White-headed lemur
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The white-headed lemur (''Eulemur albifrons''), also known as the white-headed brown lemur, white-fronted brown lemur, or white-fronted lemur, is a species of
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians ( monkeys and apes, the latter includin ...
in the family Lemuridae. It is only found in north-eastern
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Afric ...
. It is
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose nu ...
and is usually found in rainforest treetops. It was formerly recognised as a subspecies of the
common brown lemur The common brown lemur (''Eulemur fulvus'') is a species of lemur in the family Lemuridae. It is found in Madagascar and has been introduced to Mayotte. Taxonomy Five additional currently recognized species of lemur were until 2001 considered ...
(''Eulemur fulvus'').


Physical characteristics

The white-headed lemur is a medium-sized
lemur Lemurs ( ) (from Latin ''lemures'' – ghosts or spirits) are wet-nosed primates of the superfamily Lemuroidea (), divided into 8 families and consisting of 15 genera and around 100 existing species. They are endemic to the island of Madagas ...
and has a horizontal posture, which is suited to its way of movement. It has a long furry tail assisting it in maintaining its balance as it lands from leaping at a considerable distance. Males have gray-brown upper parts, with darker lower limbs and tail, paler gray upper parts, gray head and face and a darker crown. Females have redder-brown upper parts, paler underparts and darker feet than males. The cheeks and beards are white, bushy and pronounced in males, reddish-brown and less bushy in females. The head, face and muzzle of the female are dark gray, but without the bushy cheeks of the male. The white-headed lemur has an average body weight of , and body length of , and its tail can grow up to . White-headed lemurs can live up to 30 years. The species was previously classified as ''Eulemur fulvus albifrons'', a subspecies of the
common brown lemur The common brown lemur (''Eulemur fulvus'') is a species of lemur in the family Lemuridae. It is found in Madagascar and has been introduced to Mayotte. Taxonomy Five additional currently recognized species of lemur were until 2001 considered ...
, and although very similar in appearance genetic analysis supports distinct species status.


Habitat and range

This lemur is mostly found in moist lowland and montane rainforests. The white-headed lemur is
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose nu ...
and spends most of its time in the upper layers of the forest. It is only found in north-eastern
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Afric ...
.


Behaviour and mating

This species is cathemeral, meaning it is active at varying times throughout the day and night. It has an omnivorous diet consisting of fruit, mature leaves, flowers, bark, sap, soil, insects,
centipede Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, ...
s and millipedes. The diet changes depending on the reproductive stage of the females, during gestation both males and females supplement their diet with flowers, during lactation, more flowers and young leaves and when nonreproductive they eat more mature leaves and miscellaneous other items like millipedes. The mating system of this lemur has not been reported. However, other species in the genus '' Eulemur'' are either
monogamous Monogamy ( ) is a form of dyadic relationship in which an individual has only one partner during their lifetime. Alternately, only one partner at any one time ( serial monogamy) — as compared to the various forms of non-monogamy (e.g., pol ...
or
polygynous Polygyny (; from Neoclassical Greek πολυγυνία (); ) is the most common and accepted form of polygamy around the world, entailing the marriage of a man with several women. Incidence Polygyny is more widespread in Africa than in any ...
. It is likely that the white-headed lemur is similar. For the first three weeks of its life, a young lemur hangs onto its mother's belly, altering its grasp only to nurse. After three weeks have passed, it shifts and rides on the mother's back. It then starts to take its first steps. Following this, it starts to sample solid food, nibbling on whatever the other members of the group happen to be eating. This is its first sign of independence. Nursing continues but its importance in the infant's diet tapers. The young lemur is weaned after approximately four months, usually by January. Unlike other members of the genus, females are not usually dominant to males, so the degree to which females exert active
mate choice Mate choice is one of the primary mechanisms under which evolution can occur. It is characterized by a "selective response by animals to particular stimuli" which can be observed as behavior.Bateson, Paul Patrick Gordon. "Mate Choice." Mate Choic ...
is not known. It forms multi-male multi-female groups. Depending on the population, the size of these groups can vary, they usually number three to twelve individuals, although five to seven seems to be average.


Status

''Eulemur albifrons'' is classified as Vulnerable on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
.


References

{{Taxonbar, from=Q1150284 True lemurs Mammals described in 1796 Taxa named by Étienne Geoffroy Saint-Hilaire