelope
Tibetan antelope
(P. hodgsonii)

Saiga

Saiga antelope
Saiga antelope
(S. tatarica)

Neotragini

Dorcatragus

Beira (D. megalotis)

Madoqua

Günther's dik-dik
Günther's dik-dik
(M. guentheri) Kirk's dik-dik
Kirk's dik-dik
(M. kirkii) Silver dik-dik
Silver dik-dik
(M. piacentinii) Salt's dik-dik
Salt's dik-dik
(M. saltiana)

Neotragus

Bates's pygmy antelope
Bates's pygmy antelope
(N. batesi) Suni
Suni
(N. moschatus) Royal antelope
Royal antelope
(N. pygmaeus)

Oreotragus

Klipspringer
Klipspringer
(O. oreotragus)

Ourebia

Oribi
Oribi
(O. ourebi)

Raphicerus

Steenbok
Steenbok
(R. campestris) Cape grysbok
Cape grysbok
(R. melanotis) Sharpe's grysbok
Sharpe's grysbok
(R. sharpei)

Suborder Suina

Suidae

Babyrousa

Buru babirusa
Buru babirusa
(B. babyrussa) North Sulawesi babirusa
North Sulawesi babirusa
(B. celebensis) Togian babirusa
Togian babirusa
(B. togeanensis)

Hylochoerus

Giant forest hog
Giant forest hog
(H. meinertzhageni)

Phacochoerus

Desert
Desert
warthog (P. aethiopicus) Common warthog
Common warthog
(P. africanus)

Porcula

Pygmy hog
Pygmy hog
(P. salvania)

Potamochoerus

Bushpig
Bushpig
(P. larvatus) Red river hog
Red river hog
(P. porcus)

Sus (Pigs)

Palawan bearded pig
Palawan bearded pig
(S. ahoenobarbus) Bornean bearded pig
Bornean bearded pig
(S. barbatus) Indo-chinese warty pig (S. bucculentus) Visayan warty pig
Visayan warty pig
(S. cebifrons) Celebes warty pig
Celebes warty pig
(S. celebensis) Flores
Flores
warty pig (S. heureni) Oliver's warty pig
Oliver's warty pig
(S. oliveri) Philippine warty pig
Philippine warty pig
(S. philippensis) Wild boar
Wild boar
(S. scrofa) Timor
Timor
warty pig (S. timoriensis) Javan warty pig
Javan warty pig
(S. verrucosus)

Tayassuidae

Tayassu

White-lipped peccary
White-lipped peccary
(T. pecari)

Catagonus

Chacoan peccary
Chacoan peccary
(C. wagneri)

Pecari

Collared peccary
Collared peccary
(P. tajacu) Giant peccary (P. maximus)

Suborder Tylopoda

Camelidae

Lama

Llama
Llama
(L. glama) Guanaco
Guanaco
(L. guanicoe)

Vicugna

Vicuña
Vicuña
(V. vicugna) Alpaca
Alpaca
(V. pacos)

Camelus

Dromedary
Dromedary
(C. dromedarius) Bactrian camel
Bactrian camel
(C. bactrianus) Wild Bactrian camel
Bactrian camel
(C. ferus)

Whippomorpha
Whippomorpha
(unranked clade)

Hippopotamidae

Hippopotamus

Hippopotamus
Hippopotamus
(H. amphibius)

Choeropsis

Pygmy hippopotamus
Pygmy hippopotamus
(C. liberiensis)

v t e

Heraldry

Armiger Law of heraldic arms Grant of arms Blazon

Officers and officials

Authorities Officers of arms

King of Arms, Herald, Pursuivant

Private Officer of Arms

Conventional elements of coats of arms

Escutcheon Chief Field (Tincture) Division Supporter Supporter Slogan (battle cry) Crest Torse Mantling Helmet/Galero Crown/Coronet Compartment Order Ordinaries Charges Motto Dexter Sinister (right) (left)

Types of coats of arms

National coat of arms Ecclesiastical heraldry Burgher arms Civic heraldry Canting arms Attributed arms

Heraldic achievement

Escutcheon (shield)

Field

Divisions Variations

Charge Chief Lines Ordinary Augmentation of honour Cadency Marshalling

Quartering Impalement

Pale Bar Bend Bordure Canton Chevron Cross Fess Flaunch Gyron Label Lozenge Orle Pall Roundel Saltire

Creatures

Beasts

Bear Boar Bull/Ox Dog/Hound Camelopard (giraffe) Hind/Stag (deer) Kangaroo Leopard Lion Wolf

Birds

Cock Dove Eagle Martlet Pelican Rook

Sea creatures

Dolphin Ged Seahorse Lucy (esox) Scallop

Legendary creatures

Allocamelus Alphyn Basilisk Biscione Chollima Cockatrice Dragon Enfield Garuda Griffin/Keythong Harpy Konrul Lampago Lindworm Manticore Mermaid Pantheon Panther Pegasus Phoenix Salamander Hippocampus Hippogriff Sea-lion Turul Tyger Unicorn Wyvern Yale

Others

Bat Bee Crapaudy (toad) Emmet (ant) Serpent

Knots

Bourchier knot Bowen knot Cavendish knot Dacre knot Harrington knot Hastings knot Heneage knot Hinckaert knot Hungerford knot Lacy knot Medici knot Morvillier knot Ormonde knot Savoy knot Shakespeare knot Stafford knot Trafford knot Tristram knot Wake knot

Tinctures Rules Tricking Hatching

Metals

     Argent
Argent
(white)      Or (gold)

Colours

     Gules (red)      Sable (black)      Azure (blue)      Vert (green)      Purpure (purple)

Furs

Ermines

Erminois Erminites Pean

Vair

Potent

Stains

     Murrey
Murrey
(mulberry)      Sanguine (blood red)      Tenné

Non-traditional1

Metals

     Copper

Colours

     Bleu-celeste      Carnation      Cendrée      Orange      Rose

External

Crowns and coronets Crest Compartment Helmet Mantling Motto Supporter Torse Mantle and pavilion

See also

List of oldest heraldry Heraldic flag
Heraldic flag
(Banner of arms) Heraldic badge Women in heraldry Socialist heraldry Vexillology

1 Non-traditional, rarely used traditions in italic (typically regional or modern, considered unheraldic by some) Heraldry
Heraldry
portal Portal:Heraldry/Web resources

Authority control

LCCN: sh85146670 GND: 4066129-5

Taxon identifiers

Wd: Q58697 ADW: Sus_scrofa ARKive: sus-scrofa EoL: 328663 EPPO: SUXXSC Fauna Europaea: 305279 Fossilworks: 104172 GBIF: 7705930 iNaturalist: 42134 ITIS: 180722 IUCN: 41775 MSW: 1

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